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The skuas are a group of predatory
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...
s with seven species forming the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Stercorarius'', the only genus in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Stercorariidae. The three smaller skuas, the
long-tailed skua The long-tailed skua or long-tailed jaeger (''Stercorarius longicaudus'') is a seabird in the skua family Stercorariidae. Etymology The word "jaeger" is derived from the German word ''Jäger'', meaning "hunter". The English word "skua" comes fr ...
, the
Arctic skua The parasitic jaeger (''Stercorarius parasiticus''), also known as the Arctic skua, Arctic jaeger or parasitic skua, is a seabird in the skua family Stercorariidae. It is a migratory species that breeds in Northern Scandinavia, Scotland, Iceland ...
, and the
pomarine skua The pomarine jaeger (''Stercorarius pomarinus''), pomarine skua, or pomatorhine skua, is a seabird in the skua family Stercorariidae. It is a migrant, wintering at sea in the tropical oceans. Taxonomy Its relationships are not fully resolved; i ...
are called jaegers in
North American English North American English (NAmE, NAE) is the most generalized variety of the English language as spoken in the United States and Canada. Because of their related histories and cultures, plus the similarities between the pronunciations (accents), v ...
. The English word "skua" comes from the Faroese name for the
great skua The great skua (''Stercorarius skua''), sometimes known by the name bonxie in Britain, is a large seabird in the skua family Stercorariidae. It is roughly the size of a herring gull. It mainly eats fish caught at the sea surface or taken fr ...
, , with the island of
Skúvoy Skúvoy or Skúgvoy ( da, Skuø) is an island in the central Faroe Islands, located to the south of Sandoy. It is named after the large number of great skua present on the island (who have a habit of attacking intruders). There is only one se ...
renowned for its colony of that bird. The general Faroese term for skuas is . The word "jaeger" is derived from the
German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ger ...
word , meaning "hunter". The genus name ''Stercorarius'' is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
and means "of dung"; because the food disgorged by other birds when pursued by skuas was once thought to be excrement. Skuas nest on the ground in
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
and
Arctic region The Arctic ( or ) is a polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenland), Finland, Iceland, N ...
s, and are long-distance migrants. They have even been sighted at the
South Pole The South Pole, also known as the Geographic South Pole, Terrestrial South Pole or 90th Parallel South, is one of the two points where Earth's axis of rotation intersects its surface. It is the southernmost point on Earth and lies antipod ...
.


Biology and habits

Outside the breeding season, skuas take fish,
offal Offal (), also called variety meats, pluck or organ meats, is the organs of a butchered animal. The word does not refer to a particular list of edible organs, which varies by culture and region, but usually excludes muscle. Offal may also refe ...
, and
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
. Many practice
kleptoparasitism Kleptoparasitism (etymologically, parasitism by theft) is a form of feeding in which one animal deliberately takes food from another. The strategy is evolutionarily stable when stealing is less costly than direct feeding, which can mean when foo ...
, which comprises up to 95% of the feeding methods of wintering skuas, by chasing gulls, terns and other seabirds to steal their catches, regardless of the size of the species attacked (up to three times heavier than the attacking skua). The larger species, such as the
great skua The great skua (''Stercorarius skua''), sometimes known by the name bonxie in Britain, is a large seabird in the skua family Stercorariidae. It is roughly the size of a herring gull. It mainly eats fish caught at the sea surface or taken fr ...
, also regularly kill and eat adult birds, such as
puffin Puffins are any of three species of small alcids (auks) in the bird genus ''Fratercula''. These are pelagic seabirds that feed primarily by diving in the water. They breed in large colonies on coastal cliffs or offshore islands, nesting in crev ...
s and
gull Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century, m ...
s, and have been seen killing birds as large as a
grey heron The grey heron (''Ardea cinerea'') is a long-legged wading bird of the heron family, Ardeidae, native throughout temperate Europe and Asia and also parts of Africa. It is resident in much of its range, but some populations from the more northern ...
. On the breeding grounds, the three, more slender northern breeding species commonly eat
lemming A lemming is a small rodent, usually found in or near the Arctic in tundra biomes. Lemmings form the subfamily Arvicolinae (also known as Microtinae) together with voles and muskrats, which form part of the superfamily Muroidea, which also include ...
s. Those species that breed in the southern oceans largely feed on fish that can be caught near their colonies. The eggs and chicks of other seabirds, primarily
penguin Penguins (order (biology), order List of Sphenisciformes by population, Sphenisciformes , family (biology), family Spheniscidae ) are a group of Water bird, aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: on ...
s, are an important food source for most skua species during the nesting season. In the southern oceans and
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
region, some skua species (especially the
south polar skua The south polar skua (''Stercorarius maccormicki'') is a large seabird in the skua family, Stercorariidae. An older name for the bird is MacCormick's skua, after explorer and naval surgeon Robert McCormick, who first collected the type specimen. ...
) will readily scavenge carcasses at breeding colonies of both
penguin Penguins (order (biology), order List of Sphenisciformes by population, Sphenisciformes , family (biology), family Spheniscidae ) are a group of Water bird, aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: on ...
s and
pinniped Pinnipeds (pronounced ), commonly known as seals, are a widely distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammals. They comprise the extant families Odobenidae (whose only living member is the walru ...
s. Skuas will also kill live penguin chicks. In these areas, the skuas will often forfeit their catches to the considerably larger and very aggressive
giant petrel Giant petrels form a genus, ''Macronectes'', from the family Procellariidae, which consists of two species. They are the largest birds of this family. Both species are restricted to the Southern Hemisphere, and though their distributions overlap ...
s. Skuas have also been observed to directly pilfer
milk Milk is a white liquid food produced by the mammary glands of mammals. It is the primary source of nutrition for young mammals (including breastfed human infants) before they are able to digestion, digest solid food. Immune factors and immune ...
from the
elephant seal Elephant seals are very large, oceangoing earless seals in the genus ''Mirounga''. Both species, the northern elephant seal (''M. angustirostris'') and the southern elephant seal (''M. leonina''), were hunted to the brink of extinction for oil ...
's teat. They are medium to large
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s, typically with grey or brown plumage, often with white markings on the wings. The skuas range in size from the
long-tailed skua The long-tailed skua or long-tailed jaeger (''Stercorarius longicaudus'') is a seabird in the skua family Stercorariidae. Etymology The word "jaeger" is derived from the German word ''Jäger'', meaning "hunter". The English word "skua" comes fr ...
, ''Stercorarius longicauda'', at , to the
brown skua The brown skua (''Stercorarius antarcticus''), also known as the Antarctic skua, subantarctic skua, southern great skua, southern skua, or hākoakoa (Māori), is a large seabird that breeds in the subantarctic and Antarctic zones and moves furthe ...
, ''Stercorarius antarcticus'', at . On average, a skua is about long, and across the wings. They have longish bills with a hooked tip, and webbed feet with sharp claws. They look like large dark gulls, but have a fleshy
cere The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food, ...
above the upper mandible. The skuas are strong, acrobatic fliers. They are generally aggressive in disposition. Potential predators approaching their nests will be quickly attacked by the parent birds, which usually target the heads of intruders – a practice known as ' divebombing'.


Taxonomy

The
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Stercorarius'' was introduced by the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published works ...
in 1760 with the
parasitic jaeger The parasitic jaeger (''Stercorarius parasiticus''), also known as the Arctic skua, Arctic jaeger or parasitic skua, is a seabird in the skua family Stercorariidae. It is a migratory species that breeds in Northern Scandinavia, Scotland, Iceland, ...
(''Stercorarius parasiticus'') as the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
. Skuas are related to
gull Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century, m ...
s,
wader 245px, A flock of Dunlins and Red knots">Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflat ...
s,
auk An auk or alcid is a bird of the family Alcidae in the order Charadriiformes. The alcid family includes the murres, guillemots, auklets, puffins, and murrelets. The word "auk" is derived from Icelandic ''álka'', from Old Norse ''alka'' (a ...
s, and skimmers. In the three smaller species, all nesting exclusively in the
Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical region ...
, breeding adults have the two central tail feathers obviously elongated, and at least some adults have white on the underparts and pale yellow on the neck. These characteristics are not shared by the larger species, all native to the Southern Hemisphere except for the great skua. Therefore, the skuas are often split into two genera, with only the smaller species retained in ''Stercorarius'', and the large species placed in ''Catharacta''. However, based on
genetics Genetics is the study of genes, genetic variation, and heredity in organisms.Hartl D, Jones E (2005) It is an important branch in biology because heredity is vital to organisms' evolution. Gregor Mendel, a Moravian Augustinian friar wor ...
, behavior, and
feather lice A bird louse is any chewing louse (small, biting insects) of order Phthiraptera which parasitizes warm-blooded animals, especially birds. Bird lice may feed on feathers, skin, or blood. They have no wings, and their biting mouth parts distingui ...
, the overall relationship among the species is best expressed by placing all in a single genus. The pomarine and great skuas'
mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
(inherited from the mother) is in fact more closely related to each other than it is to either Arctic or long-tailed skuas, or to the Southern Hemisphere species.Cohen, Baker, Belchschmidt, Dittmann, Furness, Gerwin, Helbig, de Korte, Marshall, Palma, Peter, Ramli, Siebold, Willcox, Wilson and Zink (1997)
Enigmatic phylogeny of skuas.
''Proc. Biol. Sci.'' 264(1379):181–190.
Thus,
hybridization Hybridization (or hybridisation) may refer to: *Hybridization (biology), the process of combining different varieties of organisms to create a hybrid *Orbital hybridization, in chemistry, the mixing of atomic orbitals into new hybrid orbitals *Nu ...
must have played a considerable role in the evolution of the diversity of Northern Hemisphere skuas.


Species

The genus contains seven species:


References


External links

*
Skua videos
on the Internet Bird Collection {{Authority control Seabirds Taxa named by Mathurin Jacques Brisson