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The Sepsidae are a
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of
flies Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced ...
, commonly called the black scavenger flies or ensign flies. Over 300 species are described worldwide. They are usually found around dung or decaying plant and animal material. Many species resemble ants, having a "waist" and glossy black body. Many Sepsidae have a curious wing-waving habit made more apparent by dark patches at the wing end. Many species have a very wide distribution, reflecting the
coprophagous Coprophagia () or coprophagy () is the consumption of feces. The word is derived from the grc, κόπρος , "feces" and , "to eat". Coprophagy refers to many kinds of feces-eating, including eating feces of other species (heterospecifics), of ...
habit of most Sepsidae. Some species have been spread over large territories in association with livestock. Adult flies are found mostly on mammal
excrement Feces ( or faeces), known colloquially and in slang as poo and poop, are the solid or semi-solid remains of food that was not digested in the small intestine, and has been broken down by bacteria in the large intestine. Feces contain a relati ...
, including that of humans (less often on other rotting organic matter), where eggs are laid and
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e develop, and on nearby vegetation, carrion, fermenting tree sap, and shrubs and herbs. Many Sepsidae apparently play an important biological role as decomposers of mammal and other animal excrement. Some species may have a limited hygienic importance because of their association with human feces. Others are useful tools in
forensic entomology Forensic entomology is the scientific study of the colonization of a dead body by arthropods. This includes the study of insect types commonly associated with cadavers, their respective life cycles, their ecological presences in a given environme ...
.


Description

For terms, see
Morphology of Diptera Dipteran morphology differs in some significant ways from the broader morphology of insects. The Diptera is a very large and diverse order of mostly small to medium-sized insects. They have prominent compound eyes on a mobile head, and (at most) ...
.
Sepsids are slender flies that resemble minute, winged ants. They are usually black in color, sometimes lustrous, and sometimes with silvery hairs on the thorax. The head is rounded. Sepsids have one or more bristles at the posteroventral margin of the posterior spiracle of the thorax, a character that distinguishes the family from other acalyptrates. The postvertical bristles are divergent or sometimes absent. Up to three pairs of frontal bristles are seen. They have ocelli with ocellar bristles. Vibrissae and palpi are poorly developed. The front legs of the male often have extrusions, spurs, teeth, or other ornamentation. The tibia has a dorsal preapical bristle in most genera. The abdomen is usually constricted in the basal part. The larva is slender, tapering at the front end, and smooth except for ventral creeping welts. The larva is amphipneustic: it has two pairs of spiracles, one toward the head and one at the tail. The bulbous posterior end with its pair of spiracles distinguishes it from the larvae of other acalyptrates. The
pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...
is enclosed within a puparium.


Classification

Genera include: *''
Adriapontia ''Adriapontia'' is a genus of flies in the family Sepsidae The Sepsidae are a family of flies, commonly called the black scavenger flies or ensign flies. Over 300 species are described worldwide. They are usually found around dung or decaying ...
'' Ozerov, 1996 *'' Afromeroplius'' Ozerov, 1996 *''
Afronemopoda ''Afronemopoda'' is a genus of flies in the family Sepsidae The Sepsidae are a family of flies, commonly called the black scavenger flies or ensign flies. Over 300 species are described worldwide. They are usually found around dung or decaying ...
'' Ozerov, 2004 *''
Afrosepsis ''Afrosepsis'' is a genus of flies in the family Sepsidae The Sepsidae are a family of flies, commonly called the black scavenger flies or ensign flies. Over 300 species are described worldwide. They are usually found around dung or decaying p ...
'' Ozerov, 1996 *''
Archisepsis ''Archisepsis'' is a genus of flies in the family Sepsidae. Species *'' Archisepsis armata'' ( Schiner, 1868) *'' Archisepsis bolivica'' Ozerov, 2004 *'' Archisepsis discolor'' (Bigot, 1857) *'' Archisepsis diversiformis'' (Ozerov, 1993) *'' Arc ...
'' Silva, 1993 *''
Australosepsis ''Australosepsis'' is a genus of flies in the family Sepsidae. Species *'' Australosepsis frontalis'' (Walker Walker or The Walker may refer to: People * Walker (given name) *Walker (surname) * Walker (Brazilian footballer) (born 1982), Brazi ...
'' Malloch, 1925 *''
Brachythoracosepsis ''Brachythoracosepsis'' is a genus of flies in the family Sepsidae. Species *'' Brachythoracosepsis butikensis'' (Vanschuytbroeck, 1963) *'' Brachythoracosepsis freidbergi'' Ozerov, 1996 *'' Brachythoracosepsis nodosa'' (Walker Walker or The Wa ...
'' Ozerov, 1996 *'' Decachaetophora''
Duda ) (Polish, Ukrainian Carpathians) *Diple (Dalmatian Coast) *Tulum (Turkish and Pontic) *Tsambouna (Dodecanese and Cyclades) *Askambandoura (Crete) *Gajdy (Polish/Czech/Slovak) *Gaita ( Galician) *Surle (Serbian/Croatian) *Mezoued/Zukra (Northern A ...
, 1926
*''
Diploosmeteriosepsis ''Diploosmeteriosepsis'' is a genus of flies in the family Sepsidae The Sepsidae are a family of flies, commonly called the black scavenger flies or ensign flies. Over 300 species are described worldwide. They are usually found around dung or ...
'' Ozerov, 1996 *''
Dicranosepsis ''Dicranosepsis'' is a genus of flies in the family Sepsidae. Species *'' Dicranosepsis barbata'' Iwasa, 2008 *'' Dicranosepsis bicolor'' ( Wiedemann, 1830) *'' Dicranosepsis biformis'' Iwasa, 2012 *'' Dicranosepsis breviappendiculata'' ( Meijer ...
''
Duda ) (Polish, Ukrainian Carpathians) *Diple (Dalmatian Coast) *Tulum (Turkish and Pontic) *Tsambouna (Dodecanese and Cyclades) *Askambandoura (Crete) *Gajdy (Polish/Czech/Slovak) *Gaita ( Galician) *Surle (Serbian/Croatian) *Mezoued/Zukra (Northern A ...
, 1926
*''
Dudamira ''Dudamira'' is a genus of flies in the family Sepsidae. Species *'' Dudamira abyssinica'' (Duda ) (Polish, Ukrainian Carpathians) *Diple ( Dalmatian Coast) * Tulum (Turkish and Pontic) *Tsambouna (Dodecanese and Cyclades) *Askambandoura (Cret ...
'' Ozerov, 1996 *''
Idiosepsis ''Idiosepsis'' is a genus of flies in the family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain ...
'' Ozerov, 1990 *'' Lasionemopoda''
Duda ) (Polish, Ukrainian Carpathians) *Diple (Dalmatian Coast) *Tulum (Turkish and Pontic) *Tsambouna (Dodecanese and Cyclades) *Askambandoura (Crete) *Gajdy (Polish/Czech/Slovak) *Gaita ( Galician) *Surle (Serbian/Croatian) *Mezoued/Zukra (Northern A ...
, 1926
*'' Lasiosepsis''
Duda ) (Polish, Ukrainian Carpathians) *Diple (Dalmatian Coast) *Tulum (Turkish and Pontic) *Tsambouna (Dodecanese and Cyclades) *Askambandoura (Crete) *Gajdy (Polish/Czech/Slovak) *Gaita ( Galician) *Surle (Serbian/Croatian) *Mezoued/Zukra (Northern A ...
, 1926
*''
Lateosepsis ''Lateosepsis'' is a genus of flies in the family Sepsidae. Species *'' Lateosepsis laticornis'' (Duda ) (Polish, Ukrainian Carpathians) *Diple ( Dalmatian Coast) * Tulum (Turkish and Pontic) *Tsambouna (Dodecanese and Cyclades) *Askambandoura ...
'' Ozerov, 2004 *''
Leptomerosepsis ''Leptomerosepsis'' is a genus of flies in the family Sepsidae. Species *'' Leptomerosepsis improvisa'' Ozerov, 1996 *'' Leptomerosepsis rubiginosa'' Ozerov, 1996 *'' Leptomerosepsis ruwenzoriensis'' (Vanschuytbroeck, 1963) *'' Leptomerosepsis s ...
''
Duda ) (Polish, Ukrainian Carpathians) *Diple (Dalmatian Coast) *Tulum (Turkish and Pontic) *Tsambouna (Dodecanese and Cyclades) *Askambandoura (Crete) *Gajdy (Polish/Czech/Slovak) *Gaita ( Galician) *Surle (Serbian/Croatian) *Mezoued/Zukra (Northern A ...
, 1926
*'' Meropliosepsis''
Duda ) (Polish, Ukrainian Carpathians) *Diple (Dalmatian Coast) *Tulum (Turkish and Pontic) *Tsambouna (Dodecanese and Cyclades) *Askambandoura (Crete) *Gajdy (Polish/Czech/Slovak) *Gaita ( Galician) *Surle (Serbian/Croatian) *Mezoued/Zukra (Northern A ...
, 1926
*'' Meroplius'' Rondani, 1874 *'' Microsepsis'' Silva, 1993 *''
Mucha Mucha (; Czech and Slovak feminine: Muchová) is a Slavic surname, derived from ''mucha'', meaning " fly".''Dictionary of American Family Names''"Mucha Family History" Oxford University Press, 2013. Retrieved on 4 January 2016. Mucha is the standa ...
'' Ozerov, 1992 *'' Nemopoda'' Robineau-Desvoidy, 1830 *'' Ortalischema''
Frey Freyr (Old Norse: 'Lord'), sometimes anglicized as Frey, is a widely attested god in Norse mythology, associated with kingship, fertility, peace, and weather. Freyr, sometimes referred to as Yngvi-Freyr, was especially associated with Sweden and ...
, 1925
*'' Orygma'' Meigen, 1830 *'' Palaeosepsioides'' Ozerov, 1992 *'' Palaeosepsis''
Duda ) (Polish, Ukrainian Carpathians) *Diple (Dalmatian Coast) *Tulum (Turkish and Pontic) *Tsambouna (Dodecanese and Cyclades) *Askambandoura (Crete) *Gajdy (Polish/Czech/Slovak) *Gaita ( Galician) *Surle (Serbian/Croatian) *Mezoued/Zukra (Northern A ...
, 1926
*'' Parapalaeosepsis''
Duda ) (Polish, Ukrainian Carpathians) *Diple (Dalmatian Coast) *Tulum (Turkish and Pontic) *Tsambouna (Dodecanese and Cyclades) *Askambandoura (Crete) *Gajdy (Polish/Czech/Slovak) *Gaita ( Galician) *Surle (Serbian/Croatian) *Mezoued/Zukra (Northern A ...
, 1926
*'' Paratoxopoda''
Duda ) (Polish, Ukrainian Carpathians) *Diple (Dalmatian Coast) *Tulum (Turkish and Pontic) *Tsambouna (Dodecanese and Cyclades) *Askambandoura (Crete) *Gajdy (Polish/Czech/Slovak) *Gaita ( Galician) *Surle (Serbian/Croatian) *Mezoued/Zukra (Northern A ...
, 1926
*'' Perochaeta''
Duda ) (Polish, Ukrainian Carpathians) *Diple (Dalmatian Coast) *Tulum (Turkish and Pontic) *Tsambouna (Dodecanese and Cyclades) *Askambandoura (Crete) *Gajdy (Polish/Czech/Slovak) *Gaita ( Galician) *Surle (Serbian/Croatian) *Mezoued/Zukra (Northern A ...
, 1926
* '' Protorygma'' Hennig, 1965 *'' Pseudonemopoda''
Duda ) (Polish, Ukrainian Carpathians) *Diple (Dalmatian Coast) *Tulum (Turkish and Pontic) *Tsambouna (Dodecanese and Cyclades) *Askambandoura (Crete) *Gajdy (Polish/Czech/Slovak) *Gaita ( Galician) *Surle (Serbian/Croatian) *Mezoued/Zukra (Northern A ...
, 1926
*'' Pseudopalaeosepsis'' Ozerov, 1992 *'' Saltella'' Robineau-Desvoidy, 1830 *''
Sepsis Sepsis, formerly known as septicemia (septicaemia in British English) or blood poisoning, is a life-threatening condition that arises when the body's response to infection causes injury to its own tissues and organs. This initial stage is follo ...
'' Fallén, 1810 *'' Susanomira'' Pont, 1987 *'' Themira'' Robineau-Desvoidy, 1830 *'' Toxopoda'' Macquart, 1851 *'' Xenosepsis'' Malloch, 1925 *'' Zuskamira'' Pont, 1987


See also

* List of sepsid fly species recorded in Europe


References


Further reading


Identification

*
Duda ) (Polish, Ukrainian Carpathians) *Diple (Dalmatian Coast) *Tulum (Turkish and Pontic) *Tsambouna (Dodecanese and Cyclades) *Askambandoura (Crete) *Gajdy (Polish/Czech/Slovak) *Gaita ( Galician) *Surle (Serbian/Croatian) *Mezoued/Zukra (Northern A ...
, O. 1926 Monographie der Sepsiden (Dipt.). ''Ann. Naturhist. Mus. Wien'' 39 (1925): 1-153 and 40 (1926) : 1-110.This work is partly out of date but still the only review of world
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
. *
Willi Hennig Emil Hans Willi Hennig (20 April 1913 – 5 November 1976) was a Germans, German biologist and zoologist who is considered the founder of Phylogenesis, phylogenetic systematics, otherwise known as cladistics. In 1945 as a POWs in World War II, p ...
, 1949: 39a. ''Sepsidae''. In
Erwin Lindner Erwin Lindner (7 April 1888 – 30 November 1988) was a German entomologist mainly interested in Diptera. He was born in Böglins, Memmingen and died in Stuttgart, aged 100 years. In 1913 Erwin Lindner joined the Palaearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
and
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
. *Adrian C. Pont and Rudolf Meier ''The Sepsidae (Diptera) of Europe''.
Fauna Entomologica Scandinavica ''Fauna Entomologica Scandinavica'' is a scientific book series of entomological identification manuals for insects (and other terrestrial arthropods) in North-West Europe, mainly Fennoscandia and Denmark. The series is used by a number of groups, ...
Volume 37. 198 pages. *A.L. Ozerov ''Sepsid Flies (Diptera, Sepsidae) of Russia's Fauna''. Studies on the fauna; Archives of the Zoological Museum of Moscow State University: Zool. Mus. Moscow. Univ. Publ.Language: Russian, title, contents and a summary in English. 184 pages.A very well illustrated guide to all 57 species from 11 genera of Sepsidae flies occurring in Russia, with keys to adults and pre-imaginal stages, and accounts concerning anatomy, phylogeny and distribution. *Silva, V. C. . Revision of the family Sepsidae of the Neotropical region. ii. The genus ''Meropliosepsis'' Duda, 1926 (Diptera, Schizophora). Revista Brasileira de Entomologia, v. 36, n. 3, p. 549-552, 1992. *K. G. V. Smith, 1989 An introduction to the immature stages of British Flies. Diptera Larvae, with notes on eggs, puparia and pupae.''
Handbooks for the Identification of British Insects ''Handbooks for the Identification of British Insects'' is a series of books produced by the Royal Entomological Society (RES). The aim of the Handbooks is to provide illustrated identification keys to the insects of Britain, together with concise ...
'' Vol 10 Part 14
pdf
download manual (two parts Main text and figures index) * Lives in ocky Mountainsof Colorado, in the United States.


Species lists


West Palaearctic including RussiaJapan


External links


Family Sepsidae at EOLImages of Sepsidae on Diptera.info
{{Taxonbar, from=Q1685942 Brachycera families Articles containing video clips