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The order Sordariales is one of the most diverse taxonomic groups within the Sordariomycetes (subdivision
Pezizomycotina Pezizomycotina make up most of the Ascomycota fungi and include most lichenized fungi too. Pezizomycotina contains the filamentous ascomycetes and is a subdivision of the Ascomycota (fungi that form their spores in a sac-like ''ascus''). It is mo ...
,
division Division or divider may refer to: Mathematics *Division (mathematics), the inverse of multiplication *Division algorithm, a method for computing the result of mathematical division Military *Division (military), a formation typically consisting ...
Ascomycota Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The def ...
). Species in the order Sordariales have a broad range of ecological diversity, containing lignicolous, herbicolous and coprophilous taxa. Most Sordariales are
saprobic Saprotrophic nutrition or lysotrophic nutrition is a process of chemoheterotrophic extracellular digestion involved in the processing of decayed (dead or waste) organic matter. It occurs in saprotrophs, and is most often associated with fungi ( ...
, producing solitary perithecial
ascomata An ascocarp, or ascoma (), is the fruiting body ( sporocarp) of an ascomycete phylum fungus. It consists of very tightly interwoven hyphae and millions of embedded asci, each of which typically contains four to eight ascospores. Ascocarps are mos ...
. They are commonly found on dung or decaying plant matter. The order contains a number of ecologically important species, including the model filamentous fungal genera
Podospora ''Podospora'' is a genus of fungi in the family Podosporaceae. Fossils of ''Podospora'' have been reported from 12 million year old rocks from central England. Species *''Podospora adelura'' *''Podospora alexandri'' *''Podospora ampullacea'' ...
and
Neurospora ''Neurospora'' is a genus of Ascomycete fungi. The genus name, meaning "nerve spore" refers to the characteristic striations on the spores that resemble axons. The best known species in this genus is '' Neurospora crassa'', a common model organ ...
, as well as potentially industrial-relevant fungi, such as members of the Chaetomiaceae family, which often produce biologically active secondary metabolites. The order Sordariales furthermore contains the highest diversity of thermophilic fungal species, with isolates present in seven different genera.


Families in the order Sordariales

Recent phylogenetic studies have aimed to contribute to the natural classification of this order. The most recent phylogenetic tree divides the order into eleven families, based on molecular data from four marker genes. * Bombardiaceae * Chaetomiaceae * Diplogelasinosporaceae *
Lasiosphaeriaceae The Lasiosphaeriaceae are a family of fungi in the Ascomycota, class Sordariomycetes. Genera *'' Anopodium'' *'' Apiosordaria'' *'' Apodospora'' *''Apodus'' *'' Arecacicola'' *'' Arniella'' *'' Arnium'' *'' Bombardia'' *'' Bombardioidea'' *'' C ...
* Lasiosphaeridaceae * Naviculisporaceae * Podosporaceae * Neoschizotheciaceae * Strattoniaceae (includes Strattonia) *
Sordariaceae The Sordariaceae are a family of perithecial fungi within the Sordariales order. The family includes the important model organism ''Neurospora crassa'' that is used in genetic research. Members of the family include the red bread molds in the ...
* Zygospermellaceae


Genera ''incertae sedis''

There are many genera in the Sordariomycetes that are not well known, and are of uncertain familial classification. These include: *'' Abyssomyces'' – 1 sp. *'' Acanthotheciella'' – 3 spp. *'' Arnium'' – 34 spp. *'' Ascolacicola'' – 1 sp. *'' Biconiosporella'' – 1 sp. *'' Bombardiella'' – 1 sp. *'' Camptosphaeria'' – 4 spp. *'' Coronatomyces'' – 1 sp. *'' Cuspidatispora'' – 1 sp. *'' Diffractella'' – 1 sp. *'' Emblemospora'' – 2 spp. *'' Eosphaeria'' – 2 spp. *'' Globosphaeria'' – 1 sp. *''
Isia Isia may refer to: * ''Isia'' (moth), a genus of moths in the subfamily Arctiinae * isiA, a photosynthetic protein * International Ski Instructors Association *Istituto Superiore per le Industrie Artistiche ISIA - Istituto Superiore per le Indust ...
'' – 2 spp. *'' Lockerbia'' – 2 spp. *'' Lunulospora'' – 2 spp. *'' Onygenopsis'' – 1 sp. *'' Periamphispora'' – 1 sp. *'' Phaeosporis'' – 2 spp. *'' Reconditella'' – 1 sp. *'' Rhexodenticula'' – 5 spp. *'' Rhexosporium'' – 1 sp. *'' Roselliniomyces'' – 7 spp. *'' Roselliniopsis'' – 7 spp. *'' Synaptospora'' – 5 spp. *'' Tripterosporella'' – 5 spp. *'' Utriascus'' – 1 sp. *'' Ypsilonia'' – 3 spp.


References

Ascomycota orders {{Sordariales-stub