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''Silesaurus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
silesaurid Silesauridae is an extinct family of Triassic dinosauriforms. It is most commonly considered to be a clade of non-dinosaur dinosauriforms, and the sister group of dinosaurs. Some studies have instead suggested that most or all silesaurids compri ...
dinosauriform from the
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
, of what is now
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...
.


Discovery

Fossilized remains of ''Silesaurus'' have been found in the
Keuper Claystone The Keuper is a lithostratigraphic unit (a sequence of rock strata) in the subsurface of large parts of west and central Europe. The Keuper consists of dolomite, shales or claystones and evaporites that were deposited during the Middle and Late T ...
in Krasiejów near
Opole Opole (; german: Oppeln ; szl, Ôpole) ; * Silesian: ** Silesian PLS alphabet: ''Ôpole'' ** Steuer's Silesian alphabet: ''Uopole'' * Silesian German: ''Uppeln'' * Czech: ''Opolí'' * Latin: ''Oppelia'', ''Oppolia'', ''Opulia'' is a city loc ...
,
Silesia Silesia (, also , ) is a historical region of Central Europe that lies mostly within Poland, with small parts in the Czech Republic and Germany. Its area is approximately , and the population is estimated at around 8,000,000. Silesia is split ...
, Poland, which is also the origin of its name. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
, ''Silesaurus opolensis'', was described by
Jerzy Dzik Jerzy Dzik (born 25 February 1950) is a Polish paleontologist. He has described many species, genera, and families of conodonts, including the order Ozarkodinida (in 1976). In 2003, he described the dinosauriform '' Silesaurus'', from the Trias ...
in 2003. It is known from some 20 skeletons, making it one of the best-represented species of early dinosauriformes.


Description

''Silesaurus'' measured approximately in length. Lightly built, it was probably a fast and agile animal with an active lifestyle. The snout was narrow with forward-pointing nostrils, and the large orbits likely provided ''Silesaurus'' with acute vision. Initially, ''Silesaurus'' was thought to be strictly
herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
, but later research on
coprolite A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name is de ...
contents indicates that it may have been insectivorous, feeding on insects such as the
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
'' Triamyxa''. The teeth of the animal were small, conical, and serrated, and were distributed irregularly in its jaws. The tip of the
dentary In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...
had no teeth, and evidence suggests that it was covered by a keratinous
beak The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food ...
.


Classification

Scientists think that ''Silesaurus'' was not a
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
, but rather a dinosauriform. Dinosaurian features lacking in ''Silesaurus'' include an enlarged deltopectoral crest (a muscle attachment on the
humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...
), and
epipophyses Epipophyses are bony projections of the cervical vertebrae found in archosauromorphs, particularly dinosaurs (including some basal birds). These paired processes sit above the postzygapophyses on the rear of the vertebral neural arch. Their morp ...
(enlarged
tendon A tendon or sinew is a tough, high-tensile-strength band of dense fibrous connective tissue that connects muscle to bone. It is able to transmit the mechanical forces of muscle contraction to the skeletal system without sacrificing its ability ...
attachment above the
postzygapophysis The articular processes or zygapophyses (Greek ζυγον = "yoke" (because it links two vertebrae) + απο = "away" + φυσις = "process") of a vertebra are projections of the vertebra that serve the purpose of fitting with an adjacent vertebr ...
) on the
cervical vertebrae In tetrapods, cervical vertebrae (singular: vertebra) are the vertebrae of the neck, immediately below the skull. Truncal vertebrae (divided into thoracic and lumbar vertebrae in mammals) lie caudal (toward the tail) of cervical vertebrae. In ...
. However, ''Silesaurus'' has some dinosaurian characteristics as well: * a brevis shelf (a bone surface on the ilium that functions as an attachment site for tail muscles) *
ischium The ischium () form ...
with a slender shaft *
femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with ...
with a reduced tuberosity that borders the
ligament A ligament is the fibrous connective tissue that connects bones to other bones. It is also known as ''articular ligament'', ''articular larua'', ''fibrous ligament'', or ''true ligament''. Other ligaments in the body include the: * Peritoneal li ...
of the
femoral head The femoral head (femur head or head of the femur) is the highest part of the thigh bone (femur). It is supported by the femoral neck. Structure The head is globular and forms rather more than a hemisphere, is directed upward, medialward, and a l ...
* a prominent
lesser trochanter The lesser trochanter is a conical posteromedial bony projection of the femoral shaft. it serves as the principal insertion site of the iliopsoas muscle. Structure The lesser trochanter is a conical posteromedial projection of the shaft of the fe ...
* an overlap of the ascending process of the
astragalus ''Astragalus'' is a large genus of over 3,000 species of herbs and small shrubs, belonging to the legume family Fabaceae and the subfamily Faboideae. It is the largest genus of plants in terms of described species. The genus is native to tempe ...
with the
tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...
* a concave proximal articular surface for the reception of the distal end of the
fibula The fibula or calf bone is a leg bone on the lateral side of the tibia, to which it is connected above and below. It is the smaller of the two bones and, in proportion to its length, the most slender of all the long bones. Its upper extremity is ...
on the
calcaneum In humans and many other primates, the calcaneus (; from the Latin ''calcaneus'' or ''calcaneum'', meaning heel) or heel bone is a bone of the tarsus of the foot which constitutes the heel. In some other animals, it is the point of the hock. St ...
As a result, alternative hypotheses place ''Silesaurus'' at or near the base of the
ornithischia Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek s ...
n dinosaurs. Other scientists propose a basal link between the basal
sauropodomorph Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lon ...
s and ornithischians. Systematic position after Nesbitt (2011):


Palaeobiology

Beetle
elytra An elytron (; ; , ) is a modified, hardened forewing of beetles (Coleoptera), though a few of the true bugs (Hemiptera) such as the family Schizopteridae are extremely similar; in true bugs, the forewings are called hemelytra (sometimes alterna ...
and other arthropod remains have been found in
coprolites A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name is de ...
which have been recovered in the vicinity of ''Silesaurus'' fossil occurrences. The size and shape of the coprolites combined with the peculiar beaked architecture of ''Silesauruss skull may suggest that this animal produced the coprolites and that it could have been wholly or partially insectivorous. It might have fed on insects and other small arthropods by
pecking Pecking is the action of a bird using their beak to search for food or otherwise investigate an object or area by tapping it. Pecking can also be used by a bird to attack or fight another bird. Pecking is frequently observed in chickens and other ...
, using the beak-like tip of the snout to accurately target prey in a way similar to modern birds. If so, this would represent the earliest known occurrence of this highly derived mode of feeding and have implications for the understanding of the evolutionary adaptations that would eventually lead up to the origin of dinosaurs.


Palaeoenvironment

''Silesaurus'' lived in a subtropical environment similar to the modern
Mediterranean basin In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and w ...
with alternating summer monsoons and dry winters. The animal shared its environment of extensive swamplands and
fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...
vegetation with a wealth of
invertebrates Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
as well as dipnoan and ganoid fishes,
temnospondyls Temnospondyli (from Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carbo ...
,
phytosaurs Phytosaurs (Φυτόσαυροι in greek) are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria. Phytosauria and Phytosauridae are often considered to be equivalen ...
and early
pterosaurs Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...
.John W. M. Jagt, Grzegorz Hebda, Sławomir Mitrus, Elena Jagt-Yazykova, Adam Bodzioch, Dorota Konietzko-Meier, Klaudia Kardynał, Kamil Gruntmejer. 2015. Field Guide, Conference Paper, European Association of Vertebrate Palaeontologists XIII Annual Meeting.


References

{{Taxonbar, from=Q2322754 Silesaurids Late Triassic reptiles of Europe Fossils of Poland Opole County Fossil taxa described in 2003