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''Taohelong'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
nodosaurid Nodosauridae is a family of ankylosaurian dinosaurs, from the Late Jurassic to the Late Cretaceous period in what is now North America, South America, Europe, and Asia. Description Nodosaurids, like their close relatives the ankylosaurids, we ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
known from
Lower Cretaceous Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Lower Wick is a small hamlet located in the county of Gloucestershire, England. It is situated about five miles south west of Dursley, eig ...
rocks in north-central China. ''Taohelong'' is based on Gansu Dinosaur Museum (GSDM) 00021, fossils including a tail vertebra, ribs, a left ilium (the main bone of the hip), and bony armor recovered from the Hekou Group in the Lanzhou-Minhe Basin. The animal's armor includes part of a "sacral shield", a carpet of osteoderms over the hips found in some other
ankylosauria Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. ...
ns. ''Taohelong'' was named and described in
2013 File:2013 Events Collage V2.png, From left, clockwise: Edward Snowden becomes internationally famous for leaking classified NSA wiretapping information; Typhoon Haiyan kills over 6,000 in the Philippines and Southeast Asia; The Dhaka garment fa ...
by Yang Jing-Tao, You Hai-Lu, Li Da-Qing, and Kong De-Lai. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
is ''Taohelong jinchengensis''. The generic name means "dragon (''long'') of the river (''he'') Tao". The specific name refers to the provenance at Jincheng. The describers established some diagnostic traits. The
neural channel In biology, the nervous system is the highly complex part of an animal that coordinates its actions and sensory information by transmitting signals to and from different parts of its body. The nervous system detects environmental changes ...
of the tail vertebra has a cross-section like an inverted trapezium. In top view the profile of the outer rim of the ilium is like a mirrored "S". The osteoderms of the sacral shield are irregular in both shape and size. ''Taohelong'' was placed in the
Nodosauridae Nodosauridae is a family of ankylosaurian dinosaurs, from the Late Jurassic to the Late Cretaceous period in what is now North America, South America, Europe, and Asia. Description Nodosaurids, like their close relatives the ankylosaurids, we ...
, more precisely in the
Polacanthinae Polacanthinae is a subfamily of ankylosaurs, most often nodosaurids, from the Late Jurassic through Early Cretaceous of Europe and potentially North America and Asia. The group is defined as the largest clade closer to '' Polacanthus foxii'' tha ...
. Yang ''et al.'' performed a
phylogenetic analysis In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
and found ''Taohelong'' to be the
sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
to ''
Polacanthus foxii ''Polacanthus'', deriving its name from the Ancient Greek polys-/πολύς- "many" and akantha/ἄκανθα "thorn" or "prickle", is an early armoured, spiked, plant-eating ankylosaurian dinosaur from the early Cretaceous period of England. In ...
'', making it the first polacanthine to be described from Asia.


See also

*
Timeline of ankylosaur research This timeline of ankylosaur research is a chronological listing of events in the history of paleontology focused on the ankylosaurs, quadrupedal herbivorous dinosaurs who were protected by a covering bony plates and spikes and sometimes by a club ...


References

Nodosaurids Early Cretaceous dinosaurs of Asia Fossil taxa described in 2013 Paleontology in Gansu Ornithischian genera {{ankylosaur-stub