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The Sorraia is a rare
breed A breed is a specific group of domestic animals having homogeneous appearance (phenotype), homogeneous behavior, and/or other characteristics that distinguish it from other organisms of the same species. In literature, there exist several slig ...
of
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million y ...
indigenous to the portion of the
Iberian peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
, in the
Sorraia River The Sorraia River (), or Rio Sorraia, is a river in Santarém District, Portugal. It forms at the confluence of the Sor and Raia, near the town of Couço. Having a basin of , it flows for , joining the Tagus near Murteira. Along its length it ...
basin, in
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
. The Sorraia is known for its primitive features, including a convex profile and
dun A dun is an ancient or medieval fort. In Ireland and Britain it is mainly a kind of hillfort and also a kind of Atlantic roundhouse. Etymology The term comes from Irish language, Irish ''dún'' or Scottish Gaelic ''dùn'' (meaning "fort"), ...
coloring with
primitive markings Primitive markings are a group of hair coat markings and qualities seen in several equine species, including horses, donkeys, and asses. In horses, they are associated with primitive breeds, though not limited to such breeds. The markings are ...
. Concerning its origins, a theory has been advanced by some authors that the Sorraia is a descendant of primitive horses belonging to the naturally occurring wild
fauna Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as '' biota''. Zoo ...
of Southern
Iberia The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
. Studies are currently ongoing to discover the relationship between the Sorraia and various wild horse types, as well as its relationship with other breeds from the Iberian Peninsula and Northern Africa. Members of the breed are small, but hardy and well-adapted to harsh conditions. They were occasionally captured and used by native farmers for centuries, and a remnant population of these nearly
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
horses was discovered by a Portuguese
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...
in the early 20th century. Today, the Sorraia has become the focus of preservation efforts, with European scientists leading the way and enthusiasts from several countries forming projects and establishing herds to assist in the re-establishment of this breed from its current endangered status.


Characteristics

The Sorraia breed stands between high, although some individuals are as small as The head tends to be large, the profile convex, and the ears long. The neck is slender and long, the
withers The withers is the ridge between the shoulder blades of an animal, typically a quadruped. In many species, it is the tallest point of the body. In horses and dogs, it is the standard place to measure the animal's height. In contrast, cattle ar ...
high, and the
croup Croup, also known as laryngotracheobronchitis, is a type of respiratory infection that is usually caused by a virus. The infection leads to swelling inside the trachea, which interferes with normal breathing and produces the classic symptoms o ...
slightly sloping. The legs are strong, with long
pastern The is a part of the leg of a horse between the fetlock and the top of the hoof. It incorporates the long pastern bone (proximal phalanx) and the short pastern bone (middle phalanx), which are held together by two sets of paired ligaments to fo ...
s and well-proportioned hooves. These horses have good endurance and are
easy keeper Easy may refer to: Arts and entertainment Film and television * ''Easy'' (film), a 2003 American romantic comedy film *''Easy!'', or ''Scialla!'', a 2011 Italian comedy film * ''Easy'' (TV series), a 2016–2019 American comedy-drama anthology ...
s, thriving on relatively little
fodder Fodder (), also called provender (), is any agriculture, agricultural foodstuff used specifically to feed domesticated livestock, such as cattle, domestic rabbit, rabbits, sheep, horses, chickens and pigs. "Fodder" refers particularly to food g ...
. They have a reputation for being independent of temperament, but tractable. On adult horses, the lay of the hair can create the appearance of stars and flags on the neck and chest. Also due to the lay of the hair, newborn
foal A foal is an equine up to one year old; this term is used mainly for horses, but can be used for donkeys. More specific terms are colt for a male foal and filly for a female foal, and are used until the horse is three or four. When the foal i ...
s can appear to have stripes all over, reminiscent of
zebra Zebras (, ) (subgenus ''Hippotigris'') are African equines with distinctive black-and-white striped coats. There are three living species: the Grévy's zebra (''Equus grevyi''), plains zebra (''E. quagga''), and the mountain zebra (''E. zeb ...
stripes. The breed standard refers to this as "hair stroke".


Color

Sorraia are generally
dun A dun is an ancient or medieval fort. In Ireland and Britain it is mainly a kind of hillfort and also a kind of Atlantic roundhouse. Etymology The term comes from Irish language, Irish ''dún'' or Scottish Gaelic ''dùn'' (meaning "fort"), ...
or a dun variation called
grullo Grulla or grullo, also called blue dun, gray dun or mouse dun, is a color of horses in the dun family, characterized by tan-gray or mouse-colored hairs on the body, often with shoulder and dorsal stripes and black barring on the lower legs. In th ...
. Dun coloring includes
primitive markings Primitive markings are a group of hair coat markings and qualities seen in several equine species, including horses, donkeys, and asses. In horses, they are associated with primitive breeds, though not limited to such breeds. The markings are ...
such as a black dorsal stripe, black tipped ears, horizontal striping on the legs and a dark muzzle area. The dark muzzle area is in contrast to some other dun-colored horse breeds, who have light-colored muzzle areas and underbellies, possibly due to the presence of pangare genetics. Sorraia horses have bi-colored manes and tails with lighter colored hairs that fringe the outside of the longer growing black hair. This is a characteristic shared with other predominantly dun-colored breeds, such as the
Fjord horse The Fjord Horse or Norwegian Fjord Horse ( no, fjordhest) is a relatively small but very strong horse breed from the mountainous regions of western Norway. It is an agile breed of light draught horse build. All Fjord horses are dun in colour, ...
. Purebred Sorraia occasionally have white markings, although they are rare and undesired by the breed's
studbook A breed registry, also known as a herdbook, studbook or register, in animal husbandry and the hobby of animal fancy, is an official list of animals within a specific breed whose parents are known. Animals are usually registered by their breeders ...
.


Genetics

The relationship between the Sorraia and other
breeds A breed is a specific group of domestic animals having homogeneous appearance (phenotype), homogeneous behavior, and/or other characteristics that distinguish it from other organisms of the same species. In literature, there exist several slight ...
remains largely undetermined, as is its relationship to the
wild horse The wild horse (''Equus ferus'') is a species of the genus ''Equus'', which includes as subspecies the modern domesticated horse (''Equus ferus caballus'') as well as the endangered Przewalski's horse (''Equus ferus przewalskii''). The Europea ...
subspecies, the
Tarpan The term tarpan (''Equus ferus ferus'') refers to free-ranging horses of the Russian steppe from the 18th to the 20th century. It is generally unknown whether those horses represented genuine wild horses, feral domestic horses or hybrids. The las ...
and the
Przewalski's Horse Przewalski's horse (, , (Пржевальский ), ) (''Equus ferus przewalskii'' or ''Equus przewalskii''), also called the takhi, Mongolian wild horse or Dzungarian horse, is a rare and endangered horse originally native to the steppes of Ce ...
. The Sorraia originally developed in the southern part of the Iberian peninsula. d'Andrade hypothesized that the Sorraia would be the ancestor of the Southern Iberian breeds. Morphologically, scientists place the Sorraia as closely related to the Gallego and the Asturcon, but genetic studies using
mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
show that the Sorraia forms a cluster that is largely separated from most Iberian breeds. Some evidence links this cluster with
Konik The Konik or Polish Konik, pl, konik polski, is a Polish List of horse breeds, breed of pony. There are semi-feral populations in some regions. They are usually dun gene, mouse dun or primitive markings, striped dun in color. The Bilgoray, p ...
and domestic Mongolian horses. At the same time, one of the maternal lineages is shared with the
Lusitano The Lusitano, also known as the Pure Blood Lusitano or PSL (''Puro Sangue Lusitano''), is a Portuguese horse breed. Horses were known to be present on the Iberian Peninsula as far back as 20,000 BC, and by 800 BC the region was renowned for ...
. Genetic evidence has not supported a hypothesis that the Sorraia is related to the
Barb horse The Barb or Berber horse (Berber: ''Ayis Amaziɣ, ⴰⵢⵢⵙ ⴰⵎⴰⵣⵉⴳ''; ) is a North African breed of riding horse with great hardiness and stamina. It is closely associated with the Berber or Amazigh peoples of the Maghreb. It ha ...
, an African breed introduced to Iberia by the Moors. Multiple authors have suggested that the Sorraia might be a descendant of the
Tarpan The term tarpan (''Equus ferus ferus'') refers to free-ranging horses of the Russian steppe from the 18th to the 20th century. It is generally unknown whether those horses represented genuine wild horses, feral domestic horses or hybrids. The las ...
based on shared morphological features, principally the typical color of its coat. Other authors simply state that the Sorraia has "evident primitive characteristics", although they do not refer to a specific ancestor. However, there have been no genetic studies comparing the Sorraia with the Tarpan, and similarity of external morphology is an unreliable measure of relatedness. Genetic studies to date have been inconclusive about the closest relative of the Sorraia. On one hand, studies using mitochondrial DNA showed a relationship with the
Przewalski's Horse Przewalski's horse (, , (Пржевальский ), ) (''Equus ferus przewalskii'' or ''Equus przewalskii''), also called the takhi, Mongolian wild horse or Dzungarian horse, is a rare and endangered horse originally native to the steppes of Ce ...
, in that Przewalski's Horse has a unique haplotype (A2) not found in domestic horses, which differs by just one single
nucleotide Nucleotides are organic molecules consisting of a nucleoside and a phosphate. They serve as monomeric units of the nucleic acid polymers – deoxyribonucleic acid (DNA) and ribonucleic acid (RNA), both of which are essential biomolecules wi ...
from one of the major Sorraia haplotypes (JSO41, later A7). In comparison, genetic distances within the domestic horse are as large as 11 nucleotide differences. However, this relationship with the Przewalski's Horse was contradicted in another study using
microsatellite A microsatellite is a tract of repetitive DNA in which certain DNA motifs (ranging in length from one to six or more base pairs) are repeated, typically 5–50 times. Microsatellites occur at thousands of locations within an organism's genome. ...
data that showed that the genetic distance between the Prewalski's Horse and the Sorraia was the largest. Such conflicting results can arise when a population passes through a
genetic bottleneck A population bottleneck or genetic bottleneck is a sharp reduction in the size of a population due to environmental events such as famines, earthquakes, floods, fires, disease, and droughts; or human activities such as specicide, widespread violen ...
, and evidence suggests that the Sorraia, among other rare breeds, has recently passed through a bottleneck, effectively obscuring the position of this breed in the family tree of the domestic horse. Thus, the morphological,
physiological Physiology (; ) is the scientific study of functions and mechanisms in a living system. As a sub-discipline of biology, physiology focuses on how organisms, organ systems, individual organs, cells, and biomolecules carry out the chemical ...
, and cultural characteristics of the Sorraia are the subject of continued study to better understand the relationship between various Iberian horse breeds and wild horse subspecies.


History

Although it is known that the Sorraia developed in the southern part of the Iberian peninsula, the breed was isolated and unknown to science until the 20th century. Despite the lack of documentation, attempts have been made to reconstruct its history.
Paleolithic The Paleolithic or Palaeolithic (), also called the Old Stone Age (from Greek: παλαιός ''palaios'', "old" and λίθος ''lithos'', "stone"), is a period in human prehistory that is distinguished by the original development of stone too ...
parietal art In archaeology, rock art is human-made markings placed on natural surfaces, typically vertical stone surfaces. A high proportion of surviving historic and prehistoric rock art is found in caves or partly enclosed rock shelters; this type also m ...
images in the region depict equines with a distinct likeness to the Sorraia, with similar zebra-like markings. Analysis of
mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA ...
has been performed on
Mustang The mustang is a free-roaming horse of the Western United States, descended from horses brought to the Americas by the Spanish. Mustangs are often referred to as wild horses, but because they are descended from once-domesticated animals, they ...
s in the western United States that show similar mtDNA patterns between some Mustangs and the Sorraia breed. Spanish
conquistador Conquistadors (, ) or conquistadores (, ; meaning 'conquerors') were the explorer-soldiers of the Spanish and Portuguese Empires of the 15th and 16th centuries. During the Age of Discovery, conquistadors sailed beyond Europe to the Americas, O ...
s took
Iberian horse The Iberian horse is a designation given to a number of horse breeds native to the Iberian peninsula. At present, some breeds are officially recognized by the FAO,
s, some of whom closely resembled the modern-day Sorraia, to the Americas in their conquests, probably as pack animals. Similarities between the Sorraia and several North and South American breeds are shown in the dun and grullo coloring and various physical characteristics. This evidence suggests that the Sorraia, their ancestors, or other horses with similar features, may have had a long history in the Iberian region and a role in the creation of American breeds. Otherwise, the Sorraia breed was lost to history until 1920, when Portuguese zoologist and paleontologist Dr. Ruy d'Andrade first encountered the Sorraia horse during a hunting trip in the Portuguese lowlands. This remnant herd of primitive horses had continued to live a wild existence in these lowlands, which were rather inaccessible and had been used as a hunting preserve by Portuguese royalty until the early 1900s. At the time of d'Andrade's initial meeting the breed, the horses were ill-regarded by native farmers, although they were considered hardy native fauna that lived off of the uncultivated lands and salt marshes in the local river valleys. For centuries, peasant farmers of the area would occasionally capture the horses and use them for
agricultural Agriculture or farming is the practice of cultivating Plant, plants and livestock. Agriculture was the key development in the rise of Sedentism, sedentary human civilization, whereby farming of Domestication, domesticated species created food ...
work, including
threshing Threshing, or thrashing, is the process of loosening the edible part of grain (or other crop) from the straw to which it is attached. It is the step in grain preparation after reaping. Threshing does not remove the bran from the grain. History ...
grain A grain is a small, hard, dry fruit (caryopsis) – with or without an attached hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and legum ...
and
herding Herding is the act of bringing individual animals together into a group (herd), maintaining the group, and moving the group from place to place—or any combination of those. Herding can refer either to the process of animals forming herds in ...
bull A bull is an intact (i.e., not castrated) adult male of the species ''Bos taurus'' (cattle). More muscular and aggressive than the females of the same species (i.e., cows), bulls have long been an important symbol in many religions, includin ...
s. In the 1920s and 1930s, as mechanization became more prevalent, both wild and domesticated breeding stock diminished to almost nothing, and d'Andrade, along with his son Fernando, encouraged the conservation of the breed. In 1937, d'Andrade began a small herd of his own with five
stallion A stallion is a male horse that has not been gelded (castrated). Stallions follow the conformation and phenotype of their breed, but within that standard, the presence of hormones such as testosterone may give stallions a thicker, "cresty" nec ...
s and seven
mare A mare is an adult female horse or other equine. In most cases, a mare is a female horse over the age of three, and a filly is a female horse three and younger. In Thoroughbred horse racing, a mare is defined as a female horse more than four ...
s from horses obtained near
Coruche Coruche () is a municipality in Santarém District in Portugal. The population in 2011 was 19,944, in an area of 1115.72 km². The present Mayor is Francisco Silvestre de Oliveira, elected by the Socialist Party. The Coruche City Council ha ...
, Portugal. All Sorraias currently in captivity descend from these original horses obtained by d'Andrade, and it is believed that the remnant wild herds of the breed died out soon after. These horses were kept in a habitat similar to their native one. In 1975, two other farms took up the Sorraia's cause and acquired small herds to help with conservation. In 1976, three stallions and three mares were imported to Germany from Portugal to begin a sub-population there. In March 2004, a small breeding herd of Sorraia horses was released on the estate of a private land owner who dedicated a portion of his property so that these horses could live completely wild, as did their ancestors. The refuge created for them is in the Vale de Zebro region of south western Portugal, one of places so named because this is where the Sorraia's predecessors dwelt. Today, the breed is nearly extinct, with fewer than 200 horses existing as of 2007, including around 80 breeding mares. The
Food and Agriculture Organization The Food and Agriculture Organization of the United Nations (FAO)french: link=no, Organisation des Nations unies pour l'alimentation et l'agriculture; it, Organizzazione delle Nazioni Unite per l'Alimentazione e l'Agricoltura is an intern ...
considers it to be maintaining critical risk status. The first
studbook A breed registry, also known as a herdbook, studbook or register, in animal husbandry and the hobby of animal fancy, is an official list of animals within a specific breed whose parents are known. Animals are usually registered by their breeders ...
was published in 2004, dedicated to maintaining a written record of the bloodlines of the Sorraia. Sorraias are present mainly in Portugal, with a small population in Germany. While not bred for a specific use, the Sorraia horses are versatile and have been used in herding bulls,
dressage Dressage ( or ; a French term, most commonly translated to mean "training") is a form of horse riding performed in exhibition and competition, as well as an art sometimes pursued solely for the sake of mastery. As an equestrian sport defined b ...
riding and light harness.


American preservation efforts

Two Sorraia stallions were imported to the United States in the early 21st century. In 2006, another Sorraia stallion was imported to Canada where a Sorraia Mustang Preserve has been established on
Manitoulin Island Manitoulin Island is an island in Lake Huron, located within the borders of the Canadian province of Ontario, in the bioregion known as Laurentia. With an area of , it is the largest lake island in the world, large enough that it has over 100 ...
in Ontario. Unrelated to existing preservation efforts which work in conjunction with the Sorraia Mustang Studbook, another project by a consortium of breeders in the United States is attempting to establish a separate network and studbook. These breeders have gathered
Spanish Mustang The Spanish Mustang is an American horse breed descended from horses brought from Spain during the early conquest of the Americas. They are classified within the larger grouping of the Colonial Spanish horse, a type that today is rare in Spain. ...
s that through
mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA ...
testing show a genetic relationship with the Sorraia and are breeding them according to both
genotype The genotype of an organism is its complete set of genetic material. Genotype can also be used to refer to the alleles or variants an individual carries in a particular gene or genetic location. The number of alleles an individual can have in a ...
and
phenotype In genetics, the phenotype () is the set of observable characteristics or traits of an organism. The term covers the organism's morphology or physical form and structure, its developmental processes, its biochemical and physiological proper ...
in an attempt to help preserve what they are calling the "American Sorraia Mustang".


Naming

Dr. Ruy d'Andrade gave the breed their name of "Sorraia". D'Andrade took the name from the
Sorraia River The Sorraia River (), or Rio Sorraia, is a river in Santarém District, Portugal. It forms at the confluence of the Sor and Raia, near the town of Couço. Having a basin of , it flows for , joining the Tagus near Murteira. Along its length it ...
in Portugal. The breed had previously been known by the local Portuguese as "zebro" or "zebra", due to their markings. In the time of
Christopher Columbus Christopher Columbus * lij, Cristoffa C(or)ombo * es, link=no, Cristóbal Colón * pt, Cristóvão Colombo * ca, Cristòfor (or ) * la, Christophorus Columbus. (; born between 25 August and 31 October 1451, died 20 May 1506) was a ...
, the Sorraia was also known as the Marismeño, but the Sorraia and the
Marismeño The Marismeño is a rare breed of horse indigenous to the marshes of the Guadalquivir River, from which it takes its name. It is now found particularly in the Doñana National Park, which lies mostly in the province of Huelva, in Andalusia, sou ...
have evolved into two different breeds over time. Today, the name Marismeño refers to a population of semiferal horses living in Doñana Natural Park in Spain.


References


External links


A page dedicated to the Sorraia.
{{good article Horse breeds Feral horses Horse breeds originating in Portugal