Skorpiovenator Bustingorryi
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''Skorpiovenator'' ("scorpion hunter") is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
abelisaurid Abelisauridae (meaning "Abel's lizards") is a family (or clade) of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are found ...
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
(
Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
to
Turonian The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded by t ...
)
Huincul Formation The Huincul Formation is a geologic formation of Late Cretaceous ( Early Cenomanian to Late Turonian) age of the Neuquén Basin that outcrops in the Mendoza, Río Negro and Neuquén Provinces of northern Patagonia, Argentina.
of
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
. It is one of the most complete and informative abelisaurids yet known, described from a nearly complete and articulated skeleton.


Description

The preserved length of the excavated ''Skorpiovenator'' skeleton from the
premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has b ...
to the 12th caudal vertebra is . It was estimated to have grown up to in length. In 2010,
Gregory S. Paul Gregory Scott Paul (born December 24, 1954) is an American freelance researcher, author and illustrator who works in paleontology, and more recently has examined sociology and theology. He is best known for his work and research on theropod dino ...
gave larger estimations of and .Paul, G.S. (2010) ''The Princeton Field Guide to Dinosaurs'', Princeton University Press p. 81 In 2016, a similar size to the original estimate at has been estimated. It had short, stubby, near-useless arms, but strong legs with powerful thighs and sturdy shins over which its large body was balanced.


Skull

''Skorpiovenators skull was short, stout and covered in the ridges, furrows, tubercles and bumpy nodules that are scattered over the heads of most
abelisaurid Abelisauridae (meaning "Abel's lizards") is a family (or clade) of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are found ...
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
s. It is craniocaudally short, similar to ''
Carnotaurus ''Carnotaurus'' (; ) is a genus of theropod dinosaur that lived in South America during the Late Cretaceous period, probably sometime between 71 and 69 million years ago. The only species is ''Carnotaurus sastrei''. Known from a single well-p ...
'', and is shorter and deeper than the skulls of '' Abelisaurus'' and ''
Majungasaurus ''Majungasaurus'' (; ) is a genus of abelisaurid theropod dinosaur that lived in Madagascar from 70 to 66 million years ago, at the end of the Cretaceous Period (geology), Period, making it one of the last known non-avian dinosaurs that went ex ...
''. Notably, the maxilla and lacrimal of ''Skorpiovenator'' are wider than in the corresponding bones of the remaining abelisaurids. ''Skorpiovenator'' had 19 maxillary teeth, which is more than any other known abelisaurid. The shape of the crowns of the teeth is similar to other abelisaurids, and exhibit enamel wrinkles and marginal serrations.


Discovery and naming

The type specimen was described and named by Canale, Scanferla, Agnolin, and Novas in 2009 (though the paper was released as an advanced publication online in 2008). The name ''Skorpiovenator bustingorryi'' is derived from the Greek and Latin for "scorpion hunter," due to the abundant scorpions present at the dig site, and the specific name honors Manuel Bustingorry, who owned the farm where the specimen was found. The describers have defined a new name Brachyrostra for a
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
, to which ''Skorpiovenator'' belonged. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
, ''Skorpiovenator bustingorryi'', is known from a single, nearly complete skeleton (MMCH-PV 48K) missing only sections of the tail and the majority of the forelimbs. The specimen was recovered from the lower part of the
Huincul Formation The Huincul Formation is a geologic formation of Late Cretaceous ( Early Cenomanian to Late Turonian) age of the Neuquén Basin that outcrops in the Mendoza, Río Negro and Neuquén Provinces of northern Patagonia, Argentina.
in
Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and gl ...
, dating to the late
Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
stage, about 95 million years ago. It would have lived alongside other carnivorous dinosaurs such as the carcharodontosaurid ''
Mapusaurus ''Mapusaurus'' () was a giant carcharodontosaurid carnosaurian dinosaur from the early Late Cretaceous (early Turonian stage), approximately 93.9 to 89.6 million years ago, of what is now Argentina. Discovery ''Mapusaurus'' was excavated be ...
'' and another abelisaurid, ''
Ilokelesia ''Ilokelesia'' is an extinct genus of abelisaurid theropod,Coria, R.A.; Salgado, L. & Calvo, J.O. (1991) "Primeros restos de dinosaurios Theropoda del Miembro Huincul, Formación Río Limay (Cretácico Tardío Presenoniano), Neuquén, Argentina." ...
''. The remains are deposited in the
Ernesto Bachmann Paleontological Museum The Ernesto Bachmann Paleontological Museum (''Museo Municipal Paleontológico, Arqueológico e Histórico "Ernesto Bachmann"'' or Archaeological, Paleontological and Historical Municipal Museum) (MEB) in Villa El Chocón, Neuquén Province, Arg ...
of
Villa El Chocón Villa El Chocón is a village and municipality in Neuquén Province in southwestern Argentina. The village was initially created to house the workmen building the Ezequiel Ramos Mexía dam (colloquially known as El Chocón Dam) on the Limay River. ...
,
Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and gl ...
,
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
.


Classification

In 2008, Canale ''et al.'' published a phylogenetic analysis focusing on the South American carnotaurines. In their results, they found that all South American forms (including ''Skorpiovenator'') grouped together as a sub-clade of Carnotaurinae, which they named Brachyrostra, meaning "short snouts". They defined the
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
Brachyrostra as "all the abelisaurids more closely related to ''Carnotaurus sastrei'' than to ''Majungasaurus crenatissimus''."Canale, J.I., Scanferla, C.A., Agnolin, F., and Novas, F.E. (2008). "New carnivorous dinosaur from the Late Cretaceous of NW Patagonia and the evolution of abelisaurid theropods." ''Naturwissenschaften''. .


See also

*
Timeline of ceratosaur research This timeline of ceratosaur research is a chronological listing of events in the history of paleontology focused on the ceratosaurs, a group of relatively primitive, often horned, predatory theropod dinosaurs that became the apex predators of ...


References

{{Portal bar, Dinosaurs, Cretaceous Brachyrostrans Late Cretaceous dinosaurs of South America Cenomanian life Cretaceous Argentina Fossils of Argentina Huincul Formation Fossil taxa described in 2009 Taxa named by Fernando Novas