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''Sinoconodon'' is an extinct genus of
mammaliamorph Mammaliamorpha is a clade of cynodonts. It contains the clades Tritylodontidae and Mammaliaformes, as well as a few genera that do not belong to either of these groups. The family Tritheledontidae has also been placed in Mammaliamorpha by some ph ...
s that appears in the
fossil record A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...
of the
Lufeng Formation The Lufeng Formation (formerly Lower Lufeng Series) is a Lower Jurassic sedimentary rock formation found in Yunnan, China. It has two units: the lower Dull Purplish Beds/Shawan Member are of Hettangian age, and Dark Red Beds/Zhangjia'ao Member ar ...
of China in the
Sinemurian In the geologic timescale, the Sinemurian is an age and stage in the Early or Lower Jurassic Epoch or Series. It spans the time between 199.3 ± 2 Ma and 190.8 ± 1.5 Ma (million years ago). The Sinemurian is preceded by the Hettangian and is ...
stage of the
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-J ...
period, about 193 million years ago. While sharing many
plesiomorphic In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, and ...
traits with other non-
mammaliaform Mammaliaformes ("mammalian forms") is a clade that contains the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts. It is defined as the clade originating from the most recent com ...
cynodont The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety ...
s, it possessed a special, secondarily evolved jaw joint between the
dentary In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...
and the
squamosal bone The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. In most tetrapods, the squamosal and quadratojugal bones form the cheek series of the skull. The bone forms an ancestral co ...
s, which in more derived taxa would replace the primitive
tetrapod Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct theraps ...
one between the
articular The articular bone is part of the lower jaw of most vertebrates, including most jawed fish, amphibians, birds and various kinds of reptiles, as well as ancestral mammals. Anatomy In most vertebrates, the articular bone is connected to two oth ...
and
quadrate bone The quadrate bone is a skull bone in most tetrapods, including amphibians, sauropsids (reptiles, birds), and early synapsids. In most tetrapods, the quadrate bone connects to the quadratojugal and squamosal bones in the skull, and forms upper ...
s. The presence of a dentary-squamosal joint is a trait historically used to define mammals.


Description

This animal had skull of which suggest a presacral body length of and weight about due to the similar parameters to the
European hedgehog The European hedgehog (''Erinaceus europaeus''), also known as the West European hedgehog or common hedgehog, is a hedgehog species native to Europe from Iberia and Italy northwards into Scandinavia and westwards into the British Isles.Harris, S. ...
. ''Sinoconodon'' closely resembled early mammaliaforms like ''
Morganucodon ''Morganucodon'' (" Glamorgan tooth") is an early mammaliaform genus that lived from the Late Triassic to the Middle Jurassic. It first appeared about 205 million years ago. Unlike many other early mammaliaforms, ''Morganucodon'' is well represe ...
'', but it is regarded as more basal, differing substantially from ''Morganucodon'' in its dentition and growth habits. Like most other non-mammalian tetrapods, such as
reptiles Reptiles, as most commonly defined are the animals in the Class (biology), class Reptilia ( ), a paraphyletic grouping comprising all sauropsid, sauropsids except birds. Living reptiles comprise turtles, crocodilians, Squamata, squamates (lizar ...
and
amphibians Amphibians are four-limbed and ectothermic vertebrates of the class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial, fossorial, arbore ...
, it was
polyphyodont A polyphyodont is any animal whose teeth are continually replaced. In contrast, diphyodonts are characterized by having only two successive sets of teeth. Polyphyodonts include most toothed fishes, many reptiles such as crocodiles and geckos, and ...
, replacing many of its teeth throughout its lifetime, and it seems to have grown slowly but continuously until its death. It was thus somewhat less mammal-like than mammaliaforms such as
morganucodont Morganucodonta ("Glamorgan teeth") is an extinct order of basal Mammaliaformes, a group including crown-group mammals (Mammalia) and their close relatives. Their remains have been found in Southern Africa, Western Europe, North America, India an ...
s and
docodont Docodonta is an order of extinct mammaliaforms that lived during the Mesozoic, from the Middle Jurassic to Early Cretaceous. They are distinguished from other early mammaliaforms by their relatively complex molar teeth, from which the order get ...
s. The combination of basal tetrapod and mammalian features makes it a unique
transitional fossil A transitional fossil is any fossilized remains of a life form that exhibits traits common to both an ancestral group and its derived descendant group. This is especially important where the descendant group is sharply differentiated by gross a ...
.Mammals of the Mesozoic: The least mammal-like mammals
/ref>


Taxonomy

''Sinoconodon'' was named by Patterson and Olson in 1961. Its type is ''Sinoconodon rigneyi''. It was assigned to Triconodontidae by Patterson and Olson in 1961; to Triconodonta by Jenkins and Crompton in 1979; to Sinoconodontidae by Carroll in 1988; to Mammaliamorpha by Wible in 1991; to Mammalia by Luo and Wu in 1994; to
Mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
ia by Kielan-Jaworowska ''et al.'' in 2004; and to
Mammaliaformes Mammaliaformes ("mammalian forms") is a clade that contains the crown group mammals and their closest Extinction, extinct relatives; the group adaptive radiation, radiated from earlier probainognathian cynodonts. It is defined as the clade origin ...
by Luo ''et al.'' in 2001 and Bi ''et al.'' in 2014.


Phylogeny


References


External links


Mammaliformes from Palaeos
*http://eol.org/pages/10575421/names?all=1 *http://palaeos.com/vertebrates/mammaliformes/mammaliformes.html#Mammaliformes Prehistoric prozostrodonts Prehistoric cynodont genera Transitional fossils Jurassic synapsids Jurassic animals of Asia Sinemurian life Fossils of China Fossil taxa described in 1961 Taxa named by Bryan Patterson Taxa named by Everett C. Olson {{Paleo-therapsid-stub