''Craspedocephalus trigonocephalus'', the Sri Lankan pit viper, Ceylon pit viper, Sri Lankan green pitviper or locally, pala polonga, ( si, පළා පොළඟා) is a

venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...

pit viper

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...

to Sri Lanka. No subspecies are currently recognized.

Description

''Craspedocephalus trigonocephalus'' is a

sexually dimorphic Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...

, mid-sized, cylindrical species. These snakes measure at birth, and males grow to a maximum total length of . The neck is distinct from the flattened, triangular head. There is a loreal pit on each side of the head. Mid-sized eyes and a short, rounded, broad snout are present. The males are considerably smaller than females, which can grow up to in total length. The ground color of the snake is variable and cryptic. Typically, it is a green snake with a black variegated pattern, and a black temporal line is usually present. The wet zone snakes have these black patterns more clearly marked. The vertebral area has a tinge of yellow. The tail is black. The

ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...

are light greenish-yellow or may even be grey. Some snakes have only the black temporal line and the black tail, with the rest of the body being green. In addition, males tend to have a blue coloration, whereas the females are predominantly green. These are bulky snakes with

prehensile Prehensility is the quality of an appendage or organ that has adapted for grasping or holding. The word is derived from the Latin term ''prehendere'', meaning "to grasp". The ability to grasp is likely derived from a number of different orig ...

, short tails, suiting their

arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose nu ...

lifestyle. Sri Lanka Green Pit Viper head detail

Reproduction

These are

viviparous Among animals, viviparity is development of the embryo inside the body of the parent. This is opposed to oviparity which is a reproductive mode in which females lay developing eggs that complete their development and hatch externally from the ...

snakes. About five to 25 young are produced at once.

Scales

These snakes have two or three large supra

ocular scales In scaled reptiles, the ocular scales are those forming the margin of the eye.Wright AH, Wright AA. 1957. ''Handbook of Snakes''. Comstock Publishing Associates (7th printing, 1985). 1105 pp. . The name originates from the term ''oculus'' which ...

, and their

nasal scale In reptiles, the nasal scale refers to the scale that encloses the nostril.Mallow D, Ludwig D, Nilson G. 2003. True Vipers: Natural History and Toxinology of Old World Vipers. Malabar, Florida: Krieger Publishing Company. 359 pp. . Sometimes this ...

s are partially divided into two or may not be divided. They have three preoculars, two or three postoculars, 9–10 supralabial scales, three or four

infralabials In reptiles, the sublabial scales, also called lower-labials or infralabials, are those scales that border the mouth opening along the lower jaw. They do not include the median scaleWright AH, Wright AA. 1957. ''Handbook of Snakes''. Comstock Publis ...

, and 142–160 ventrals; The

anal scale Anal may refer to: Related to the anus *Related to the anus of animals: ** Anal fin, in fish anatomy ** Anal vein, in insect anatomy ** Anal scale, in reptile anatomy *Related to the human anus: ** Anal sex, a type of sexual activity involv ...

is not divided; their 53–69

subcaudals In snakes, the subcaudal scales are the enlarged plates on the underside of the tail.Wright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates (7th printing, 1985). 1105 pp. . These scales may be either single or divided (pair ...

are divided. At midbody, the 17–19 rows of

dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down t ...

may or may not have keels. Most head shields are small and smooth.

Common names

Common names include: Sri Lankan green pit viper,Gumprecht A, Tillack F, Orlov NL, Captain A, Ryabov S. 2004. ''Asian Pitvipers''. Geitje Books. Berlin. 1st Edition. 368 pp. . Sri Lankan pit viper,Mehrtens JM. 1987. ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . ''pala polonga'' (පළා පොළඟා), and green pit viper.

Geographic range

It is an endemic species to Sri Lanka and widely distributed in all three

climatic zone Climate classifications are systems that categorize the world's climates. A climate classification may correlate closely with a biome classification, as climate is a major influence on life in a region. One of the most used is the Köppen climate ...

s of the island, except higher hills and

arid A region is arid when it severely lacks available water, to the extent of hindering or preventing the growth and development of plant and animal life. Regions with arid climates tend to lack vegetation and are called xeric or desertic. Most ...

zones, while relatively more common in wet zone grasslands and

rain forest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainforest ...

areas and occasionally in

plantations A plantation is an agricultural estate, generally centered on a plantation house, meant for farming that specializes in cash crops, usually mainly planted with a single crop, with perhaps ancillary areas for vegetables for eating and so on. Th ...

cardamom Cardamom (), sometimes cardamon or cardamum, is a spice made from the seeds of several plants in the genera ''Elettaria'' and ''Amomum'' in the family Zingiberaceae. Both genera are native to the Indian subcontinent and Indonesia. They are r ...

, cocoa,

coffee Coffee is a drink prepared from roasted coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulating effect on humans, primarily due to its caffeine content. It is the most popular hot drink in the world. Seeds of ...

, and

tea Tea is an aromatic beverage prepared by pouring hot or boiling water over cured or fresh leaves of ''Camellia sinensis'', an evergreen shrub native to East Asia which probably originated in the borderlands of southwestern China and north ...

, from the lower altitudes from .de Silva, 1980 The type locality given is "l'île S.-Eustache" (Sri Lanka).

Behavior

It is

and nocturnal, occasionally descending to the ground in search of food such as lizards, frogs, small mammals, and birds. This sluggish pit viper is usually encountered on low shrubs during morning hours, but it mostly occupies in grasslands and rain forests. In mornings, it is seen to stay on top of trees to obtain sun rays to heat its body. It uses its tail to hold on to a tree branch. This is not a particularly defensive species, but if agitated, it will vibrate its tail tip, form a sinuous loop with the fore body, and lash and attempt to bite, seldom with a hissing sound. It produces live young mostly during June and July.

Venom

The venom is primarily haemotoxic, with victims experiencing severe pain and swelling of the bitten area,

oedema Edema, also spelled oedema, and also known as fluid retention, dropsy, hydropsy and swelling, is the build-up of fluid in the body's tissue. Most commonly, the legs or arms are affected. Symptoms may include skin which feels tight, the area ma ...

and blisters, and localised tissue necrosis; the pain of the wound may last for a few days. Ptosis and lymphadenopathy takes place. Also in some victims, polyuric renal failure and cardiac electrophysiological dysfunction occur, but fatalities have not been reported.