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Mammaliaformes
Mammaliaformes ("mammalian forms") is a clade that contains the crown group mammals and their closest Extinction, extinct relatives; the group adaptive radiation, radiated from earlier probainognathian cynodonts. It is defined as the clade originating from the most recent common ancestor of Morganucodonta and the crown group mammals; the latter is the clade originating with the most recent common ancestor of extant Monotremata, Marsupialia, and Placentalia. Besides Morganucodonta and the crown group mammals, Mammaliaformes includes Docodonta and ''Hadrocodium'' as well as the Triassic ''Tikitherium'', the earliest known member of the group. Mammaliaformes is a term of phylogenetic nomenclature. In contrast, the assignment of organisms to Mammalia has traditionally been founded on traits and, on this basis, Mammalia is slightly more inclusive than Mammaliaformes. In particular, trait-based taxonomy generally includes ''Adelobasileus'' and ''Sinoconodon'' in Mammalia, though they fa ...
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Docodonta
Docodonta is an Order (biology), order of extinct Mammaliaformes, mammaliaforms that lived during the Mesozoic, from the Middle Jurassic to Early Cretaceous. They are distinguished from other early Mammaliaformes, mammaliaforms by their relatively complex Molar (tooth), molar teeth, from which the order gets its name. Until recently, Docodonta were represented primarily by teeth and jaws found across former Laurasia, (modern-day North America, Europe, and Asia). However, recent discoveries in China include some exceptionally well preserved, almost complete body fossils. Description Skeletal traits Docodonts have a long and low mandible (lower jaw), formed primarily by the tooth-bearing dentary bone. The dentary connects to the cranium via a joint with the Squamosal bone, squamosal, a connection which is strengthened relative to earlier mammaliaforms. The other bones in the jaw, known as postdentary elements, are still connected to the dentary and lie within a groove (the postd ...
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Cynodont
The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety of lifestyles, including carnivory and herbivory. Mammals are cynodonts, as are their extinct ancestors and close relatives, having evolved from advanced probainognathian cynodonts during the Late Triassic. All other cynodont lines went extinct, with the last known non-mammalian cynodont group, the Tritylodontidae, having its youngest records in the Early Cretaceous. Description Early cynodonts have many of the skeletal characteristics of mammals. The teeth were fully differentiated and the braincase bulged at the back of the head. Outside of some crown-group mammals (notably the therians), all cynodonts probably laid eggs. The temporal fenestrae were much larger than those of their ancestors, and the widening of the zygomatic arch in ...
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Castorocauda
''Castorocauda'' is an extinct, semi-aquatic, superficially otter-like genus of docodont mammaliaforms with one species, ''C. lutrasimilis''. It is part of the Yanliao Biota, found in the Daohugou Beds of Inner Mongolia, China dating to the Middle to Late Jurassic. It was part of an explosive Middle Jurassic radiation of Mammaliaformes moving into diverse habitats and niches. Its discovery in 2006, along with the discovery of other unusual mammaliaforms, disproves the previous hypothesis of Mammaliaformes remaining evolutionarily stagnant until the extinction of the non-avian dinosaurs. Weighing an estimated , ''Castorocauda'' is the largest known Jurassic mammaliaform. It is the earliest known mammaliaform with aquatic adaptations or a fur pelt. It was also adapted for digging, and its teeth are similar to those of seals and Eocene whales, collectively suggesting it behaved similarly to the modern-day platypus and river otters and ate primarily fish. It lived in a wet, seaso ...
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Haramiyida
Haramiyida ("thief" from Arabic الحرامية (al ḥarāmiyah), "thief, bandit") is a possibly polyphyletic order of mammaliaform cynodonts or mammals of controversial taxonomic affinites. Their teeth, which are by far the most common remains, resemble those of the multituberculates. However, based on ''Haramiyavia'', the jaw is less derived; and at the level of evolution of earlier basal mammals like ''Morganucodon'' and ''Kuehneotherium'', with a groove for ear ossicles on the dentary. If they are early multituberculates, they would be the longest lived mammalian clade of all time. However, a more recent study in November 2015 may dispute this and suggested the Haramiyida were not crown mammals, but were part of an earlier offshoot of mammaliaformes instead. It is also disputed whether the Late Triassic species are closely related to the Jurassic and Cretaceous members belonging to Euharamiyida/Eleutherodontida, as some phylogenetic studies recover the two groups as unrelate ...
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Morganucodonta
Morganucodonta ("Glamorgan teeth") is an extinct order of basal Mammaliaformes, a group including crown-group mammals (Mammalia) and their close relatives. Their remains have been found in Southern Africa, Western Europe, North America, India and China. The morganucodontans were probably insectivorous and nocturnal, though like eutriconodonts some species attained large sizes and were carnivorous. Nocturnality is believed to have evolved in the earliest mammals in the Triassic (called the nocturnal bottleneck) as a specialisation that allowed them to exploit a safer, night-time niche, while most larger predators were likely to have been active during the day (though some dinosaurs, for example, were nocturnal as well). Anatomy and biology Morganucodontans had a double jaw articulation made up of the dentary-squamosal joint as well as a quadrate-articular one. This implies that they also retained one of their postdentary bones: the articular. There is a trough at the back of t ...
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Hadrocodium
''Hadrocodium wui'' is an extinct mammaliaform that lived during the Sinemurian stage of the Early Jurassic approximately in the Lufeng Formation of the Lufeng Basin in what is now the Yunnan province in south-western China (, paleocoordinates ). It is considered as the closest relative of the class Mammalia. The fossil of this mouse-like, paper-clip sized animal was discovered in 1985 but was then interpreted as a juvenile morganucodontid. ''Hadrocodium'' remained undescribed until 2001; since then its large brain and advanced ear structure have greatly influenced the interpretation of the earliest stages of mammalian evolution, as these mammalian characters could previously be traced only to some . ''Hadrocodium'' is known only from a skull long, and the body mass is estimated to have been , making it one of the smallest Mesozoic mammaliaforms (in contrast, the related ancestral mammal would have been bigger, about ). The name ''Hadrocodium'' alludes to its large cranial ca ...
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Mammal
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles (including birds) from which they diverged in the Carboniferous, over 300 million years ago. Around 6,400 extant species of mammals have been described divided into 29 orders. The largest orders, in terms of number of species, are the rodents, bats, and Eulipotyphla (hedgehogs, moles, shrews, and others). The next three are the Primates (including humans, apes, monkeys, and others), the Artiodactyla ( cetaceans and even-toed ungulates), and the Carnivora (cats, dogs, seals, and others). In terms of cladistics, which reflects evolutionary history, mammals are the only living members of the Synapsida (synapsids); this clade, together with Saur ...
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Mammalia
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles (including birds) from which they diverged in the Carboniferous, over 300 million years ago. Around 6,400 extant species of mammals have been described divided into 29 orders. The largest orders, in terms of number of species, are the rodents, bats, and Eulipotyphla (hedgehogs, moles, shrews, and others). The next three are the Primates (including humans, apes, monkeys, and others), the Artiodactyla ( cetaceans and even-toed ungulates), and the Carnivora (cats, dogs, seals, and others). In terms of cladistics, which reflects evolutionary history, mammals are the only living members of the Synapsida (synapsids); this clade, together w ...
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Morganucodon
''Morganucodon'' (" Glamorgan tooth") is an early mammaliaform genus that lived from the Late Triassic to the Middle Jurassic. It first appeared about 205 million years ago. Unlike many other early mammaliaforms, ''Morganucodon'' is well represented by abundant and well preserved (though in the vast majority of cases disarticulated) material. Most of this comes from Glamorgan in Wales (''Morganucodon watsoni''), but fossils have also been found in Yunnan Province in China (''Morganucodon oehleri'') and various parts of Europe and North America. Some closely related animals (''Megazostrodon'') are known from exquisite fossils from South Africa. The name comes from a Latinization of ''Morganuc'', the name for South Glamorgan in the Domesday Book, the county of Wales where it was discovered by Walter Georg Kühne,Walter G. Kühne, "On a Triconodont tooth of a new pattern from a Fissure-filling in South Glamorgan", ''Proceedings of the Zoological Society of London'', volume 119 (1 ...
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Probainognathia
Probainognathia is one of the two major subgroups of the clade Eucynodontia, the other being Cynognathia. The earliest forms were carnivorous and insectivorous, though some groups eventually also evolved herbivorous diets. The earliest and most basal probainognathian is the Middle Triassic (Anisian) aged ''Lumkuia'', from South Africa, though probainognathians would not become prominent until the mid Norian stage of the Late Triassic. Three groups survived the extinction at the end of Triassic: Tritheledontidae and Tritylodontidae, which both survived until the Jurassic—the latter even into the Cretaceous (''Montirictus'' and '' Xenocretosuchus'')—and Mammaliaformes, which includes the mammals. Phylogeny Below is a cladogram from Ruta, Botha-Brink, Mitchell and Benton (2013) showing one hypothesis of cynodont relationships: See also * Evolution of mammals * List of prehistoric mammals This is an incomplete list of prehistoric mammals. It does not include extant mam ...
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Tikitherium
''Tikitherium'' is an extinct genus of mammaliaforms from the Late Triassic. It is thought to be an insectivore and a close relative to Docodonta. ''Tikitherium'' refers to Tiki, the village located near the Tiki Formation where the specimen was found, and ''therium'' is Greek for “Beast”. The species was named ''copei'' in honor of Edward Drinker Cope for his pioneering discoveries towards understanding mammalian molars.Datta, P.M. 2005. Earliest Mammal with Transversely Expanded Upper Molar from the Late Triassic (Carnian) Tiki Formation, South Rewa Gondwana Basin, India. Journal of Vertebrate Paleontology. 25(1):200-207. History and discovery ''Tikitherium copei'' was first described by Datta in 2005. The first and only specimen is an upper left molar that was discovered in the lower part of the Tiki Formation, located in the South Rewa Gondwana Basin, India. Both the genus ''Tikitherium'' and the species ''copei'' were named by Datta in 2005. The currently only known ...
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Triassic
The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic Period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic. The Triassic began in the wake of the Permian–Triassic extinction event, which left the Earth's biosphere impoverished; it was well into the middle of the Triassic before life recovered its former diversity. Three categories of organisms can be distinguished in the Triassic record: survivors from the extinction event, new groups that flourished briefly, and other new groups that went on to dominate the Mesozoic Era. Reptiles, especially archosaurs, were the chief terrestrial vertebrates during this time. A specialized subgroup of archo ...
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