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''Spathiostemon javensis'' is a plant that can grow as a shrub or a tree in the tribe
Acalypheae The Acalypheae is a tribe of the subfamily Acalyphoideae, under the family Euphorbiaceae. It comprises 12 subtribes and 32 genera. Genera See also *Taxonomy of the Euphorbiaceae Here is a full taxonomy of the family Euphorbiaceae, accordin ...
of the family
Euphorbiaceae Euphorbiaceae, the spurge family, is a large family of flowering plants. In English, they are also commonly called euphorbias, which is also the name of a genus in the family. Most spurges, such as ''Euphorbia paralias'', are herbs, but some, e ...
. It is native to the region from the
Bismarck Archipelago The Bismarck Archipelago (, ) is a group of islands off the northeastern coast of New Guinea in the western Pacific Ocean and is part of the Islands Region of Papua New Guinea. Its area is about 50,000 square km. History The first inhabitants o ...
to
New Guinea New Guinea (; Hiri Motu Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea). It is a simplified version of ...
,
Wallacea Wallacea is a biogeographical designation for a group of mainly Indonesian islands separated by deep-water straits from the Asian and Australian continental shelves. Wallacea includes Sulawesi, the largest island in the group, as well as Lo ...
and into Southeast Asia. It is often common in the understorey of forests. The wood is used in constructions.


Description

The species can grow as a shrub up to a tree some 20m high, with a
diameter at breast height Diameter at breast height, or DBH, is a standard method of expressing the diameter of the trunk or bole of a standing tree. DBH is one of the most common dendrometric measurements. Tree trunks are measured at the height of an adult's breast, ...
up to 45 cm, seldom with
buttresses A buttress is an architectural structure built against or projecting from a wall which serves to support or reinforce the wall. Buttresses are fairly common on more ancient buildings, as a means of providing support to act against the lateral (si ...
up to a metre high and long, some 30 cm thick. The bark is smooth and fluted, peels off in scales and strips, its colour varies through white, grey, white mottled, dark red, pale brown, dark brown to black. The wood varies from soft but is usually hard, the sapwood is white to brownish, while the heartwood is brownish. The leaves are elliptic to somewhat obovate in
shape A shape or figure is a graphics, graphical representation of an object or its external boundary, outline, or external Surface (mathematics), surface, as opposed to other properties such as color, Surface texture, texture, or material type. A pl ...
, about 4.3-24 x 1.7-11 cm in size, smooth and glabrous, though perhaps some hairs on the lower side midrib. Flower colour from green, white, yellow to brown. Fruit is some 1-1.2 x 0.6-0.7 cm in size, and pink-brownish to reddish and yellow, with hairs. Flowering and fruiting occur all year round. ''Spathiostemon javensis'' is distinguished from its only sister species ('' Spathiostemon moniliformis'') by having: subhirsute petioles; hair tuft
domatia A domatium (plural: domatia, from the Latin "domus", meaning home) is a tiny chamber that houses arthropods, produced by a plant. Ideally domatia differ from galls in that they are produced by the plant rather than being induced by their inhabi ...
usually on leaves; the inflorescences are subhirsute, while the
staminate The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
inflorescences are up to 7.3 cm long; there are 5
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
s of
pistil Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
late flowers; and the ovary and fruit are echinate (have spines).


Habitat, ecology

The tree is often common in the understorey of primary and secondary forests in shrublands, on cliffs, river-edges and even in and alongside plantations of rubber and cocoa. It favours flat to undulating country, with ability to grow in sediments that are dry to periodically inundated. Often grows in limestone soils but also alluvial, black/brown clay, clay-loam and sand. Occurs from sea level up to 670m elevation. In the
Buol Regency Buol Regency ( id, Kabupaten Buol) is a regency (second-level administrative division) of Central Sulawesi Province of Indonesia. It covers an area of 4,043.57 km2, and had a population of 132,330 at the 2010 Census; and 145,254 (comprising 74,624 ...
, central Sulawesi, the tree occurs in both undisturbed and disturbed forest. In the south of West Papua, on the border with Papua Niugini, small ''Spathistemon javensis'' trees are positively associated with ''
Pometia pinnata ''Pometia pinnata'' is a large tropical hardwood and fruit tree species, with common names including matoa, taun tree, island lychee, tava, Pacific lychee of the plant family Sapindaceae. Naturally widespread, the trees are native to tropical So ...
'', a targeted logging species. At Aipiri,
Manokwari Manokwari is a coastal town and the capital of the Indonesian province of West Papua. It is one of only seven provincial capitals of Indonesia without a city status. It is also the administrative seat of Manokwari Regency. However, under pro ...
, West Papua, the tree did not grow on the coast, but dominated the forest 600m inland, along with ''
Horsfieldia irya ''Horsfieldia irya'' is a species of plant in the family Myristicaceae. It is found in Burma, India, Malaysia, Papua New Guinea, Singapore, Solomon Islands, Sri Lanka, Thailand, and Vietnam. The plant is a dioecious tree that grows to about 8– ...
'' and ''
Myristica fatua ''Myristica'' is a genus of trees in the family Myristicaceae. There are over 150 species, distributed in Asia and the western Pacific. The type species of the genus, and the most economically important member, is ''Myristica fragrans'' (the nut ...
''. At another site in Manokwari, the species was highly associated with natural ''
Intsia bijuga ''Intsia bijuga'', commonly known as Borneo teak, Johnstone River teak, Kwila, Moluccan ironwood, Pacific teak, scrub mahogany and vesi, is a species of flowering tree in the family Fabaceae, native to the Indo-Pacific. It ranges from Tanzania and ...
'' stands. The tallest, most impressive forests on
Waigeo Waigeo is an island in Southwest Papua province of eastern Indonesia. The island is also known as Amberi, or Waigiu. It is the largest of the four main islands in the Raja Ampat Islands Raja Ampat, or the ''Four Kings'', is an archipelago loc ...
of the
Raja Ampat Islands Raja Ampat, or the ''Four Kings'', is an archipelago located off the northwest tip of Bird's Head Peninsula on the island of New Guinea, in Indonesia's Southwest Papua province. It comprises over 1,500 small islands, cays, and shoals surrounding ...
, eastern Indonesia, contain the species as a co-dominant, along with ''
Hopea novoguineensis ''Hopea novoguineensis'' is a rainforest tree species in the Dipterocarpaceae family. It is found in New Guinea and on Halmahera, Maluku Province, Indonesia. The tree produces good timber. Description The species is a medium-sized tree with t ...
'', ''
Homalium foetidum ''Homalium foetidum'' is a species of plant in the family Salicaceae. It is found in Indonesia, Malaysia, Papua New Guinea, and the Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fi ...
'', ''
Mallotus floribundus ''Mallotus floribundus'' is a tree in the family Euphorbiaceae, in the Stylanthus section, native to Southeast Asia, Wallaceae, New Guinea and the Solomon Islands. Description The species grows a small tree, up to 18m, occasionally 25m, tall. It ...
'', ''
Pimelodendron amboinicum ''Pimelodendron amboinicum'' is a tree species in the Euphorbiaceae family. It is found from the Solomon Islands in the southwest Pacific, west to Sulawesi in Indonesia. The timber is used locally, though larger-scale illegal logging is apparent. ...
'', and ''
Vatica rassak ''Vatica rassak'' is a tree in the family Dipterocarpaceae, native to Maritime Southeast Asia. The specific epithet ''rassak'' is from , the species' Malay common name. Description ''Vatica rassak'' grows up to tall, with a trunk diameter of ...
'', with ''
Elatostema ''Elatostema'' is a genus of flowering plants containing approximately 350 known species in the nettle family Urticaceae, native to tropical forest clearings throughout Australasia, Asia and Africa. There may be as many as 1,000 species of this l ...
'' and ''
Piper Piper may refer to: People * Piper (given name) * Piper (surname) Arts and entertainment Fictional characters Comics * Piper (Morlock), in the Marvel Universe * Piper (Mutate), in the Marvel Universe Television * Piper Chapman, lea ...
'' quite common in the understorey. This forest grows on alluvial and residual sandstone and volcanic soils. The plants are spread by fruit-eating birds and mammals.


Distribution

The tree/shrub is endemic to Malesia, and is found from the Bismarck Archipelago across to parts of Southeast Asia. Countries and regions that it is found in are:
Papua New Guinea Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country i ...
(Bismarck Archipelago, mainland); Indonesia ( Maluku,
Nusa Tenggara The Lesser Sunda Islands or nowadays known as Nusa Tenggara Islands ( id, Kepulauan Nusa Tenggara, formerly ) are an archipelago in Maritime Southeast Asia, north of Australia. Together with the Greater Sunda Islands to the west they make up t ...
,
Sulawesi Sulawesi (), also known as Celebes (), is an island in Indonesia. One of the four Greater Sunda Islands, and the world's eleventh-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Mindanao and the Sulu Ar ...
,
Kalimantan Kalimantan () is the Indonesian portion of the island of Borneo. It constitutes 73% of the island's area. The non-Indonesian parts of Borneo are Brunei and East Malaysia. In Indonesia, "Kalimantan" refers to the whole island of Borneo. In 2019, ...
; Jawa, not present in
Sumatera Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
); Philippines (except
Luzon Luzon (; ) is the largest and most populous island in the Philippines. Located in the northern portion of the Philippines archipelago, it is the economic and political center of the nation, being home to the country's capital city, Manila, as ...
); and Malaysia (
Sabah Sabah () is a States and federal territories of Malaysia, state of Malaysia located in northern Borneo, in the region of East Malaysia. Sabah borders the Malaysian state of Sarawak to the southwest and the North Kalimantan province of Indone ...
where it is particularly common,
Sarawak Sarawak (; ) is a States and federal territories of Malaysia, state of Malaysia. The largest among the 13 states, with an area almost equal to that of Peninsular Malaysia, Sarawak is located in northwest Borneo Island, and is bordered by the M ...
, and
Perak Perak () is a state of Malaysia on the west coast of the Malay Peninsula. Perak has land borders with the Malaysian states of Kedah to the north, Penang to the northwest, Kelantan and Pahang to the east, and Selangor to the south. Thailand's ...
).


Conservation

As stated above the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
regards the conservation this taxa as of least concern. This is because there is a large population of the trees over a very wide distribution, with no obvious current or future threats.


Vernacular names

The plant, with a wide and common distribution, has many names: Papua New Guinea: , , , , , , , , , , , ( Amele); , , , , , , ( Biliau); ( Bogia); , , , , , , , , , ( Dumpu); , , , , , , , , ( Faita); ''sarr'' (Jal); West Papua: ( Kemtuik/Kemtoek); ( Sentani); ( Berik); (
Biak Biak is an island located in Cenderawasih Bay near the northern coast of Papua (province), Papua, an Indonesian province, and is just northwest of New Guinea. Biak is the largest island in its small archipelago, and has many atolls, reefs, and c ...
); ( Numfor/Numfur); ( Iria/Irian?); , , ( Sougb/Manikiong); ( Irarutu/Iraroetoe); (
Wain A wagon or waggon is a heavy four-wheeled vehicle pulled by draught animals or on occasion by humans, used for transporting goods, commodities, agricultural materials, supplies and sometimes people. Wagons are immediately distinguished from ...
); ''boeboeika'', ''menom'';
Halmahera Halmahera, formerly known as Jilolo, Gilolo, or Jailolo, is the largest island in the Maluku Islands. It is part of the North Maluku province of Indonesia, and Sofifi, the capital of the province, is located on the west coast of the island. Hal ...
: ( Tabaru/Tobaro); Philippines: (
Manobo The Lumad are a group of Austronesian indigenous people in the southern Philippines. It is a Cebuano term meaning "native" or "indigenous". The term is short for Katawhang Lumad (Literally: "indigenous people"), the autonym officially adopte ...
); ( Waray/Waray-waray); Sabah: ( Bonggi/Dusun Banggi); , , ( Eastern Kadazan/Dusun Kinabatangan); ( Lun Bawang/Murud); ( Sungai/Sungei); ''lengkan'' (K.); Sarawak: (
Iban IBAN or Iban or Ibán may refer to: Banking * International Bank Account Number Ethnology * Iban culture * Iban language * Iban people Given name Cycling * Iban Iriondo (born 1984) * Iban Mayo (born 1977) * Iban Mayoz (born 1981) Football * ...
); Kalimantan: ''wajan koreng''.


Uses

The wood of this species is used for constructions in sea-water in Halmahera, Indonesia, and in the Philippines for general construction. Indonesian sources describe the wood as being heavy, of strength class II (fairly strong), durability class V , and traditionally used for house pillars and bridges.tionally used as house pillars and bridges. It was identified as having a high potential as a shade-tree in merbau (Intsia bijuga, an important global commercial timber) plantations.


History

The
Braunschweig Braunschweig () or Brunswick ( , from Low German ''Brunswiek'' , Braunschweig dialect: ''Bronswiek'') is a city in Lower Saxony, Germany, north of the Harz Mountains at the farthest navigable point of the river Oker, which connects it to the Nor ...
born botanist
Carl Ludwig Blume Charles Ludwig de Blume or Karl Ludwig von Blume (9 June 1796, Braunschweig – 3 February 1862, Leiden) was a German-Dutch botanist. He was born at Braunschweig in Germany, but studied at Leiden University and spent his professional life wor ...
(1796-1862), described this species in 1826 in his publication ''Bijdragen tot de Flora van Nederlandsch Indie''.


References


Further reading

*Govaerts, R., Frodin, D.G. & Radcliffe-Smith, A. (2000). World Checklist and Bibliography of Euphorbiaceae (and Pandaceae) 1-4: 1–1622. The Board of Trustees of the Royal Botanic Gardens, Kew. * {{Taxonbar, from=Q15381941 javensis Flora of Malesia Flora of Papuasia Plants described in 1826