Snowy Grouper
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The snowy grouper (''Hyporthodus niveatus'') is a species of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...
, a
grouper Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is ...
from the subfamily
Epinephelinae Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is ...
which is part of the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Serranidae The Serranidae are a large family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, in some ca ...
, which also includes the anthias and sea basses. It is found in the western Atlantic Ocean.


Description

The snowy grouper has a robust, compressed, oval body which has its deepest point at the origin of dorsal fin. Its standard length is 2.4 to 2.8 times longer than its depth. The preopercle has a serrated edge and the serrations at its angle are enlarged, where there is also often a spine bent upwards, clothed in skin and located on its lower edge immediately in front of its angle. The upper edge of the gill cover is notably convex. There are 11 spines and 13-15 soft rays in the
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
while the anal fin contains 3 spines and 9 soft rays. The membranes between the dorsal fin spines have deep notches. The
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
has a rear margin which is convex in juveniles of standard lengths less than and is straight or concave in adults. The overall colour is dark brown with a black margin to the spiny part of the dorsal fin. Juveniles are dark brown and are marked with obvious white spots arranged in vertical rows on the rear part of the head and on the body. These extend onto the dorsal fin and like adults they have a black margin to the spiny part of the dorsal fin while the caudal and
pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
s are yellow to clear. The upper part of the base of the caudal fin has a deep black saddle-like mark that extends underneath the lateral line. This species attains a maximum published
total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...
of , although are more commonly around , and a maximum published weight of .


Distribution

The snowy grouper occurs in the western Atlantic Ocean. It is found around
Bermuda ) , anthem = "God Save the King" , song_type = National song , song = "Hail to Bermuda" , image_map = , map_caption = , image_map2 = , mapsize2 = , map_caption2 = , subdivision_type = Sovereign state , subdivision_name = , es ...
and along the eastern coast of the United States from
Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...
south into the
Gulf of Mexico The Gulf of Mexico ( es, Golfo de México) is an ocean basin and a marginal sea of the Atlantic Ocean, largely surrounded by the North American continent. It is bounded on the northeast, north and northwest by the Gulf Coast of the United ...
and the
Caribbean Sea The Caribbean Sea ( es, Mar Caribe; french: Mer des Caraïbes; ht, Lanmè Karayib; jam, Kiaribiyan Sii; nl, Caraïbische Zee; pap, Laman Karibe) is a sea of the Atlantic Ocean in the tropics of the Western Hemisphere. It is bounded by Mexico ...
. Its range extends along the coasts of
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
and South America from Mexico to the
Valdes Peninsula The Valdes Peninsula (Spanish: ''Península Valdés'') is a peninsula into the Atlantic Ocean in the Biedma Department of north-east Chubut Province, Argentina. Around in size (not taking into account the isthmus of Carlos Ameghino which connects ...
in Argentina. It has been recorded as far north as
Massachusetts Massachusetts (Massachusett: ''Muhsachuweesut Massachusett_writing_systems.html" ;"title="nowiki/> məhswatʃəwiːsət.html" ;"title="Massachusett writing systems">məhswatʃəwiːsət">Massachusett writing systems">məhswatʃəwiːsət'' En ...
.


Habitat and biology

The snowy grouper is found at depths of between over rock substrates in offshore waters, although they are most common between . The juveniles are found in shallower, coastal waters and are frequently found along the northeastern coast of the United States. Like most groupers this species is predatory and prey items recorded for adults include fishes, gastropods, cephalopods, and brachyuran crustaceans. They are
protogynous hermaphrodite Sequential hermaphroditism (called dichogamy in botany) is a type of hermaphroditism that occurs in many fish, gastropods, and plants. Sequential hermaphroditism occurs when the individual changes its sex at some point in its life. In particular, ...
s and form aggregations for
spawning Spawn is the eggs and sperm released or deposited into water by aquatic animals. As a verb, ''to spawn'' refers to the process of releasing the eggs and sperm, and the act of both sexes is called spawning. Most aquatic animals, except for aquat ...
. Females attain sexual maturity at ages between 3 and 5 years old and total lengths of , changing to males 3 to 4 years after attaining sexual maturity as females. Off North America this species spawns from May to June and a female may lay more than 2 million eggs in a season.


Taxonomy

The snowy grouper was first formally described as ''Serranus niveatus'' in 1828 by the French
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, and d ...
Achille Valenciennes (1794-1865) with the type locality given as Brazil. In 1861
Theodore Nicholas Gill Theodore Nicholas Gill (March 21, 1837 – September 25, 1914) was an American ichthyologist, mammalogist, malacologist and librarian. Career Born and educated in New York City under private tutors, Gill early showed interest in natural histo ...
described a new species ''Hyporthodus flavicauda'' and placed it in its own
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...
genus, ''
Hyporthodus ''Hyporthodus'' is a genus of marine ray-finned fish, groupers from the subfamily Epinephelinae, part of the family Serranidae The Serranidae are a large family of fishes belonging to the order Perciformes. The family contains about 450 speci ...
'' although this was later shown to be a synonym of ''Serranus niveatus'', this means that ''Serranus niveatus'' is the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
of this genus.


Utilisation

The snowy grouper is targeted by commercial fisheries in the United States, in Central America and off South America using hook-and-line, bottom longlines, traps and trawls. In United States waters the snowy grouper may be fished for from 1 January to 31 August but as its management includes an annual catch limit, the fishery may be closed if this limit has been met or is projected to be met. In 2019 the fishery was closed on 3 August.


Sources

{{Taxonbar, from=Q307102 snowy grouper Fish of the Eastern United States Fish of the Western Atlantic snowy grouper Taxonomy articles created by Polbot