Tarsal Claws
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This glossary describes the terms used in formal descriptions of
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s; where applicable these terms are used in describing other
arachnid Arachnida () is a class of joint-legged invertebrate animals (arthropods), in the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vinegaroon ...
s. Links within the glossary are shown .


Terms


A

Abdomen or opisthosoma: One of the two main body parts ( tagmata), located towards the posterior end; see also Abdomen § Other animals Accessory claw: Modified at the tip of the in web-building spiders; used with to grip strands of the web Anal tubercle: A small protuberance (tubercule) above the through which the
anus The anus (Latin, 'ring' or 'circle') is an opening at the opposite end of an animal's digestive tract from the mouth. Its function is to control the expulsion of feces, the residual semi-solid waste that remains after food digestion, which, d ...
opens Apodeme → Apophysis (plural apophyses): An outgrowth or process changing the general shape of a body part, particularly the appendages; often used in describing the male → Atrium (plural atria): An internal chamber at the entrance to the in female
haplogyne The Haplogynae or haplogynes are one of the two main groups into which araneomorph spiders have traditionally been divided, the other being the Entelegynae. Morphological phylogenetic studies suggested that the Haplogynae formed a clade; more recen ...
spiders


B

Bidentate: Having two Book lungs: Respiratory organs on the ventral side (underside) of the , in front of the , opening through narrow slits; see also Book lungs Branchial operculum → Bulbus →


C

Calamistrum (plural calamistra): Modified setae (bristles) on the of the fourth leg of spiders with a , arranged in one or more rows or in an oval shape, used to comb silk produced by the cribellum; see also
Calamistrum In spiders, the calamistrum is a row of specialized leg bristles used to comb out fine bands of silk. It is only found on cribellate spiders, that is, spiders that possess the spinning organ known as the cribellum. The calamistrum and cribellum ar ...
Caput (plural capita) → Carapace: A hardened plate (sclerite) covering the upper (dorsal) portion of the ; see also
Carapace A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tor ...
Carpoblem: The principal on the male ; also just called the tibial apophysis Cephalic region or caput: The front part of the , separated from the thoracic region by the Cephalothorax or prosoma: One of the two main body parts ( tagmata), located towards the
anterior Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
end, composed of the head ( or caput) and the thorax (thoracic region), the two regions being separated by the ; covered by the and bearing the , legs, and mouthparts Cervical groove: A shallow U-shaped groove, separating the and thoracic regions of the Chelate: A description of a where the closes against a tooth-like process Chelicera (plural chelicerae): One of two appendages at the front of the , made up of basal portion, the , and the ; sometimes called the jaw; see also
Chelicerae The chelicerae () are the mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as "jaws", chelicerae may be shaped as either articulated fangs, or similarly ...
Cheliceral furrow: A shallow groove on the basal portion of a accommodating the , usually having on its margins Cheliceral tooth: A tooth-like extension on the margin of the Chilum: A small hardened plate (sclerite) at the base of the , under the Claw → Claw tuft: A dense group of hairs or bristles (setae) underneath the paired , usually well developed in hunting spiders Clypeus: The area of the between the anterior (frontmost) and the anterior edge of the carapace Colulus: A short protuberance in the middle of the underside of the in front of the , considered to be a modification of the Conductor → Copulatory duct: An internal tube (duct) from the through which sperm enters the female; separate from the duct through which fertilized eggs pass in entelegyne spiders Copulatory opening: An opening in the ventral of female spiders; in entelegyne spiders, a double opening in the through which the is inserted; in
haplogyne The Haplogynae or haplogynes are one of the two main groups into which araneomorph spiders have traditionally been divided, the other being the Entelegynae. Morphological phylogenetic studies suggested that the Haplogynae formed a clade; more recen ...
spiders, a single opening through which male is inserted Coxa → Crenulate: Having longitudinal ridges Cribellum: A sieve-like plate in front of the , used in conjunction with the ; spiders with a cribellum are called cribellate, those without ecribellate; see also
Cribellum Cribellum literally means "little sieve", and in biology the term generally applies to anatomical structures in the form of tiny perforated plates. In certain groups of diatoms it refers to microscopically punctured regions of the frustule, or ...
Cuspule: A small spiny outgrowth ("wart") on the and of
Mygalomorphae The Mygalomorphae, or mygalomorphs, are an infraorder of spiders, and comprise one of three major groups of living spiders with over 3000 species, found on all continents except Antarctica. Many members are known as trapdoor spiders due to the ...
Cymbium (plural cymbia): The end part of the of the in a mature male, usually hollowed out and bearing the


D

Dionychous: With two on the of each leg; a feature of spiders in the clade
Dionycha The Dionycha are a clade of spiders (Araneomorphae:Entelegynae), characterized by the possession of two tarsal claws with tufts of hairs (setae) beside them, which produce strong adhesion, enabling some species to climb glass. The circumscriptio ...
Dorsal groove → Dorsum: The upper (dorsal) portion or surface of the body or ; the adjective dorsal may be applied to the upper portion or surface of any part of the body; →


E

Ecribellate → Embolus → Endite → Endosternite: An internal hardened plate (sclerite) Entelegyne: A spider whose female has an and separate ducts leading to for sperm storage and to the uterus for fertilization, creating a "flow-through" system; → ; see also
Entelegynae The Entelegynae or entelegynes are a subgroup of araneomorph spiders, the largest of the two main groups into which the araneomorphs were traditionally divided. Females have a genital plate ( epigynum) and a "flow through" fertilization system; m ...
Epigastric furrow or epigastric fold: A transverse slit towards the front (anterior) of underside of the ; the front pair of open at the edge of this furrow as do the genital openings () Epigyne or epigynum (plural epigynes): A hardened plate on the underside of the female in which the are located; only fully developed in mature females of spiders; see also
Epigyne The epigyne or epigynum is the external genital structure of female spiders. As the epigyne varies greatly in form in different species, even in closely related ones, it often provides the most distinctive characteristic for recognizing species. ...
Eyes: The basic number of eyes is eight, typically arranged in two rows (e.g. as in
Gnaphosidae Ground spiders comprise Gnaphosidae, the seventh largest spider family with over 2,000 described species in over 100 genera distributed worldwide. There are 105 species known to central Europe, and common genera include ''Gnaphosa'', ''Drassodes ...
); the front row are the anterior eyes, the row behind the posterior eyes; the four eyes to the edges are the lateral eyes, the four eyes in the centre the median eyes; the anterior median eyes are called the main eyes or direct eyes, while the other eyes are called the secondary eyes or indirect eyes; the number of eyes, their sizes and arrangement varies widely and is characteristic of spider families → ,


F

Fang: The final hinged part of the , normally folded down into a groove in the basal part of the chelicera; venom is injected via an opening near the tip of the fang Femur → Fertilization duct: A duct in female spiders leading from the to the uterus Folium: A broad leaf-like marking along the medial line of the top of the Fossa (plural fossae): A pit or depression, typically in the Fovea (also called thoracic furrow or dorsal groove): A depression or pit in the centre of the of a spider marking an inward projection of the exoskeleton to which stomach muscles are attached


G

Genital opening → Gnathocoxa → Gonopore: The genital opening; located in the
epigastric furrow The anatomy of spiders includes many characteristics shared with other arachnids. These characteristics include bodies divided into two tagmata (sections or segments), eight jointed legs, no wings or antennae, the presence of chelicerae and pedi ...
; the opening of the duct from the uterus in females and from the testes in males; see also
Gonopore A gonopore, sometimes called a gonadopore, is a genital pore in many invertebrates. Hexapods, including insects have a single common gonopore, except mayflies, which have a pair of gonopores. More specifically, in the unmodified female it is the ...


H

Haematodocha → Haplogyne: A spider whose female lacks an and in which the same ducts are used to transport sperm to the uterus and to the ; → ; see also
Haplogynae The Haplogynae or haplogynes are one of the two main groups into which araneomorph spiders have traditionally been divided, the other being the Entelegynae. Morphological phylogenetic studies suggested that the Haplogynae formed a clade; more recen ...
Heart mark: A narrow marking along the top of the roughly corresponding to the location of the heart


L

Labio-sternum mound: A mound separating the from the , found in some
tarantula Tarantulas comprise a group of large and often hairy spiders of the family Theraphosidae. , 1,040 species have been identified, with 156 genera. The term "tarantula" is usually used to describe members of the family Theraphosidae, although ...
s, where it can be a diagnostic feature Labium (plural labia): A hardened plate (sclerite) between the at the front of the ; see also Arthropod mouthparts: Labium Labrum (plural labra): A component (the "upper lip") of the mouthparts, concealed by the ; see also Arthropod mouthparts: Labrum Lateral (applied to appendages): Viewed from above or below, the sides of the leg or , i.e. the surfaces parallel to the line of sight; → , Laterigrade: With legs directed to the side, hence appearing like and moving like a crab; → Leg formula: The legs are numbered from the front from I to IV; the relative length of the legs can be represented by four numbers from the longest to the shortest; e.g. 1423 = first leg (leg I) is longest and third leg (leg III) is shortest Leg parts or segments →


M

Main eye: One of the two anterior median eyes (AME) that have the light-detecting units (rhabdomeres) pointing towards the source; particularly enlarged in the families
Salticidae Jumping spiders are a group of spiders that constitute the family Salticidae. As of 2019, this family contained over 600 described genera and over 6,000 described species, making it the largest family of spiders at 13% of all species. Jumping spi ...
and
Thomisidae The Thomisidae are a family of spiders, including about 170 genera and over 2,100 species. The common name crab spider is often linked to species in this family, but is also applied loosely to many other families of spiders. Many members of thi ...
; → , Mastidion (plural mastidia): A projection or bump on the chelicerae (not to be confused with ) Maxilla (also called endite or gnathocoxa): Modified of the , used in feeding; not the structure called by this name in other arthropods, for which see
Maxilla (arthropod mouthpart) In arthropods, the maxillae (singular maxilla) are paired structures present on the head as mouthparts in members of the clade Mandibulata, used for tasting and manipulating food. Embryologically, the maxillae are derived from the 4th and 5th seg ...
Metatarsus →


O

Operculum or branchial operculum (plural opercula): One of the plates on the surface of the , just in front of the , covering the , often pale, yellow or orange in colour; two pairs in
Mygalomorphae The Mygalomorphae, or mygalomorphs, are an infraorder of spiders, and comprise one of three major groups of living spiders with over 3000 species, found on all continents except Antarctica. Many members are known as trapdoor spiders due to the ...
, one pair in other spiders Opisthosoma →


P

Palp → Palpal bulb (also called bulbus, palpal organ, genital bulb): The copulatory organ of the male spider, carried on the modified last segment of the , used to transfer sperm to the female; see also
Palpal bulb The two palpal bulbs – also known as palpal organs and genital bulbs – are the copulatory organs of a male spider. They are borne on the last segment of the pedipalps (the front "limbs" of a spider), giving the spider an appearance often descr ...
:Conductor: A part of the palpal bulb that accompanies and supports the embolus :Embolus: The final part of the palpal bulb containing the end of the sperm duct, usually thin, sharp-tipped and strongly hardened (sclerotized) :Haematodocha (plural haematodochae): A membranous, inflatable part of the palpal bulb :Median apophysis: A projection (apophysis) of the palpal bulb, below the conductor :Subtegulum: A hardened part of the palpal bulb nearer its base than the tegulum :Tegulum: The main hardened part of the palpal bulb Paracymbium: An outgrowth of the on the male Patella → Paturon: The basal segment of a to which the connects Pedicel or petiolus: The narrow connection between the and Pedipalp (plural pedipalps or pedipalpi; also called just palp): The second appendage of the in front of the first leg; bears the in male spiders; → , see also
Pedipalp Pedipalps (commonly shortened to palps or palpi) are the second pair of appendages of chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to the chelicerae ("jaws") and ...
Plumose: Used to describe hairs () having outgrowths or appendages on two sides, giving a feather-like appearance; the appendages vary in number, size and arrangement Pluridentate: Having multiple Procurved: Used to describe a structure which is curved in such a way that the outer edges are in front of the central part; opposite Prolateral: Viewed from above or below, the side of a leg or nearest the mouth, i.e. the side facing forward; opposite (includes diagram) Promarginal: The side of the facing forward; particularly used for describing ; opposite Prosoma →


R

Rastellum (plural rastella): An often rake-like structure at end of the in
mygalomorph The Mygalomorphae, or mygalomorphs, are an infraorder of spiders, and comprise one of three major groups of living spiders with over 3000 species, found on all continents except Antarctica. Many members are known as trapdoor spiders due to the ...
spiders; used in burrowing Rebordered: Having a thickened edge (i.e. border) (more rarely seen as reborded, from the French , e.g. in Levy (1984)); particularly used of the Receptaculum (plural receptacula) → Recurved: Used to describe a structure which is curved in such a way that the outer edges are behind the central part; opposite Retrolateral: Viewed from above or below, the side of a leg or furthest from the mouth, i.e. the side facing backwards; opposite Retrolateral tibial apophysis: A backward-facing projection on the tibia of the male ; distinguishing feature of the
RTA clade The RTA clade is a clade of araneomorph spiders, united by the possession of a retrolateral tibial apophysis – a backward-facing projection on the tibia of the male pedipalp. The clade contains over 21,000 species, almost half the current total ...
Retromarginal: The side of the facing backward (towards the posterior end of the spider); particularly used for describing ; opposite


S

Scape: An elongated process or appendage of some Sclerite: A single hardened (sclerotized) part of the external covering (tegument, exoskeleton) Scopula (plural scopulae): A brush of hairs (setae); called a when on the end of the foot (tarsus), where it improves adhesion Scutum (plural scuta): A hardened (sclerotized) plate on the of some spiders Secondary eye: An eye belonging to the three pairs – anterior lateral eyes (ALE), posterior median eyes (PME) and posterior lateral eyes (PLE) – that are primarily movement detectors and have the light-detecting units (rhabdomeres) pointing away from the source; → , Segments or articles of the legs and : :Coxa (plural coxae): First leg segment, between body and trochanter; the coxa of the is heavily modified to form the or endite :Trochanter: Second leg segment, between coxa and femur :Femur (plural femora): Third leg segment, between trochanter and patella :Patella (plural patellae): Fourth leg segment, between femur and tibia :Tibia (plural tibiae): Fifth leg segment, between patella and metatarsus :Metatarsus (plural metatarsi; also called basitarsus): Sixth leg segment, between tibia and tarsus; absent in the :Tarsus (plural tarsi; also called telotarsus): Seventh (last) leg segment, after the metatarsus Seta (plural setae): A bristle; spiders have a variety of hair-like structures of increasing size that are referred to as hairs, bristles (setae) or Sigillum (plural sigilla): A circular indentation on the outside of the spider, showing where an internal muscle is attached; particularly on the in some
Mygalomorphae The Mygalomorphae, or mygalomorphs, are an infraorder of spiders, and comprise one of three major groups of living spiders with over 3000 species, found on all continents except Antarctica. Many members are known as trapdoor spiders due to the ...
and on the in some
Araneomorphae The Araneomorphae (also called the Labidognatha) are an infraorder of spiders. They are distinguishable by chelicerae (fangs) that point diagonally forward and cross in a pinching action, in contrast to the Mygalomorphae (tarantulas and their cl ...
Sperm duct: A duct in the male used to store sperm Spermatheca (plural spermathecae; also called receptulacum, receptulacum seminis): A structure in the of female spiders used to store sperm after insemination and before fertilization; see also
Spermatheca The spermatheca (pronounced plural: spermathecae ), also called receptaculum seminis (plural: receptacula seminis), is an organ of the female reproductive tract in insects, e.g. ants, bees, some molluscs, oligochaeta worms and certain other ...
Spigot: A small pointed or cylindrical structure at the tip of a from which silk emerges Spine: A pointed, rigid structure on body and legs, usually with a basal joint; spiders have a variety of hair-like structures of increasing size that are referred to as hairs, bristles (setae) or spines Spinneret: An appendage borne on the , typically one of six arranged in three pairs: anterior (anterior median, AMS), median (posterior median, PMS) and posterior (posterior lateral, PLS); silk emerges from small on the spinnerets; see also
Spinneret A spinneret is a silk-spinning organ of a spider or the larva of an insect. Some adult insects also have spinnerets, such as those borne on the forelegs of Embioptera. Spinnerets are usually on the underside of a spider's opisthosoma, and are ...
Sternum: The lower (ventral) portion of the Stridulating organ: A series of thin ridges on a hardened part of the body; rubbing this with a matching series of short, stiff bristles (setae) elsewhere on the body creates a sound Subtegulum →


T

Tapetum (plural tapeta): A light-reflecting layer in a making the eye appear pale Tarsal claw (claw): One of a set of claws at the tip of the ; there may be a single pair, often concealed in a , or an additional third central claw, much smaller than the other two Tarsal organ: a small pit, usually spherical and on the surface of each , believed to respond to
humidity Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present. Humidity depe ...
Tarsus → Teeth: Pointed growths or bumps along the margins of the Tegulum → Thoracic furrow → Tibia → Trachea (plural tracheae): A thin hardened internal tube, part of the respiratory system in many
araneomorph The Araneomorphae (also called the Labidognatha) are an infraorder of spiders. They are distinguishable by chelicerae (fangs) that point diagonally forward and cross in a pinching action, in contrast to the Mygalomorphae (tarantulas and their cl ...
spiders; opens on the underside of the via a tracheal spiracle; see Trachea § Invertebrates Trichobothrium (plural trichobothria): A slender hair-like structure of variable length on the legs and , arising from a special socket; used to detect air movements, including sounds; → , Trochanter →


U

Unidentate: Having a single tooth


V

Venter (or ventrum): The lower (ventral) portion or surface of the body or ; the adjective ventral may be applied to the lower portion or surface of any part of the body; →


Abbreviations

Some abbreviations commonly found in descriptions of spider anatomy include: *ALE: anterior lateral eyes → *ALS: anterior lateral spinnerets → *AME: anterior median eyes → *DTA: dorsal tegular apophysis, on the back of the *DTiA: dorsal tibial apophysis, on the back of a *LTA: lateral tegular apophysis'', apophysis'' on the side of the *MOQ: median ocular quadrangle, the quadrangle formed by the four median eyes, → *PLE: posterior lateral eyes → *PLS: posterior lateral spinnerets → *PME: posterior median eyes → *PMS: posterior median spinnerets → *RCF: retrolateral cymbial fold, fold on the surface of the *RTA: retrolateral tibial apophysis, on the surface of a *VTA: ventral tegular apophysis, on the underside of the *VTiA: ventral tibial apophysis, on the lower surface of a


See also

*
Spider anatomy The anatomy of spiders includes many characteristics shared with other arachnids. These characteristics include bodies divided into two tagmata (sections or segments), eight jointed legs, no wings or antennae, the presence of chelicerae and pedi ...
*
Glossary of entomology terms This glossary of entomology describes terms used in the formal study of insect species by entomologists. A–C A synthetic chlorinated hydrocarbon insecticide, toxic to vertebrates. Though i ...
*
Anatomical terms of location Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...


References


Bibliography

* * * * * * * {{cite book , editor1-last=Ubick , editor1-first=Darrell , editor2-last=Paquin , editor2-first=Pierre , editor3-last=Cushing , editor3-first=Paula E. , editor4-last=Roth , editor4-first=Vincent , title=Spiders of North America: An Identification Manual , date=2009 , publisher=American Arachnological Society , isbn=978-0977143900 , edition=1st Spider terms