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The scarlet macaw (''Ara macao'') is a large red, yellow, and blue Central and South American parrot, a member of a large group of Neotropical parrots called
macaw Macaws are a group of New World parrots that are long-tailed and often colorful. They are popular in aviculture or as companion parrots, although there are conservation concerns about several species in the wild. Biology Of the many differe ...
s. It is native to humid evergreen forests of the Neotropics. Its range extends from south-eastern Mexico to Peru, Ecuador,
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
,
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
, Venezuela and Brazil in lowlands of (at least formerly) up to , the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
island of Trinidad, as well as the Pacific island of Coiba. Formerly, it ranged north to southern Tamaulipas. In some areas, it has suffered
local extinction Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
because of habitat destruction, or capture for the
parrot trade The international trade in parrots is a lucrative enterprise, and forms an important part of the international wildlife trade. As parrots have become increasingly endangered, many countries have placed restrictions on the trade and/or prohibited t ...
, but in other areas, it remains fairly common. It is the national bird of
Honduras Honduras, officially the Republic of Honduras, is a country in Central America. The republic of Honduras is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Oce ...
. Like its relative the blue-and-yellow macaw, the scarlet macaw is a popular bird in aviculture as a result of its striking plumage.


Taxonomy

The scarlet macaw was formally described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
'' under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Psittacus macao''. The scarlet macaw is now placed in the genus ''Ara'' (
Lacépède Lacepede can refer to: * Bernard Germain de Lacépède (1756–1825), French naturalist and politician. * Lacepede Bay, a bay in South Australia. * Lacépède, Lot-et-Garonne, a ''commune'' in France * Lacepede Islands, a group of four islands in t ...
, 1799), one of 6 genera of Central and South American macaws. The two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
can be recognized by size and color detail in the feathers on the wings: * ''A. m. macao'' (Linnaeus, 1758): South American scarlet macaw, the nominate subspecies. In the wings the medium and secondary coverts have green tip. * ''A. m. cyanopterus'' Wiedenfeld, 1995: North Central American scarlet macaw. The Central American scarlet macaw is larger and has blue on its wings instead of green.


Genetics

In May 2013, it was announced that a team of scientists, led by Dr. Christopher M. Seabury and Dr. Ian Tizard of Texas A&M University had sequenced the complete genome of the scarlet macaw. Based on this genome, species-specific
microsatellite A microsatellite is a tract of repetitive DNA in which certain DNA motifs (ranging in length from one to six or more base pairs) are repeated, typically 5–50 times. Microsatellites occur at thousands of locations within an organism's genome. ...
genetic markers were developed to aid genetic studies throughout the range of the species. These genetic markers were later validated on the trace amount of DNA acquired from feathers, and applied to study red-and-green macaws in a tropical landscape where DNA can degrade very quickly. These markers were proved to be useful to study their population genetics and identification of individuals in the landscape of the Peruian Amazon.


Description

It is about long, of which more than half is the pointed, graduated tail typical of all
macaw Macaws are a group of New World parrots that are long-tailed and often colorful. They are popular in aviculture or as companion parrots, although there are conservation concerns about several species in the wild. Biology Of the many differe ...
s, though the scarlet macaw has a larger percentage of tail than the other large macaws. The average weight is about . The
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
is mostly
scarlet Scarlet may refer to: * Scarlet (cloth), a type of woollen cloth common in medieval England * Scarlet (color), a bright tone of red that is slightly toward orange, named after the cloth * Scarlet (dye), the dye used to give the cloth its color * ...
, but the rump and tail-covert
feather Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and a premier ...
s are light blue, the greater upper wing coverts are yellow, the upper sides of the flight feathers of the wings are dark blue as are the ends of the tail feathers, and the undersides of the wing and tail
flight feather Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tail ...
s are dark red with metallic gold iridescence. Some individuals may have green in the wings. There is bare white skin around the eye and from there to the bill. Tiny white feathers are contained on the face patch. The upper mandible is mostly pale horn in color and the lower is black. Juveniles have dark eyes; adults have light yellow eyes. It is frequently confused with the slightly larger green-winged macaw, which has more distinct red lines in the face and no yellow in the wing. Scarlet macaws make very loud, high and sometimes low-pitched, throaty squawks, squeaks and screams designed to carry many kilometers to call for their groups. The scarlet macaw can live up to 75 or even 90 years in captivity, although a more typical lifespan is 40 to 50 years.Robert Arking: ''Biology of Aging: Observations and Principles''. Oxford University Press, 2006, , p
129
/ref>
at the biology website of the Lamar University (retrieved 2019-02-24)


Behavior

A typical sighting is of a single bird or a pair flying above the forest canopy, though in some areas flocks can be seen. They often gather at
clay lick A mineral lick (also known as a salt lick) is a place where animals can go to lick essential mineral nutrients from a deposit of salts and other minerals. Mineral licks can be naturally occurring or artificial (such as blocks of salt that ...
s. Scarlet macaws communicate primarily through raucous honks; however, vocal communication is highly variable, and captive macaws are known to be adept mimics of human speech.


Feeding

Wild scarlet macaws feed on fruits, nuts, seeds, flowers and nectar. They also love to eat insects and larvae. They are seen feeding heavily on bugs, snails and foliage. Snails and bugs are great source of protein, as they need additional protein during breeding seasons. In Costa Rica's Central Pacific they have learned to feed on introduced Teak trees ('' Tectona grandis'') and Almond Beach Trees (Terminalia sp). The local non-profit organizations have planted hundreds of those trees along the coastline from Tarcoles River basin to Esterillos Beach which had helped increase the population drastically. The combined efforts and the correct ecotourism has also an important role in the conservation of such majestic birds. Tour companies along the Tarcoles River and its mangroves have bet on the importance of birdwatching as an asset for the growth on its population.


Breeding

While comparatively docile at most times of the year, scarlet macaws may be formidably aggressive during periods of breeding. Scarlet macaws are monogamous birds, with individuals remaining with one partner throughout their lives. The hen lays two or three white eggs in a large tree cavity. The female incubates the eggs for about five weeks, and the chicks
fledge Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnerable c ...
from the nest about 90 days after hatching and leave their parents about a year later. Juveniles reach sexual maturity at five years of age.


Distribution and habitat

The South American range is extensive and covers the Amazon forest, extending to Peru east of the Andes, to
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
. In Bolivia, it is very present in the Aquicuana Reserve, located in the Beni Department, near the city of
Riberalta Riberalta is a town in the Beni Department in northern Bolivia, situated where the Madre de Dios River joins the Beni River. Riberalta is on the south (south east) bank of the Beni River. As the capital of the province of Vaca Diez, the city ...
, the Capital of the Bolivian Amazon. In Central America, the range extends from extreme eastern and southern Mexico and Panama through
Guatemala Guatemala ( ; ), officially the Republic of Guatemala ( es, República de Guatemala, links=no), is a country in Central America. It is bordered to the north and west by Mexico; to the northeast by Belize and the Caribbean; to the east by H ...
,
El Salvador El Salvador (; , meaning " The Saviour"), officially the Republic of El Salvador ( es, República de El Salvador), is a country in Central America. It is bordered on the northeast by Honduras, on the northwest by Guatemala, and on the south b ...
,
Honduras Honduras, officially the Republic of Honduras, is a country in Central America. The republic of Honduras is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Oce ...
and Belize, the island of Coiba and infrequently on the mainland of Panama, and in Costa Rica in isolated regions on the Pacific Coast; the Nicoya Peninsula the Carara National Park and Peninsula de Osa. The scarlet macaw has escaped or been deliberately released in to Florida, but there is no evidence that the population is breeding and may only persist due to continuing releases or escapes. Scarlet macaws inhabit humid lowland
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
rain forests, open woodlands, river edges, and savannas.


Conservation status

The habitat of scarlet macaws is also considered to have the greatest latitudinal range for any bird in the genus ''Ara'', as the estimated maximum territorial range covers 6,700,000 km2. Nevertheless, the scarlet macaw's habitat is fragmented, and the bird is mostly confined to tiny populations scattered throughout its original range in Middle America.Juniper, T., and M. Parr., (1998). ''Parrots: A Guide to Parrots of the World.'' Yale University Press. However, as they still occur in large numbers over most of their original range in South America, the species is classified by IUCN as least concern. Its wild population is currently estimated between 20,000 and 50,000 individuals. Commercial international trade in the species (including parts and derivatives) is prohibited by the bird's listing under CITES Appendix 1 due to predation for the pet and cage bird trade. Both subspecies are listed by USFWS as endangered.


Aviculture

The scarlet macaw is an early example of a parrot breeding in captivity. Captive breeding occurred in Northern Mexico at Paquime (also called Casas Grandes) and very likely Southwest New Mexico
Mimbres Valley The Mimbres Valley AVA is an American Viticultural Area located in southwestern New Mexico near the towns of Deming and Silver City. The AVA includes the Mimbres Watershed of the Mimbres River, and most vineyards are planted at elevations ...
in the 11th century. Breeding pens, perches, bones, and eggshell fragments have been uncovered. The straightforward nature of scarlet macaw breeding and the value of their plumes in trade created a market for trade wherein the animals were used in religious rites north to the Colorado Plateau region. Today the scarlet macaw is found worldwide in captivity, but is best represented in captivity in the Americas. Captive techniques developed from the pet trade have positively affected wild populations: in areas with low macaws populations, the "extra" babies that typically die in the nest may be reared by humans hands and released into the wild to bolster the population, as has been done by the
Tambopata Macaw Project The Macaw Society (formerly known as the Tambopata Macaw Project) is a long-term research project on the ecology and conservation of macaws and parrots under the direction of Donald Brightsmith and Gabriela Vigo of the Schubot Center for Avian H ...
. Their captive diet, egg incubation, assisted hatching, hand rearing, co-parenting, parent-rearing, fledgling, maturation, and breeding are well understood within the avicultural community (AFA Watchbird magazine).


See also

* The Ara Project (macaw reintroduction) * The Macaw Society *
List of macaws The list of macaws includes 19 species of macaws including extinct and critically endangered species, and does not include several hypothetical extinct species that have been proposed based on very little evidence. Species in taxonomic order '' ...
* Iago (Disney)


References


External links


Scarlet Macaw
in-depth research article.
Tambopata Macaw project
long term research project in south east Peru
World Parrot Trust
Parrot Encyclopedia — Species Profiles
Scarlet macaw videos
on the Internet Bird Collection

* ttp://www.pbase.com/image/19378619 Scarlet macaw photo on PBase
Stamps
(15 countries) with RangeMap
''Ara macao''
conservation and reintroduction in Costa Rica


The Scarlet Macaw Genome Project

Scarlet macaw information


in-depth research and facts scarlet-macaws] {{Authority control Ara (genus), scarlet macaw scarlet macaw Birds of Central America Birds of Colombia Birds of the Amazon Basin Birds of Ecuador Birds of Venezuela Birds of Trinidad and Tobago Birds of the Caribbean Talking birds National symbols of Honduras scarlet macaw scarlet macaw Articles containing video clips Species endangered by the pet trade