The stapedius is the smallest
skeletal muscle
Skeletal muscle (commonly referred to as muscle) is one of the three types of vertebrate muscle tissue, the others being cardiac muscle and smooth muscle. They are part of the somatic nervous system, voluntary muscular system and typically are a ...
in the
human
Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...
body.
At just over one millimeter in length, its purpose is to stabilize the smallest bone in the body, the
stapes
The ''stapes'' or stirrup is a bone in the middle ear of humans and other tetrapods which is involved in the conduction of sound vibrations to the inner ear. This bone is connected to the oval window by its annular ligament, which allows the f ...
or stirrup bone of the middle ear.
Structure
The stapedius emerges from a pinpoint
foramen
In anatomy and osteology, a foramen (; : foramina, or foramens ; ) is an opening or enclosed gap within the dense connective tissue (bones and deep fasciae) of extant and extinct amniote animals, typically to allow passage of nerves, artery, ...
or opening in the apex of the
pyramidal eminence
The pyramidal eminence is a hollow conical projection upon the posterior wall of the tympanic cavity of the middle ear. The stapedius muscle arises in the hollow of the eminence and its tendon exits through its apex.
The pyramidal eminence is si ...
(a hollow, cone-shaped prominence in the posterior wall of the
tympanic cavity
The tympanic cavity is a small cavity surrounding the bones of the middle ear. Within it sit the ossicles, three small bones that transmit vibrations used in the detection of sound.
Structure
On its lateral surface, it abuts the external audit ...
), and inserts into the neck of the stapes.
Nerve supply
The stapedius is supplied by the
nerve to stapedius
The nerve to the stapedius is a branch of the facial nerve (CN VII) which innervates the stapedius muscle. It arises from the CN VII within the facial canal, opposite the pyramidal eminence. It passes through a small canal in this eminence to re ...
, a branch of the
facial nerve
The facial nerve, also known as the seventh cranial nerve, cranial nerve VII, or simply CN VII, is a cranial nerve that emerges from the pons of the brainstem, controls the muscles of facial expression, and functions in the conveyance of ta ...
.
[
]
Function
The stapedius dampens the vibrations of the stapes by pulling on the neck of that bone. As one of the muscles involved in the acoustic reflex
The acoustic reflex (also known as the stapedius reflex, stapedial reflex, auditory reflex, middle-ear-muscle reflex (MEM reflex, MEMR), attenuation reflex, cochleostapedial reflex or intra-aural reflex) is an involuntary muscle contraction that ...
it prevents excess movement of the stapes
The ''stapes'' or stirrup is a bone in the middle ear of humans and other tetrapods which is involved in the conduction of sound vibrations to the inner ear. This bone is connected to the oval window by its annular ligament, which allows the f ...
, helping to control the amplitude of sound waves from the general external environment to the inner ear.
Clinical significance
Paralysis
Paralysis (: paralyses; also known as plegia) is a loss of Motor skill, motor function in one or more Skeletal muscle, muscles. Paralysis can also be accompanied by a loss of feeling (sensory loss) in the affected area if there is sensory d ...
of the stapedius allows wider oscillation
Oscillation is the repetitive or periodic variation, typically in time, of some measure about a central value (often a point of equilibrium) or between two or more different states. Familiar examples of oscillation include a swinging pendulum ...
of the stapes, resulting in heightened reaction of the auditory ossicles to sound vibration. This condition, known as hyperacusis
Hyperacusis is an increased Hearing, sensitivity to sound and a low tolerance for environmental noise. Definitions of hyperacusis can vary significantly; it often revolves around damage to or dysfunction of the Stapes, ''stapes'' bone, stapedius ...
, causes normal sounds to be perceived as very loud. Paralysis of the stapedius muscle may result when the nerve to the stapedius
The nerve to the stapedius is a branch of the facial nerve (CN VII) which innervates the stapedius muscle. It arises from the CN VII within the facial canal, opposite the pyramidal eminence. It passes through a small canal in this eminence to rea ...
, a branch of the facial nerve
The facial nerve, also known as the seventh cranial nerve, cranial nerve VII, or simply CN VII, is a cranial nerve that emerges from the pons of the brainstem, controls the muscles of facial expression, and functions in the conveyance of ta ...
, is damaged, or when the facial nerve itself is damaged before the nerve to stapedius branches. In cases of Bell's palsy
Bell's palsy is a type of facial paralysis that results in a temporary inability to control the facial muscles on the affected side of the face. In most cases, the weakness is temporary and significantly improves over weeks. Symptoms can vary f ...
, a unilateral paralysis of the facial nerve, the stapedius is paralyzed and hyperacusis may result.
Other animals
Like the stapes
The ''stapes'' or stirrup is a bone in the middle ear of humans and other tetrapods which is involved in the conduction of sound vibrations to the inner ear. This bone is connected to the oval window by its annular ligament, which allows the f ...
bone to which it attaches, the stapedius muscle shares evolutionary history with other vertebrate structures.
The mammalian stapedius evolved from a muscle called the depressor mandibulae in other tetrapods, the function of which was to open the jaws (this function was taken over by the digastric muscle
The digastric muscle (also digastricus) (named ''digastric'' as it has two 'bellies') is a bilaterally paired suprahyoid muscle located under the jaw. Its posterior belly is attached to the mastoid notch of temporal bone, and its anterior belly ...
in mammals). The depressor mandibulae arose from the levator operculi Levator muscle can refer to:
* Levator scapulae muscle
* Levator palpebrae superioris muscle
* Levator ani
The levator ani is a broad, thin muscle group, situated on either side of the pelvis. It is formed from three muscle components: the puboc ...
in bony fish, and is equivalent to the epihyoidean in sharks. Like the stapedius, all of these muscles derive from the hyoid arch
The pharyngeal arches, also known as visceral arches'','' are transient structures seen in the embryonic development of humans and other vertebrates, that are recognisable precursors for many structures. In fish, the arches support the gills a ...
and are innervated by cranial nerve VII
The facial nerve, also known as the seventh cranial nerve, cranial nerve VII, or simply CN VII, is a cranial nerve that emerges from the pons of the brainstem, controls the muscles of facial expression, and functions in the conveyance of tast ...
.
See also
* Hearing
Hearing, or auditory perception, is the ability to perceive sounds through an organ, such as an ear, by detecting vibrations as periodic changes in the pressure of a surrounding medium. The academic field concerned with hearing is auditory sci ...
* Middle ear
The middle ear is the portion of the ear medial to the eardrum, and distal to the oval window of the cochlea (of the inner ear).
The mammalian middle ear contains three ossicles (malleus, incus, and stapes), which transfer the vibrations ...
* Ossicles
The ossicles (also called auditory ossicles) are three irregular bones in the middle ear of humans and other mammals, and are among the smallest bones in the human body. Although the term "ossicle" literally means "tiny bone" (from Latin ''ossi ...
* Tensor tympani
The tensor tympani is a muscle within the middle ear, located in the bony canal above the bony part of the auditory tube, and connects to the malleus bone. Its role is to dampen loud sounds, such as those produced from chewing, shouting, or thun ...
the other major muscle in the middle ear
* Stapes
The ''stapes'' or stirrup is a bone in the middle ear of humans and other tetrapods which is involved in the conduction of sound vibrations to the inner ear. This bone is connected to the oval window by its annular ligament, which allows the f ...
the other bone to which the muscle attaches
References
Rodríguez-Vázquez JF. Development of the stapedius muscle and pyramidal eminence in humans. J Anat. 2009 Sep;215(3):292-9. doi: 10.1111/j.1469-7580.2009.01105.x
External links
{{DEFAULTSORT:Stapedius Muscle
Muscles of the head and neck
Otorhinolaryngology
Auditory system