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A stinger (or sting) is a sharp organ found in various
animal Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, ...
s (typically
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s and other
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s) capable of injecting
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
, usually by piercing the
epidermis The epidermis is the outermost of the three layers that comprise the skin, the inner layers being the dermis and Subcutaneous tissue, hypodermis. The epidermal layer provides a barrier to infection from environmental pathogens and regulates the ...
of another animal. An
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
sting is complicated by its introduction of
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
, although not all stings are venomous. Bites, which can introduce saliva as well as additional pathogens and diseases, are often confused with stings, and vice versa. Specific components of venom are believed to give rise to an
allergic reaction Allergies, also known as allergic diseases, are various conditions caused by hypersensitivity of the immune system to typically harmless substances in the environment. These diseases include Allergic rhinitis, hay fever, Food allergy, food al ...
, which in turn produces skin lesions that may vary from a small itching weal, or slightly elevated area of the skin, to large areas of inflamed skin covered by vesicles and crusted lesions. Stinging insects produce a painful swelling of the skin, the severity of the lesion varying according to the location of the sting, the identity of the insect and the sensitivity of the subject. Many species of bees and
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder ...
s have two poison glands, one gland secreting a
toxin A toxin is a naturally occurring poison produced by metabolic activities of living cells or organisms. They occur especially as proteins, often conjugated. The term was first used by organic chemist Ludwig Brieger (1849–1919), derived ...
in which formic acid is one recognized constituent, and the other secreting an
alkaline In chemistry, an alkali (; from the Arabic word , ) is a basic salt of an alkali metal or an alkaline earth metal. An alkali can also be defined as a base that dissolves in water. A solution of a soluble base has a pH greater than 7.0. The ...
neurotoxin; acting independently, each toxin is rather mild, but when they combine through the sting, the combination has strong irritating properties. In a small number of cases, the second occasion of a bee or wasp sting causes a severe allergic reaction known as anaphylaxis. While the overwhelming majority of insects withdraw their stingers from their victims, a few insects leave them in the wounds. For example, of the 20,000 species of bees worldwide, only the half-dozen species of honeybees ('' Apis'') are reported to have a barbed stinger that cannot be withdrawn; of wasps, nearly all are reported to have smooth stingers with the exception of two species, '' Polybia rejecta'' and '' Synoeca surinama''. A few non-insect arthropods, such as
scorpion Scorpions are predatory arachnids of the Order (biology), order Scorpiones. They have eight legs and are easily recognized by a pair of Chela (organ), grasping pincers and a narrow, segmented tail, often carried in a characteristic forward cur ...
s, also sting.


Arthropods

Among
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s, a sting or stinger is a sharp organ, often connected with a venom gland and adapted to inflict a wound by piercing, as with the caudal sting of a
scorpion Scorpions are predatory arachnids of the Order (biology), order Scorpiones. They have eight legs and are easily recognized by a pair of Chela (organ), grasping pincers and a narrow, segmented tail, often carried in a characteristic forward cur ...
. Stings are usually located at the rear of the animal. Animals with stings include bees, wasps (including
hornet Hornets (insects in the genus ''Vespa'') are the largest of the Eusociality, eusocial wasps, and are similar in appearance to yellowjackets, their close relatives. Some species can reach up to in length. They are distinguished from other Vespi ...
s), some
ant Ants are Eusociality, eusocial insects of the Family (biology), family Formicidae and, along with the related wasps and bees, belong to the Taxonomy (biology), order Hymenoptera. Ants evolved from Vespoidea, vespoid wasp ancestors in the Cre ...
s like fire ants, and
scorpion Scorpions are predatory arachnids of the Order (biology), order Scorpiones. They have eight legs and are easily recognized by a pair of Chela (organ), grasping pincers and a narrow, segmented tail, often carried in a characteristic forward cur ...
s, as well as a single beetle species ('' Onychocerus albitarsis'') that can deliver a venomous sting from its antennae, whose terminal segments have evolved to resemble a scorpion's tail. In all stinging Hymenoptera the sting is a modified ovipositor. Unlike most other stings,
honey bee A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to mainland Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the ...
workers' stings are strongly barbed and lodge in the flesh of
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s upon use, tearing free from the honey bee's body, killing the bee within minutes. The sting has its own ganglion, and it continues to saw into the target's flesh and release venom for several minutes. This trait is of obvious disadvantage to the individual but protects the hive from attacks by large animals; aside from the effects of the venom, the remnant also marks the stung animal with honey bee alarm pheromone. The barbs of a honey bee's attack are only suicidal if the skin is elastic, as is characteristic of vertebrates such as birds and mammals; honey bees can sting other insects repeatedly without dying. The sting of nearly all other bees and other sting-bearing organisms is not barbed and can be used to sting repeatedly. The description of barbed or unbarbed is not precise: there are barbs on the stings of yellowjacket wasps and the Mexican honey wasp, but the barbs are so small that the wasp can sometimes withdraw its sting apparatus from victim's skin. The stings of some wasps, such as those of the '' Polistes versicolor'', contain relatively large amounts of 5-hydroxytryptamine (5-HT) in its venoms. The 5-HT in these venoms has been found to play at least two roles: one as a pain-producing agent and the other in the distribution and penetration of the paralyzing components to vulnerable sites in the offender. This helps in the rapid immobilization of the animal or of the body parts receiving the venom. Spiders only bite, although some tarantulas have barbed bristles called urticating hairs. Certain caterpillars also have urticating hairs.
Centipede Centipedes (from Neo-Latin , "hundred", and Latin , "foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', "lip", and Neo-Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, ...
s also possess a venomous bite rather than a sting, inflicted with a highly modified first pair of legs, called forcipules.


Stingrays, platypuses and jellyfish

Organs that perform similar functions in non-arthropods are often referred to as "stings". These organs include the modified dermal denticle of the stingray, the venomous spurs on the hind legs of the male platypus, and the cnidocyte tentacles of the
jellyfish Jellyfish, also known as sea jellies or simply jellies, are the #Life cycle, medusa-phase of certain gelatinous members of the subphylum Medusozoa, which is a major part of the phylum Cnidaria. Jellyfish are mainly free-swimming marine animal ...
.


See also

* Bee sting * Cnidocyte * Chelicerae * Forcipule * Insect bite * Schmidt Sting Pain Index * Starr sting pain scale * Stinging plant


References


External links

*{{Commons category-inline, Stingers Animal anatomy