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The southern pygmy perch (''Nannoperca australis''), also known as the Tasmanian pygmy perch, is a species of freshwater
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...
, a temperate perch from the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Percichthyidae which is native to southeastern
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
and
Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...
.


Description

The southern pygmy perch has an oblong and moderately compressed body with a convex dorsal profile and a straight ventral profile. It has a large head the top of which bulges slightly and a blunt snout. It has a slightly oblique, terminal mouth which is protractile with the
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
reaching to a level near the centre of the eye. There are thin bands of villiform teeth on the jaws and the roof of the mouth. There is a
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...
made up of an irregular series of pored scales. The dorsal fin has 7-9 spines in its front part, separated from the rear part by a deep notch, the rear part contains 7-10 soft rays. The
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
is similar in shape to the soft part of the
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
and has 3 spines and 7-8 soft rays. The large
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
is rounded. They are variable in colour, the colour apparently being determined by their environment and can be pale cream through to greenish-brown, with paler underparts. They can also show irregular markings on the flanks such as dark spotting or horizontal stripes. In the breeding season the males are brighter as their dorsal, anal and caudal fins develop a red colour and have black margins and the
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two en ...
s and the area around the anus become black. These fish attain a maximum
total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...
of but are more commonly recorded at standard lengths of around .


Distribution

Southern pygmy perch are
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to temperate rivers systems in southeastern Australia, especially the
Murray Murray may refer to: Businesses * Murray (bicycle company), an American manufacturer of low-cost bicycles * Murrays, an Australian bus company * Murray International Trust, a Scottish investment trust * D. & W. Murray Limited, an Australian who ...
- Murrumbidgee river system from Brinagagee bear
Darlington Point Darlington Point is a small town on the banks of the Murrumbidgee River in the Riverina district of western New South Wales, Australia. It is part of the Murrumbidgee Council local government area.  The centre of town is four kilometres from ...
in
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
, in
Victoria (Australia) Victoria is a state in southeastern Australia. It is the second-smallest state with a land area of , the second most populated state (after New South Wales) with a population of over 6.5 million, and the most densely populated state in ...
west to
Ewens Ponds Ewens Ponds is a series of three water-filled limestone sinkholes in the Australian state of South Australia located in the gazetted locality of Eight Mile Creek, on the watercourse of Eight Mile Creek about south of Mount Gambier and eas ...
in
South Australia South Australia (commonly abbreviated as SA) is a state in the southern central part of Australia. It covers some of the most arid parts of the country. With a total land area of , it is the fourth-largest of Australia's states and territories ...
. They also occur on Flinders Island and on
King Island King Island, Kings Island or King's Island may refer to: Australia * King Island (Queensland) * King Island, at Wellington Point, Queensland * King Island (Tasmania) ** King Island Council, the local government area that contains the Tasmanian is ...
in the
Bass Strait Bass Strait () is a strait separating the island state of Tasmania from the Australian mainland (more specifically the coast of Victoria, with the exception of the land border across Boundary Islet). The strait provides the most direct waterwa ...
and in the northward draining rivers in Tasmania.


Habitat and biology

Southern pygmy perch are found in a wide variety of freshwater habitats so long as they have aquatic vegetation. They can be found in both still or slow flowing waters and they have been recorded in low current streams, lakes, billabongs, ditches, impoundments,
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s and
ephemeral Ephemerality (from the Greek word , meaning 'lasting only one day') is the concept of things being transitory, existing only briefly. Academically, the term ephemeral constitutionally describes a diverse assortment of things and experiences, fr ...
wetland A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The ...
s. In all of these habitat they prefer the vegetated marginal areas. It is a mainly
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
species with its diet mostly consisting of small
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s such as
mosquito larva Mosquitoes (or mosquitos) are members of a group of almost 3,600 species of small flies within the family Culicidae (from the Latin ''culex'' meaning "gnat"). The word "mosquito" (formed by ''mosca'' and diminutive ''-ito'') is Spanish for "litt ...
e and other
aquatic insect Aquatic insects or water insects live some portion of their life cycle in the water. They feed in the same ways as other insects. Some ''diving'' insects, such as predatory diving beetles, can hunt for food underwater where land-living insects c ...
s, as well as '' Daphnia'' and other
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s. As they mature they prey on larger insects including
mayflies Mayflies (also known as shadflies or fishflies in Canada and the upper Midwestern United States, as Canadian soldiers in the American Great Lakes region, and as up-winged flies in the United Kingdom) are aquatic insects belonging to the orde ...
but their small mouth means that they do not consume many other fishes. Southern pygmy perch attain sexual maturity during the first year after hatching, mature males are mature at about and females at around . The breeding season runs from the late winter into the early spring once the water temperatures rise above . Breeding males are
territorial A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
and defend a small area of water from other males. The female lays small clutches of eggs in the aquatic vegetation or onto the riverbed within the male's territory. These hatch into larvae of 3-4mm in length 2-4 days after fertilisation.


Conservation

The Southern pygmy perch was formerly widespread in New South Wales but has now largely been extirpated from the State, there are only three known locations where it can still be found. It is still common in coastal Victoria but it is rare in the Murray catchment of that state and it is rare and threatened in South Australia. In Tasmania it remains relatively common. The reasons for the decline and continuing threats to this species includes invasive alien fish such as
brown trout The brown trout (''Salmo trutta'') is a European species of salmonid fish that has been widely introduced into suitable environments globally. It includes purely freshwater populations, referred to as the riverine ecotype, ''Salmo trutta'' morph ...
, '' Gambusia'' and European perch as well as habitat fragmentation and destruction The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
assessed the conservation status of this species as
Near Threatened A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify fo ...
in 2019. The Department of Primary Industries in New South Wales has undertaken a captive breeding programme for this species.


Taxonomy

The Southern pygmy perch was first formally described in 1861 by Albert Günther with the type locality given as the Murray River. Molecular studies have shown that there are two cryptic species within ''Nannoperca australis'' one from the eastern coastal streams and the other from the Murray–Darling basin and the western coastal streams.


Aquarium

The Southern pygmy perch is in the
aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
trade within Australia where it is kept in cold water aquaria and in ponds.


References


External links


Southern Pygmy Perch
{{Taxonbar, from=Q5747401 Nannoperca Freshwater fish of Australia Fish described in 1861 Taxa named by Albert Günther