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botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek w ...
, a bract is a modified or specialized
leaf A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ste ...
, especially one associated with a reproductive structure such as a
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
,
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
axis or
cone scale A conifer cone (in formal botanical usage: strobilus, plural strobili) is a seed-bearing organ on gymnosperm plants. It is usually woody, ovoid to globular, including scales and bracts arranged around a central axis, especially in conifers an ...
. Bracts are usually different from foliage leaves. They may be smaller, larger, or of a different color, shape, or texture. Typically, they also look different from the parts of the flower, such as the petals or
sepals A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
. A plant having bracts is referred to as bracteate or bracteolate, while one that lacks them is referred to as ebracteate and ebracteolate, without bracts.


Variants

Some bracts are brightly-coloured and serve the function of attracting pollinators, either together with the
perianth The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower, and structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepals when ...
or instead of it. Examples of this type of bract include those of ''
Euphorbia pulcherrima The poinsettia ( or ) (''Euphorbia pulcherrima'') is a commercially important flowering plant species of the diverse spurge family Euphorbiaceae. Indigenous to Mexico and Central America, the poinsettia was first described by Europeans in 1834 ...
'' (poinsettia) and ''
Bougainvillea ''Bougainvillea'' ( , ) is a genus of thorny ornamental vines, bushes, and trees belonging to the four o' clock family, Nyctaginaceae. It is native to eastern South America, found from Brazil, west to Peru, and south to southern Argentina. ...
'': both of these have large colourful bracts surrounding much smaller, less colourful flowers. In
grasses Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns and ...
, each floret (flower) is enclosed in a pair of papery bracts, called the lemma (lower bract) and palea (upper bract), while each spikelet (group of florets) has a further pair of bracts at its base called glumes. These bracts form the
chaff Chaff (; ) is the dry, scaly protective casing of the seeds of cereal grains or similar fine, dry, scaly plant material (such as scaly parts of flowers or finely chopped straw). Chaff is indigestible by humans, but livestock can eat it. In agri ...
removed from
cereal A cereal is any Poaceae, grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, Cereal germ, germ, and bran. Cereal Grain, grain crops are grown in greater quantit ...
grain during
threshing Threshing, or thrashing, is the process of loosening the edible part of grain (or other crop) from the straw to which it is attached. It is the step in grain preparation after reaping. Threshing does not remove the bran from the grain. History ...
and
winnowing Winnowing is a process by which chaff is separated from grain. It can also be used to remove pests from stored grain. Winnowing usually follows threshing in grain preparation. In its simplest form, it involves throwing the mixture into the ...
. Bats may detect acoustic signals from dish-shaped bracts such as those of ''
Marcgravia evenia ''Marcgravia evenia'' is a species of flowering vine in the family Marcgraviaceae. Within this family it belongs to the Galetae group, which is characterized by a long inflorescence axis and boat shaped nectaries. The plant is endemism, endemic t ...
''. A prophyll is a leaf-like structure, such as a bracteole, subtending (extending under) a single flower or
pedicel Pedicle or pedicel may refer to: Human anatomy *Pedicle of vertebral arch, the segment between the transverse process and the vertebral body, and is often used as a radiographic marker and entry point in vertebroplasty and kyphoplasty procedures ...
. The term can also mean the lower bract on a peduncle. The frequently showy pair of bracts of ''
Euphorbia ''Euphorbia'' is a very large and diverse genus of flowering plants, commonly called spurge, in the family Euphorbiaceae. "Euphorbia" is sometimes used in ordinary English to collectively refer to all members of Euphorbiaceae (in deference to t ...
'' species in subgenus ''Lacanthis'' are the cyathophylls. Bracts subtend the cone scales in the seed cones of many
conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
s, and in some cases, such as ''
Pseudotsuga ''Pseudotsuga'' is a genus of evergreen coniferous trees in the family Pinaceae (subfamily Laricoideae). Common names for species in the genus include Douglas fir, Douglas-fir, Douglas tree, Oregon pine and Bigcone spruce. '' Pseudotsuga menz ...
'', they extend beyond the cone scales. File:Castilleja levisecta bracts.jpg, Bright yellow bracts of ''
Castilleja levisecta ''Castilleja levisecta'' is a rare species of flowering plant in the family Orobanchaceae known by the common name golden paintbrush, or golden Indian paintbrush, listed under the Endangered Species Act in 1997. It is native to British Columbia a ...
'' are more tightly pressed against the stem than bracts of other ''
Castilleja ''Castilleja'', commonly known as paintbrush, Indian paintbrush, or prairie-fire, is a genus of about 200 species of annual and perennial herbaceous plants native to the west of the Americas from Alaska south to the Andes, northern Asia, and one ...
'' species. File:Bougainvillea glabra.JPG, Bracts of ''
Bougainvillea glabra ''Bougainvillea glabra'', the lesser bougainvillea or paperflower, is the most common species of bougainvillea used for bonsai. The epithet 'glabra' comes from Latin and means "bald". Description It is an evergreen, climbing shrub with thick ...
'', differ in colour from the non-bract leaves, and attract pollinators. File:Banana bract.jpg, Bracts along a
banana A banana is an elongated, edible fruit – botanically a berry – produced by several kinds of large herbaceous flowering plants in the genus ''Musa''. In some countries, bananas used for cooking may be called "plantains", distinguis ...
flower stalk surround the rows of flowers File:E milii vulcanii ies.jpg, ''
Euphorbia milii ''Euphorbia milii'', the crown of thorns, Christ plant, or Christ thorn, is a species of flowering plant in the spurge family Euphorbiaceae, native to Madagascar. The species name commemorates Baron Milius, once Governor of Réunion, who introdu ...
'' var. ''vulcanii'' cyathia bearing a pair of pinkish cyathophylls. File:Ananas bracteatus var tricolor.jpg, Colourful bracts of ''
Ananas bracteatus The pineapple (''Ananas comosus'') is a tropical plant with an edible fruit; it is the most economically significant plant in the family Bromeliaceae. The pineapple is indigenous to South America, where it has been cultivated for many centurie ...
''. File:Erigeron peregrinus 6287.JPG, The many green
involucral bract In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
s of ''
Erigeron peregrinus ''Erigeron peregrinus'' is a North American species of flowering plants in the family Asteraceae known by the common name wandering fleabane. ''Erigeron peregrinus'' is native to northwestern North America from Alaska to Oregon. Some populations ...
'' are linear, loose, taper to a point, about the same length, and help to distinguish this species. File:Blooming Dogwood.jpg,
Dogwood ''Cornus'' is a genus of about 30–60 species of woody plants in the family Cornaceae, commonly known as dogwoods, which can generally be distinguished by their blossoms, berries, and distinctive bark. Most are deciduous trees or shrub ...
species ''
Cornus florida ''Cornus florida'', the flowering dogwood, is a species of flowering tree in the family Cornaceae native to eastern North America and northern Mexico. An endemic population once spanned from southernmost coastal Maine south to northern Florida ...
'' inflorescence showing four large white bracts and central flower cluster. File:2013-05-10 08 26 08 Closeup of pink dogwoods at the Brendan T. Byrne State Forest headquarters.jpg, ''
Cornus florida ''Cornus florida'', the flowering dogwood, is a species of flowering tree in the family Cornaceae native to eastern North America and northern Mexico. An endemic population once spanned from southernmost coastal Maine south to northern Florida ...
'' dogwood
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...
with pink bracts surrounding small flower cluster. File:豔紅苞鳳梨 20190712192436.jpg, The red bracts of ''
Aechmea bracteata ''Aechmea bracteata'' is a plant species in the genus ''Aechmea''. This species is native to Central America, Mexico, Colombia, and Venezuela; it is also reportedly naturalized in the Bahamas The Bahamas (), officially the Commonwealth ...
''. File:女王鬱金 20190715203108 01.jpg, The pink bracts of ''
Curcuma petiolata ''Curcuma petiolata'' (also known as jewel of Thailand, Siam tulip, hidden ginger, pastel hidden ginger, hidden lily or queen lily) is a plant of the Zingiberaceae or ginger family. It is native to Thailand, Myanmar, Laos and Java. Average heig ...
''. File:火鶴 20190729181532.jpg, The pink bract of a new cultivar: '' Anthurium andraeanum'' 'Pink Lady'. File:豔山姜花序總苞 20190508131956.jpg, The inflorescence of '' Alpinia zerumbet'' protected by two bracts before flower bloom. File:地湧金蓮 Musella lasiocarpa 20201028130818 01.jpg, The golden bracts of ''
Musella lasiocarpa ''Musella lasiocarpa'' ( syn. ''Musa lasiocarpa''), commonly known as Chinese dwarf banana, golden lotus banana or Chinese yellow banana, is the sole species in the genus ''Musella''. It is thus a close relative of bananas, and also a member of ...
''. File:Hydrangea involucrata (9344379310).jpg, Large bracts of ''
Hydrangea involucrata ''Hydrangea involucrata'' is a species of flowering plant in the family Hydrangeaceae, native to Japan and Taiwan.Pages 67-69; ''Hydrangeas for American Gardens'' by Michael Dirr. Timber Press, 2004. /. It is a deciduous In the fields of hor ...
'' surround tiny flowers and demonstrate the "lacecap" style
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
that has made various ''
Hydrangea ''Hydrangea'', () commonly named the hortensia, is a genus of over 75 species of flowering plants native to Asia and the Americas. By far the greatest species diversity is in eastern Asia, notably China, Korea, and Japan. Most are shrubs tall, ...
'' species and
cultivars A cultivar is a type of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and when Plant propagation, propagated retain those traits. Methods used to propagate cultivars include: division, root and st ...
popular with gardeners. File:Viburnum opulus RF.jpg, Like many ''
Hydrangea ''Hydrangea'', () commonly named the hortensia, is a genus of over 75 species of flowering plants native to Asia and the Americas. By far the greatest species diversity is in eastern Asia, notably China, Korea, and Japan. Most are shrubs tall, ...
'' species (
Hydrangeaceae Hydrangeaceae is a family of flowering plants in the order Cornales, with a wide distribution in Asia and North America, and locally in southeastern Europe. Description The genera are characterised by leaves in opposite pairs (rarely whorled ...
family), ''
Viburnum opulus ''Viburnum opulus'', the guelder-rose or guelder rose () is a species of flowering plant in the family (biology), family Adoxaceae (formerly Caprifoliaceae) native plant, native to Europe, northern Africa and central Asia. Description ''Viburn ...
'' (
Adoxaceae Adoxaceae, commonly known as moschatel family, is a small family of flowering plants in the order Dipsacales, now consisting of five genera and about 150–200 species. They are characterised by opposite toothed leaves, small five- or, more rare ...
family) has bracts that surround tiny flowers, creating a "lacecap" style
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
.


Bracteole

A small bract is called a bracteole or bractlet. Technically this is any bract that arises ''on'' a pedicel instead of subtending it.


Involucral bracts

Bracts that appear in a
whorl A whorl ( or ) is an individual circle, oval, volution or equivalent in a whorled pattern, which consists of a spiral or multiple concentric objects (including circles, ovals and arcs). Whorls in nature File:Photograph and axial plane floral ...
subtending an
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
are collectively called an involucre. An involucre is a common feature beneath the inflorescences of many
Apiaceae Apiaceae or Umbelliferae is a family of mostly aromatic flowering plants named after the type genus ''Apium'' and commonly known as the celery, carrot or parsley family, or simply as umbellifers. It is the 16th-largest family of flowering plants ...
,
Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae w ...
,
Dipsacaceae The Dipsacaceae have been recognized as a family (the teasel family) of the order Dipsacales containing 350 species of perennial or biennial herbs and shrubs in eleven genera. The species are currently placed in the family Caprifoliaceae. Native ...
and
Polygonaceae The Polygonaceae are a family of flowering plants known informally as the knotweed family or smartweed—buckwheat family in the United States. The name is based on the genus ''Polygonum'', and was first used by Antoine Laurent de Jussieu in 1789 ...
. Each flower in an inflorescence may have its own whorl of bracts, in this case called an involucel. In this case they may be called ''chaff'', ''paleas'', or ''receptacular bracts'' and are usually minute scales or bristles. Many asteraceous plants have bracts at the base of each inflorescence. The term involucre is also used for a highly conspicuous bract or bract pair at the base of an inflorescence. In the family
Betulaceae Betulaceae, the birch family, includes six genera of deciduous nut-bearing trees and shrubs, including the birches, alders, hazels, hornbeams, hazel-hornbeam, and hop-hornbeams numbering a total of 167 species. They are mostly natives of the t ...
, notably in the genera ''
Carpinus Hornbeams are hardwood trees in the flowering plant genus ''Carpinus'' in the birch family Betulaceae. The 30–40 species occur across much of the temperate regions of the Northern Hemisphere. Origin of names The common English name ''hornbea ...
'' and ''
Corylus The hazel (''Corylus'') is a genus of deciduous trees and large shrubs native to the temperate Northern Hemisphere. The genus is usually placed in the birch family Betulaceae,Germplasmgobills Information Network''Corylus''Rushforth, K. (1999). ...
'', the involucre is a leafy structure that protects the developing nuts. Beggar-tick ('' Bidens comosa'') has narrow involucral bracts surrounding each inflorescence, each of which also has a single bract below it. There is then a pair of leafy bracts on the main stem and below those a pair of leaves.


Epicalyx

An epicalyx, which forms an additional whorl around the calyx of a single flower, is a modification of bracteoles In other words, the epicalyx is a ''group of bracts resembling a calyx'' or ''bracteoles forming a whorl outer to the calyx''. It is a calyx-like extra whorl of floral appendages. Each individual segment of the epicalyx is called an episepal because they resemble the sepals in them. They are present in the hibiscus family,
Malvaceae Malvaceae, or the mallows, is a family of flowering plants estimated to contain 244 genera with 4225 known species. Well-known members of economic importance include okra, cotton, cacao and durian. There are also some genera containing familiar ...
. ''
Fragaria ''Fragaria'' () is a genus of flowering plants in the rose family, Rosaceae, commonly known as strawberries for their edible fruits. There are more than 20 described species and many hybrids and cultivars. The most common strawberries grown com ...
'' (strawberries) may or may not have an epicalyx.


Spathe

A ''spathe'' is a large bract or pair of bracts forming a sheath to enclose the flower cluster of such plants as
palm Palm most commonly refers to: * Palm of the hand, the central region of the front of the hand * Palm plants, of family Arecaceae **List of Arecaceae genera * Several other plants known as "palm" Palm or Palms may also refer to: Music * Palm (ba ...
s,
arum ''Arum'' is a genus of flowering plants in the family Araceae, native to Europe, northern Africa, and western and central Asia, with the highest species diversity in the Mediterranean region. Frequently called arum lilies, they are not closely ...
s, irises,Richard Lynch
crocus ''Crocus'' (; plural: crocuses or croci) is a genus of seasonal flowering plants in the family Iridaceae (iris family) comprising about 100 species of perennials growing from corms. They are low growing plants, whose flower stems remain undergro ...
es,Alex Casha and
dayflower ''Commelina'' is a genus of approximately 170 species commonly called dayflowers due to the short lives of their flowers. They are less often known as widow's tears. It is by far the largest genus of its family, Commelinaceae. The Swedish taxonom ...
s (''
Commelina ''Commelina'' is a genus of approximately 170 species commonly called dayflowers due to the short lives of their flowers. They are less often known as widow's tears. It is by far the largest genus of its family, Commelinaceae. The Swedish taxonom ...
''). ''
Zephyranthes tubispatha ''Zephyranthes tubispatha'', synonym ''Habranthus tubispathus'', the Rio Grande copperlily or Barbados snowdrop, is a species of flowering plant in the family Amaryllidaceae. It is a perennial bulb native to southern South America (Brazil, Arge ...
'' in the
Amaryllidaceae The Amaryllidaceae are a family of herbaceous, mainly perennial and bulbous (rarely rhizomatous) flowering plants in the monocot order Asparagales. The family takes its name from the genus ''Amaryllis'' and is commonly known as the amaryllis fa ...
derives its specific name from its tubular spathe. In many arums (family
Araceae The Araceae are a family of monocotyledonous flowering plants in which flowers are borne on a type of inflorescence called a spadix. The spadix is usually accompanied by, and sometimes partially enclosed in, a spathe (or leaf-like bract). Also ...
), the spathe is petal-like, attracting pollinators to the flowers arranged on a type of spike called a spadix.


References

{{Botany Plant morphology Leaves