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''Silvisaurus'', from the Latin silva "woodland" and Greek sauros "lizard", is a nodosaurid ankylosaur from the
Early Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early ...
to Late Cretaceous period.


Discovery and species

A fossil of the species was discovered in the fifties by rancher Warren H. Condray of Wells, KS. He notified senator Frank Carlson who directed him to the chancellor of the University of Kansas,
Franklin David Murphy Franklin David Murphy (January 29, 1916 – June 16, 1994) was an American administrator, educator, and medical doctor. During his life, he served as Chancellor of the University of Kansas (KU) and Chancellor of the University of California, Los A ...
. Murphy sent the preparator of the paleontology of vertebrates department of the natural history museum of the university, Russell R. Camp, to investigate the matter. In July 1955 Camp, with help from Condray, recovered the skeleton of a dinosaur. It was further prepared by Camp and Glenn H. Marihugh. In 1961 the find was described and named by
Theodore H. Eaton Theodore may refer to: Places * Theodore, Alabama, United States * Theodore, Australian Capital Territory * Theodore, Queensland, a town in the Shire of Banana, Australia * Theodore, Saskatchewan, Canada * Theodore Reservoir, a lake in Saskatche ...
jr., also from the University of Kansas, as the type species ''Silvisaurus condrayi''. The generic name is derived from Latin ''silva'', "wood", in reference to the probably densely forested habitat of the animal. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
honours Condray.Eaton, T.H., Jr. 1960.
A new armored dinosaur from the Cretaceous of Kansas
. ''The University of Kansas Paleontological Contributions: Vertebrata'' 8: 1-24.
To date, ''Silvisaurus'' includes only the type species. The holotype, KU 10296, was found in exposures of the
Terracota Clay Member Terracotta, terra cotta, or terra-cotta (; ; ), in its material sense as an earthenware substrate, is a clay-based unglazed or glazed ceramic where the fired body is porous. In applied art, craft, construction, and architecture, terra ...
of the Dakota Formation (late Albian-early
Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
) in Kansas, and consists of an incomplete skeleton with skull. It includes the mandible, eight neck vertebrae, ten dorsal vertebrae, a
sacrum The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30. The sacrum situates at the upper, back part ...
of six sacral vertebrae, three tail vertebrae, a left pubis fragment, the lower end of the right femur, and a toe phalanx. Additionally disarticulated plates and spikes from the body armour were discovered. The condition of the fossil was poor as the bones had been exposed at the bottom of a dry riverbed and had been weathered and trampled by cattle. Some elements were only present as impressions or natural casts.


Paleobiology

Based on the remains, the animal is estimated to have been approximately in length. Its skull measures in length and is wide. The bony secondary palate is poorly developed in ''Silvisaurus'', the dentary includes at least twenty-five teeth, the basal tubera of the basioccipital are bulbous, and each premaxilla holds eight to nine teeth. The presence of teeth at the front of the jaw suggests that this may have been a relatively primitive nodosaur, since most later forms had a toothless beak instead. In addition to the usual rounded and polygonal osteoderms, ''Silvisaurus'' may have also sported bony spines on its shoulders and tail. The head contained large air passages, which may have been used for loud vocalisations, presumably for communication.


Classification

This taxon represents a relatively primitive nodosaurid, and Vickaryous ''et al.'' (2004) have stated that "'' Sauropelta edwardsorum'', ''Silvisaurus condrayi'', and ''
Pawpawsaurus ''Pawpawsaurus'', meaning "Pawpaw Lizard", is a nodosaurid ankylosaur from the Cretaceous (late Albian) of Tarrant County, Texas, discovered in May 1992. The only species yet assigned to this taxon, ''Pawpawsaurus campbelli,'' is based on a ...
campbelli'' form a
basal Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''. Science * Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure * Basal (medicine), a minimal level that is nec ...
polytomy nested deep to ''
Cedarpelta ''Cedarpelta'' is a extinct genus of basal ankylosaurid dinosaur from Utah that lived during the Late Cretaceous period (Cenomanian to lower Turonian stage, 98.2 to 93 Ma) in what is now the Mussentuchit Member of the Cedar Mountain Formation. Th ...
''."


See also

* Timeline of ankylosaur research


References

*Vickaryous, Maryanska, and Weishampel 2004. Chapter Seventeen: Ankylosauria. in The Dinosauria (2nd edition), Weishampel, D. B., Dodson, P., and Osmólska, H., editors. University of California Press. {{Taxonbar, from=Q133169 Nodosaurids Late Cretaceous dinosaurs of North America Fossil taxa described in 1960 Paleontology in Kansas Ornithischian genera