Surgeonfish
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Acanthuridae are the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of surgeonfishes, tangs, and unicornfishes. The family includes about 86 extant species of marine fish living in tropical seas, usually around
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. Co ...
s. Many of the species are brightly colored and popular in aquaria.


Morphology

The distinctive characteristic of the family is that they have scalpel-like modified scales, one or more on either side of the peduncle of the tail. The spines are dangerously sharp and may seriously injure anyone who carelessly handles such a fish. The
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal c ...
,
anal Anal may refer to: Related to the anus *Related to the anus of animals: ** Anal fin, in fish anatomy ** Anal vein, in insect anatomy ** Anal scale, in reptile anatomy *Related to the human anus: ** Anal sex, a type of sexual activity involving s ...
, and
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
s are large, extending for most of the length of the body. The mouths are small and have a single row of teeth adapted to grazing on algae. Surgeonfishes sometimes feed as solitary individuals, but they often travel and feed in schools. Feeding in schools may be a mechanism for overwhelming the highly aggressive defense responses of small territorial
damselfish Damselfish are those within the subfamilies Abudefdufinae, Chrominae, Lepidozyginae, Pomacentrinae, and Stegastenae within the family Pomacentridae. Most species within this group are relatively small, with the largest species being about 30 ...
es that vigorously guard small patches of algae on coral reefs. Most species are fairly small, with a maximum length of , but some in the genus ''Acanthurus'', some in the genus ''Prionurus'', and most species in the genus ''Naso'' may grow larger; the
whitemargin unicornfish The whitemargin unicornfish (''Naso annulatus'') is a tropical fish found throughout the Indo-Pacific. It can reach a length of 100 cm, making it one of the largest members of the family Acanthuridae. The species itself exists beyond the Pak ...
(''Naso annulatus'') is the largest species in the family, reaching a length of up to . These fishes may grow quickly in aquaria, so average growth size and suitability should be checked before adding them to any marine aquarium.


Evolution and fossil record

There are several extinct genera known from fossils dating from the
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
to
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
:


Eocene genera

*'' Proacanthurus'' *'' Tylerichthys'' *'' Gazolaichthys'' *''
Naseus ''Naseus'' is an extinct genus of surgeonfishes from the Lutetian-aged Monte Bolca Lagerstätte A Lagerstätte (, from '' Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extr ...
'' *'' Tauichthys'' *'' Eorandallius'' *'' Metacanthurus''


Oligocene genera

*'' Glarithurus'' *''
Caprovesposus ''Caprovesposus parvus'' is an extinct, prehistoric surgeonfish from what is now the Caucasus Mountains. Its very small fossils (up to 3 centimeters in length) are thought to represent a pelagic larval or juvenile stage. The adult form is, as ...
'' *'' Arambourgthurus'' *?'' Eonaso''


Miocene genera

*'' Marosichthys''


Etymology and taxonomic history

The name of the family is derived from the Greek words ''akantha'' and ''oura'', which loosely translate to "thorn" and "tail", respectively. This refers to the distinguishing characteristic of the family, the "
scalpel A scalpel, lancet, or bistoury is a small and extremely sharp bladed instrument used for surgery, anatomical dissection, podiatry and various arts and crafts (either called a hobby knife or an X-acto knife.). Scalpels may be single-use disposa ...
" found on the
caudal peduncle Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
. In the early 1900s, the family was called Hepatidae.


In the aquarium

Tangs are very sensitive to disease in the home aquarium. However, if the tang is fed enough algae and the aquarium is properly maintained disease should not be a problem. It is usually necessary to quarantine the animals for a period before introducing them to the aquarium. Adults range from in length and most grow quickly even in aquaria. When considering a tang for an aquarium it is important to consider the size to which these fish can grow. Larger species such as the popular Pacific blue tang surgeonfish (of ''
Finding Nemo ''Finding Nemo'' is a 2003 American computer-animated comedy-drama adventure film produced by Pixar Animation Studios and released by Walt Disney Pictures. Directed by Andrew Stanton with co-direction by Lee Unkrich, the screenplay was writ ...
'' fame), '' Naso'' or lipstick tang, lined surgeonfish, Sohal surgeonfish and
Atlantic blue tang surgeonfish ''Acanthurus coeruleus'' is a surgeonfish found commonly in the Atlantic Ocean. It can grow up to long.Figueiredo, J.L. and N.A. Menezes, (2000). ''Manual de peixes marinhos do sudeste do Brasil. VI. Teleostei (5)''. Museu de Zoologia, Universid ...
can grow to and require swimming room and hiding places. Many also suggest adding aggressive tangs to the aquarium last as they are territorial and may fight and possibly kill other fish. Tangs primarily graze on
macroalgae Seaweed, or macroalgae, refers to thousands of species of macroscopic, multicellular, marine algae. The term includes some types of '' Rhodophyta'' (red), ''Phaeophyta'' (brown) and ''Chlorophyta'' (green) macroalgae. Seaweed species such as ...
from genera such as ''
Caulerpa ''Caulerpa'' is a genus of seaweeds in the family Caulerpaceae (among the green algae). They are unusual because they consist of only one cell with many nuclei, making them among the biggest single cells in the world. A species in the Mediterran ...
'' and ''
Gracilaria ''Gracilaria'' is a genus of red algae (Rhodophyta) notable for its economic importance as an agarophyte, as well as its use as a food for humans and various species of shellfish. Various species within the genus are cultivated among Asia, South ...
'', although they have been observed in an aquarium setting to eat meat-based fish foods. A popular technique for aquarists, is to grow macroalgae in a
sump A sump is a low space that collects often undesirable liquids such as water or chemicals. A sump can also be an infiltration basin used to manage surface runoff water and recharge underground aquifers. Sump can also refer to an area in a cave ...
or refugium. This technique not only is economically beneficial, but serves to promote enhanced water quality through
nitrate Nitrate is a polyatomic ion A polyatomic ion, also known as a molecular ion, is a covalent bonded set of two or more atoms, or of a metal complex, that can be considered to behave as a single unit and that has a net charge that is not zer ...
absorption. The growth of the algae can then be controlled by feeding it to the tang.


Gallery

File:Acanthurus achilles1.jpg, Achilles tang, ''Acanthurus achilles'' File:Acanthurus dussumieri.jpg, Eyestripe surgeonfish, ''Acanthurus dussumieri'' File:Acanthurusleucosternon-Weißkehl.jpg, Powderblue surgeonfish, ''Acanthurus leucosternon'' File:Acanthurus tennenti Kreisdorn-Doktorfisch3.jpg, Doubleband surgeonfish, ''Acanthurus tennenti'' File:Ctenochaetus strigosus 1.jpg,
Kole tang ''Ctenochaetus strigosus'', known by the common names kole tang or spotted surgeonfish or goldring surgeonfish or yellow-eyed tang, is a marine (ocean), marine reef Tang (fish), tang in the fish Family (biology), family Acanthuridae which is Endem ...
, ''Ctenochaetus strigosus'' File:Naso vlamingii - Masken-Nasendoktorfisch imponierend.jpg, Bignose unicornfish, ''Naso vlamingii'' File:Nasounicornis-Blauklingen.jpg, Bluespine unicornfish, ''Naso unicornis'' File:Zebrasoma desjardinii Indischer Segelflossendoktora.jpg, Red Sea sailfin tang, ''Zebrasoma desjardinii'' File:OrangeBand Surgeonfish.jpg, Orangespot surgeonfish, ''Acanthurus olivaceus'', being cleaned by two
Hawaiian cleaner wrasse The Hawaiian cleaner wrasse or golden cleaner wrasse (''Labroides phthirophagus''), is a species of wrasse (genus '' Labroides'') found in the waters surrounding the Hawaiian Islands. The fish is endemic to Hawaii. These cleaner fish inhabit cor ...
, ''Labroides phthirophagus'' File:Paracanthurus hepatus (Regal Tang).jpg, Regal tang, ''Paracanthurus hepatus'' in an aquarium


References


External links

* * * {{Authority control Perciformes families Marine fish families Taxa named by Charles Lucien Bonaparte Articles which contain graphical timelines