Sinusonasus Magnodens
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''Sinusonasus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s from the Early
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
Period, recovered from the Yixian Formation. It lived in what is now the
Liaoning Province Liaoning () is a coastal provinces of China, province in Northeast China that is the smallest, southernmost, and most populous province in the region. With its capital at Shenyang, it is located on the northern shore of the Yellow Sea, and i ...
of
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
. ''Sinusonasus'' was a theropod, specifically a troodontid dinosaur. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
, ''Sinusonasus magnodens'', was named and described by Xu Xing and Wang Xiaolin in 2004.Xu X. and Wang X.-l. 2004. "A New Troodontid (Theropoda: Troodontidae) from the Lower Cretaceous Yixian Formation of Western Liaoning, China". ''Acta Geologica Sinica'' 78(1): 22-26 The generic name, derived from
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''sinus'', "wave", and ''nasus'', "nose", refers to the
sinusoid A sine wave, sinusoidal wave, or just sinusoid is a mathematical curve defined in terms of the ''sine'' trigonometric function, of which it is the graph. It is a type of continuous wave and also a smooth periodic function. It occurs often in ma ...
form, in lateral view, of the nasals. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
means "big-toothed" from Latin ''magnus'', "large" and ''dens'', "tooth". In a later publication the species is referred to as "Sinucerasaurus" but this is a junior objective synonym. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
, IVPP V 11527, was found in the Lujiatun Member of the Yixian Formation, dating from the
Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.9 ± 2 Ma and 129.4 ± 1.5 Ma (million years ago). The Hauterivian is preceded by the Va ...
. It consists of a partial skeleton including skull and lower jaw fragments and partial tail, pelvis and hindlimbs. The fossil is compressed and partially articulated. ''Sinusonasus'' is a small troodontid. In 2010, Gregory S. Paul estimated its length at one metre, its weight at 2.5 kilogrammes.Paul, G.S., 2010, ''The Princeton Field Guide to Dinosaurs'', Princeton University Press p. 141 The
femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with ...
is 141 millimetres long. In 2004, several distinguishing traits were established. An interantorbital channel, connecting the antorbital fenestrae at each skull side, is lacking. The nasal bone has an undulating profile. The middle
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
ry teeth are rather large. The
chevron Chevron (often relating to V-shaped patterns) may refer to: Science and technology * Chevron (aerospace), sawtooth patterns on some jet engines * Chevron (anatomy), a bone * ''Eulithis testata'', a moth * Chevron (geology), a fold in rock lay ...
s on the rear
caudal vertebrae The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
are so long, measured from the front to the back, that they connect, forming a continuous plate at the underside of the tail. The neck of the thighbone is elongated. More generally, the head is relatively short, equalling 77% of the length of the thighbone. There are at least nineteen maxillary teeth per side. The front teeth are not serrated: those more to the rear only have denticles at the trailing edge. Five
sacral vertebra The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30. The sacrum situates at the upper, back part ...
e are present; the tail probably consisted of about thirty vertebrae. The pubic bone probably pointed obliquely to the front. The
ischium The ischium () form ...
is elongated. ''Sinusonasus'' has a long lower leg, indicating a good running capacity. The foot is "
arctometatarsal An arctometatarsalian organism is one in which the proximal part of the middle metatarsal is pinched between the surrounding metatarsals. The trait appears to be highly homoplastic, common in certain sorts of dinosaurs accustomed to running (among ...
", with a 'pinched' upper third metatarsal. The second metatarsal is distinctly shorter than the fourth. The second toe bears a retractable 'sickle claw'. ''Sinusonasus'' was in 2004 placed in the Troodontidae. It was presumed to have had a rather derived position, despite living in the
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
. This was by the describing authors not interpreted as an indication for a long
ghost lineage A ghost lineage is a hypothesized ancestor in a species lineage that has left no fossil evidence yet can be inferred to exist because of gaps in the fossil record or genomic evidence. The process of determining a ghost lineage relies on fossilized ...
, troodontids developing earlier during the Jurassic than had been thought, but explained by rapid evolutionary change after a Cretaceous origin of the group.


See also

*
Timeline of troodontid research This timeline of troodontid research is a chronological listing of events in the History of paleontology, history of paleontology focused on the troodontids, a group of bird-like theropod dinosaurs including animals like ''Troodon''. Troodontid re ...


References

{{Taxonbar, from=Q690483 Early Cretaceous dinosaurs of Asia Yixian fauna Troodontids Fossil taxa described in 2004 Taxa named by Xu Xing