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The takin (''Budorcas taxicolor''; ), also called cattle chamois or gnu goat, is a large species of
ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, cam ...
of the subfamily Caprinae found in the eastern
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
. It includes four
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
: the
Mishmi takin The Mishmi takin (''Budorcas taxicolor taxicolor'') is an endangered goat-antelope native to India, Myanmar and the People's Republic of China. It is a subspecies of takin. The Mishmi takin lives in Northeast India and eats bamboo and willow sho ...
(''B. t. taxicolor''), the
golden takin The golden takin (''Budorcas taxicolor bedfordi'') is an endangeredgoat-antelope (takin), native to the Qin Mountains in China's southern Shaanxi province. Golden takins have unique adaptations that help them stay warm and dry during cold Himala ...
(''B. t. bedfordi''), the Tibetan (or Sichuan) takin (''B. t. tibetana''), and the Bhutan takin (''B. t. whitei''). Whilst the takin has in the past been placed together with the muskox in the tribe Ovibovini, more recent
mitochondrial A mitochondrion (; ) is an organelle found in the cells of most Eukaryotes, such as animals, plants and fungi. Mitochondria have a double membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is use ...
research shows a closer relationship to ''
Ovis ''Ovis'' is a genus of mammals, part of the Caprinae subfamily of the ruminant family Bovidae. Its seven highly sociable species are known as sheep or ovines. Domestic sheep are members of the genus, and are thought to be descended from the wil ...
'' (sheep). Its physical similarity to the muskox is therefore an example of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
. The takin is the national animal of
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainous ...
.


Etymology

The specific name ''taxicolor'' comes from la, taxus, , badger and la, color, , hue, label=none referring to badger-like coloration.


Appearance

The takin rivals the muskox as the largest and stockiest of the subfamily Caprinae, which includes
goat The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the a ...
s,
sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus ''Ovis'', in everyday usage it almost always refers to domesticated s ...
, and similar species. Its short legs are supported by large, two-toed hooves, which each have a highly developed spur. It has a stocky body and a deep chest. Its large head is distinctive by its long, arched nose and stout
horn Horn most often refers to: *Horn (acoustic), a conical or bell shaped aperture used to guide sound ** Horn (instrument), collective name for tube-shaped wind musical instruments *Horn (anatomy), a pointed, bony projection on the head of various ...
s, which are ridged at the base. These horns are present in both sexes, and run parallel to the skull before turning upwards to a short point; they are about long, but can grow up to . Its long, shaggy coat is light in color with a dark stripe along the back, and males (bulls) also have dark faces. Four subspecies of takin are currently recognised, and these tend to show a variation in coat colour. Their thick
wool Wool is the textile fibre obtained from sheep and other mammals, especially goats, rabbits, and camelids. The term may also refer to inorganic materials, such as mineral wool and glass wool, that have properties similar to animal wool. As ...
often turns black in colour on their undersides and legs. Their overall coloration ranges from dark blackish to reddish-brown suffused with greyish-yellow in the eastern Himalayas to lighter yellow-gray in the Sichuan Province to mostly golden or (rarely) creamy-white with fewer black hairs in the
Shaanxi Province Shaanxi (alternatively Shensi, see § Name) is a landlocked province of China. Officially part of Northwest China, it borders the province-level divisions of Shanxi (NE, E), Henan (E), Hubei (SE), Chongqing (S), Sichuan (SW), Gansu (W), Ningx ...
. The legend of the ' golden fleece' sought by Jason and the
Argonauts The Argonauts (; Ancient Greek: ) were a band of heroes in Greek mythology, who in the years before the Trojan War (around 1300 BC) accompanied Jason to Colchis in his quest to find the Golden Fleece. Their name comes from their ship, '' Argo'', ...
may have been inspired by the lustrous coat of the
golden takin The golden takin (''Budorcas taxicolor bedfordi'') is an endangeredgoat-antelope (takin), native to the Qin Mountains in China's southern Shaanxi province. Golden takins have unique adaptations that help them stay warm and dry during cold Himala ...
(''B. t. bedfordi''). Hair length can range from , on the flanks of the body in summer, up to on the underside of the head in winter. In height, takin stand at the shoulder, but measure a relatively short in head-and-body length, with the tail adding only an additional . Measurements of weights vary, but according to most reports, the males are slightly larger, weighing against in females. Sources including Betham (1908) report that females are larger, with the largest captive takin known to the author, at , having been female. Takin can weigh up to or in some cases. Instead of relying on localized scent glands, the takin secretes an oily, strong-smelling substance over its whole body, enabling it to mark objects such as trees.Macdonald, D. (2001) ''The New Encyclopedia of Mammals''. Oxford University Press, Oxford. via arkive.org A prominent nose with a swollen appearance caused biologist
George Schaller George Beals Schaller (born 1933) is a German-born American mammalogist, biologist, conservationist and author. Schaller is recognized by many as the world's preeminent field biologist, studying wildlife throughout Africa, Asia and South Americ ...
to liken the takin to a "bee-stung
moose The moose (in North America) or elk (in Eurasia) (''Alces alces'') is a member of the New World deer subfamily and is the only species in the genus ''Alces''. It is the largest and heaviest extant species in the deer family. Most adult mal ...
." Features reminiscent of familiar domesticated species have earned takins such nicknames as "cattle chamois" and "gnu goat."


Distribution and habitat

Takin are found from forested valleys to rocky, grass-covered alpine zones, at altitudes between above sea level. The Mishmi takin occurs in eastern Arunachal Pradesh, while the Bhutan takin is in western Arunachal Pradesh and Bhutan.
Dihang-Dibang Biosphere Reserve Dihang-Dibang or Dehang-Debang is a biosphere reserve constituted in 1998. It is in the Indian state of Arunachal Pradesh. The Mouling National Park and the Dibang Wildlife Sanctuary are located fully or partly within this biosphere reserve. The ...
in Arunachal Pradesh, India is a stronghold of both Mishmi, Upper Siang (Kopu) and Bhutan takins.


Behaviour and ecology

Takin are found in small family groups of around 20 individuals, although older males may lead more solitary existences. In the summer, herds of up to 300 individuals gather high on the mountain slopes. Groups often appear to occur in largest numbers when favorable feeding sites, salt licks, or hot springs are located. Mating takes place in July and August. Adult males compete for dominance by sparring head-to-head with opponents, and both sexes appear to use the scent of their own urine to indicate dominance. A single young is born after a gestation period of around eight months. Takin migrate from the upper pasture to lower, more forested areas in winter and favor sunny spots upon sunrise.''Animal Diversity Web'' (November, 2002
"Budorcas taxicolor"
(University of Michigan Museum of Zoology) via arkive.org
When disturbed, individuals give a 'cough' alarm call and the herd retreats into thick bamboo thickets and lies on the ground for camouflage.Huffman, Brent

''Ultimate Ungulate'' via arkive.org
Takin feed in the early morning and late afternoon, grazing on a variety of leaves and grasses, as well as bamboo shoots and flowers. They have been observed standing on their hind legs to feed on leaves over high. Salt is also an important part of their diets, and groups may stay at a mineral deposit for several days.


Threats

The takin is listed as Vulnerable on the IUCN Red List and considered Endangered in China. It is threatened by overhunting and the destruction of its natural habitat. It is not a common species naturally, and the population appears to have been reduced considerably. Takin horns have appeared in the illegal wildlife trade in Myanmar; and during three surveys carried out from 1999 to 2006 in the Tachilek market, a total of 89 sets of horns were observed openly for sale.


References

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Further reading

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External links

*
Wild creamy-white ''golden takins'' (b. t. bedfordi) in China
{{Taxonbar, from1=Q193469, from2=Q10741556 Caprids Mammals described in 1850 Taxa named by Brian Houghton Hodgson