Szechuanosaurus Campi - IMG 1247
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''Szechuanosaurus'' (" Szechuan lizard") is an extinct Chure, D.J. (2000) ''A new species of'' Allosaurus ''from the Morrison Formation of Dinosaur National Monument (UT–CO) and a revision of the theropod family Allosauridae''. Unpublished Ph.D. dissertation, Columbia University, New York
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of carnivorous
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
. Fossils referred to the genus have been found in
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
,
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
in the Oxfordian-?
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
(Peng ''et al''., 2005). Its
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is largely based on several undiagnostic teeth from the
Shangshaximiao Formation The Shaximiao Formation () is a Middle to Late Jurassic aged geological formation in Sichuan, China, most notable for the wealth of dinosaurs fossils that have been excavated from its strata. The Shaximiao Formation is exposed in and around the ...
and it is possibly also known from the
Kuangyuan Series The Kuangyuan Series is a Late Jurassic ( Oxfordian-?Tithonian) geologic formation in Sichuan (Szechuan), China. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, et al. (2004). "Dinosaur distribution." ...
and the
Kalaza Formation The Kalaza Formation is a geological formation in Xinjiang, China whose strata date back to the Late Jurassic. There is some confusion with the stratigraphy of this unit, as the term is used for sediments of equivalent age in both the Junggar Basin ...
, both also located in China.


Discovery and species

Three species have been assigned to this genus over the years. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''Szechuanosaurus campi'', named by
Yang Zhongjian Yang Zhongjian, also Yang Chung-chien (; 1 June 1897 – 15 January 1979), courtesy name Keqiang (), also known as C.C. (Chung Chien) Young, was a Chinese paleontologist and zoologist. He was one of China's foremost vertebrate paleontologists. H ...
("Chung Chien Young") in 1942 for four isolated tooth specimens: IVPP V235, two partial teeth; IVPP V236, a partial tooth; IVPP V238, several tooth fragments; and IVPP V239, a single tooth. The teeth form a
syntype In biological nomenclature, a syntype is any one of two or more biological types that is listed in a description of a taxon where no holotype was designated. Precise definitions of this and related terms for types have been established as part of ...
series and were not found together. Also a very fragmentary skeleton, specimen UCMP 32102, was referred to the genus by Yang. Some of the teeth indicate a large body-size. These fossils, although possibly
metriacanthosaurid Metriacanthosauridae is an extinct family (biology), family of Allosauroidea, allosauroid theropod dinosaurs that lived from the Middle Jurassic to the Early Cretaceous. When broken down into its Greek roots, it means "moderately-spined lizards". ...
, are now considered to be non-diagnostic, making ''S. campi'' a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a s ...
''.Brusatte, S. L., Benson, R. B. J., and Xu, X. 2010. "The evolution of large-bodied theropod dinosaurs during the Mesozoic in Asia". ''
Journal of Iberian Geology ''Journal of Iberian Geology'' (formerly ''Cuadernos de Geología Ibérica'') is a triannual peer-reviewed scientific journal published by the Universidad Complutense de Madrid. The journal covers the field of geology and related earth sciences ...
'', 36: 275-296
The generic name refers to Szechuan. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
honours the American paleontologist
Charles Lewis Camp Charles Lewis Camp (March 12, 1893 Jamestown, North Dakota – August 14, 1975 San Jose, California) was a palaeontologist and zoologist, working from the University of California, Berkeley. He took part in excavations at the 'Placerias Quarry', in ...
who had discovered UCMP 32102. A partial skeleton, CV 00214, was initially listed by Dong ''et al.'' (1978) in a faunal list as a new species of ''Szechuanosaurus'', ''Szechuanosaurus yandonensis''. At first, there was no description or illustration of it, making ''S. yandonensis'' at the time a ''
nomen nudum In taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate descr ...
''. Later,
Dong Zhiming Dong Zhiming (Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; born January 1937) is a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) in Beijing. He began working at the ...
''et alii'' (1983) described it, and assigned it to ''Szechuanosaurus campi'' The affinities of this skeleton are uncertain, and it has only been briefly described. Holtz ''et al''. (2004) included it in their
phylogenetic analysis In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
and found it to be the most basal
tetanuran Tetanurae (/ˌtɛtəˈnjuːriː/ or "stiff tails") is a clade that includes most theropod dinosaurs, including megalosauroids, allosauroids, tyrannosauroids, ornithomimosaurs, compsognathids and maniraptorans (including birds). Tetanurans ar ...
. This individual was a medium-sized theropod, with an
ischium The ischium () form ...
(a pelvic bone) of ; for comparison, an ischium of ''
Piatnitzkysaurus ''Piatnitzkysaurus'' ( meaning "Piatnitzky's lizard") is a genus of megalosauroid theropod dinosaur that lived approximately 179 to 177 million years ago during the lower part of the Jurassic Period in what is now Argentina. ''Piatnitzkysaurus'' ...
'' estimated to weigh is long. In 2000, Daniel Chure referred the specimen to "
Szechuanoraptor ''Yangchuanosaurus'' is an extinct genus of metriacanthosaurid theropod dinosaur that lived in China from the Middle to Late Jurassic periods (Bathonian to Oxfordian stages), and was similar in size and appearance to its North American and Eur ...
dongi", itself an invalid ''nomen ex dissertatione''. Carrano, Benson & Sampson (2012) synonymized it with ''
Yangchuanosaurus ''Yangchuanosaurus'' is an extinct genus of metriacanthosaurid theropod dinosaur that lived in China from the Middle to Late Jurassic periods (Bathonian to Oxfordian stages), and was similar in size and appearance to its North American and Eur ...
shangyouensis'' from the same formation. The third species is ''Szechuanosaurus zigongensis'', named by Gao Yuhui in 1993 for an almost complete skeleton, specimen ZDM 9011.Gao, Y., 1993, "A new species of ''Szechuanosaurus'' from the Middle Jurassic of Dashanpu, Zigong, Sichuan", ''Vertebrata PalAsiatica'' 31(4): 308-314 It is an older species, from the
Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...
, appearing to be distinct from the type species and therefore requires its own genus name. It was reassigned to ''
Yangchuanosaurus ''Yangchuanosaurus'' is an extinct genus of metriacanthosaurid theropod dinosaur that lived in China from the Middle to Late Jurassic periods (Bathonian to Oxfordian stages), and was similar in size and appearance to its North American and Eur ...
'', as a ''Yangchuanosaurus zigongensis'', by Matthew Carrano, Roger Benson & Scott Sampson in 2012.


Notes


References


''Szechuanosaurus campi'' in The Theropod Database
{{Taxonbar, from=Q637535 Fossil taxa described in 1942 Late Jurassic dinosaurs of Asia Prehistoric neotheropods Paleontology in Sichuan Taxa named by Yang Zhongjian Nomina dubia