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The Sambirano region, also known as the Sambirano domain, is a
biogeographic Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
region of northwestern
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
. It has a distinctive lowland climate with year-round rain, differing from the rest of seasonally-dry western and northern Madagascar. The climate supports humid lowland rainforests, with greater similarity to those of eastern Madagascar than the dry deciduous forests characteristic of the Madagascar's western lowlands.Donque, G. (1972). The Climatology of Madagascar. In: Battistini, R., Richard-Vindard, G. (eds) ''Biogeography and Ecology in Madagascar''. Monographiae Biologicae, vol 21. Springer, Dordrecht. https://doi-org.wikipedialibrary.idm.oclc.org/10.1007/978-94-015-7159-3_3Koechlin, J. (1972). Flora and Vegetation of Madagascar. In: Battistini, R., Richard-Vindard, G. (eds) ''Biogeography and Ecology in Madagascar''. Monographiae Biologicae, vol 21. Springer, Dordrecht. https://doi-org.wikipedialibrary.idm.oclc.org/10.1007/978-94-015-7159-3_4Chauvet, B. (1972). The Forests of Madagascar. In: Battistini, R., Richard-Vindard, G. (eds) ''Biogeography and Ecology in Madagascar''. Monographiae Biologicae, vol 21. Springer, Dordrecht. https://doi-org.wikipedialibrary.idm.oclc.org/10.1007/978-94-015-7159-3_5


Geography

The region takes its name from the
Sambirano River The Sambirano is a river of northwestern Madagascar in the region of Diana. It has its sources at the Maromokotra peak and flows through the Tsaratanana Reserve to the Indian Ocean. Its delta occupies 250 km. The ecosystem of the river, ma ...
, which runs through the center of the region. The Sambirano region extends from the Ifasy River in the north to the Ampasindava Peninsula in the south, in the southwestern portion of
Diana Region Diana is a region in Madagascar at the northeast part of the island. It borders the regions of Sava to the southeast and Sofia to the southwest. It covers an area of 19,266 km2, and had a population of 889,736 in 2018. The regional capital i ...
. It extends from the coast to the slopes of the
Tsaratanana Massif Madagascar is a large island in the Indian Ocean off the eastern coast of southern Africa, east of Mozambique. It has a total area of with of land and of water. Madagascar is the fourth largest island and the 2nd largest island country in th ...
, which bounds the region on the east. It also includes the islands of
Nosy Be Nosy Be (formerly Nossi-bé and Nosse Be) is an island off the northwest coast of Madagascar. Nosy Be is Madagascar's largest and busiest tourist resort. It has an area of , and its population was 109,465 according to the provisional results of t ...
and
Nosy Komba Nosy Komba is a volcanic island off the north west coast of Madagascar, lying between Nosy Be Nosy Be (formerly Nossi-bé and Nosse Be) is an island off the northwest coast of Madagascar. Nosy Be is Madagascar's largest and busiest tourist re ...
.


Climate

The region receives summer monsoon rains characteristic of the west coast, but also rain during the winter months from the easterly trade winds. The trade winds bring winter rain to eastern Madagascar but across most of the west are intercepted by Madagascar's central highlands. The convergence of easterly wind currents in the lee of the Tsaratanana Massif is responsible for the Sambirano region's higher rainfall. Average annual rainfall in
Ambanja Ambanja is a city and commune in northern Madagascar. According to 2001 census the population of Ambanja was 28,468. Geography Ambanja is located on the northern berth of the Sambirano River and is crossed by the Route Nationale 6 ( Antsiranan ...
is 2171 mm. January is rainiest month, averaging 538.4 mm. May to October are drier months, with the lowest rainfall (24.6 mm) in July. Mean annual temperature averages 25.7° C at Ambanja, with little seasonal variation and humidity throughout most of the year. As one ascends the slopes of Tsaratanana, rainfall and humidity increase, and temperature and seasonal rainfall variation decrease.


Flora

The predominant natural vegetation in the region is lowland rainforest, although little remains. Mature forests have a canopy of trees 25 to 30 meters high, festooned with abundant
liana A liana is a long- stemmed, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the canopy in search of direct sunlight. The word ''liana'' does not refer to a ta ...
s. There is a middle stratum of smaller, shade-tolerant trees, and a sparse understory of ferns, dwarf palms, shrubs, and herbaceous plants. Trees of the families
Sarcolaenaceae The Sarcolaenaceae are a family of flowering plants endemic to Madagascar. The family includes 79 species of mostly evergreen trees and shrubs in ten genera. Recent DNA studies indicate that the Sarcolaenaceae are a sibling taxon to the family ...
and
Myristicaceae The Myristicaceae are a family of flowering plants native to Africa, Asia, Pacific islands, and the Americas and has been recognized by most taxonomists. It is sometimes called the "nutmeg family", after its most famous member, ''Myristica fragra ...
, and genus ''
Anthostema ''Anthostema'' is a flowering plant genus in the Family Euphorbiaceae (spurge family) first described as a genus in 1824. It is native to Africa and Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg ...
'' are characteristic canopy trees, along with trees of families
Fabaceae The Fabaceae or Leguminosae,International Code of Nomenc ...
(''
Cynometra ''Cynometra'' is genus of tropical forest trees with a pantropical distribution. It is particularly important as a forest component in west Africa and the neotropics. '' Cynometra alexandri'' (muhimbi) is a familiar timber tree of central and ea ...
''),
Lauraceae Lauraceae, or the laurels, is a plant family that includes the true laurel and its closest relatives. This family comprises about 2850 known species in about 45 genera worldwide (Christenhusz & Byng 2016 ). They are dicotyledons, and occur ma ...
(''
Ocotea ''Ocotea'' is a genus of flowering plants belonging to the family Lauraceae. Many are evergreen trees with lauroid leaves. There are over 520 species currently accepted within the genus, distributed mostly in tropical and subtropical areas of t ...
'' and '' Cryptocarya agathophylla''),
Ebenaceae The Ebenaceae are a family of flowering plants belonging to order Ericales. The family includes ebony and persimmon among about 768 species of trees and shrubs. It is distributed across the tropical and warmer temperate regions of the world. It ...
(''
Diospyros ''Diospyros'' is a genus of over 700 species of deciduous and evergreen trees and shrubs. The majority are native to the tropics, with only a few species extending into temperate regions. Individual species valued for their hard, heavy, dark tim ...
''),
Euphorbiaceae Euphorbiaceae, the spurge family, is a large family of flowering plants. In English, they are also commonly called euphorbias, which is also the name of a genus in the family. Most spurges, such as ''Euphorbia paralias'', are herbs, but some, e ...
,
Arecaceae The Arecaceae is a family of perennial flowering plants in the monocot order Arecales. Their growth form can be climbers, shrubs, tree-like and stemless plants, all commonly known as palms. Those having a tree-like form are called palm trees ...
,
Rubiaceae The Rubiaceae are a family of flowering plants, commonly known as the coffee, madder, or bedstraw family. It consists of terrestrial trees, shrubs, lianas, or herbs that are recognizable by simple, opposite leaves with interpetiolar stipules ...
,
Sapindaceae The Sapindaceae are a family of flowering plants in the order Sapindales known as the soapberry family. It contains 138 genera and 1858 accepted species. Examples include horse chestnut, maples, ackee and lychee. The Sapindaceae occur in tempera ...
, and
Anacardiaceae The Anacardiaceae, commonly known as the cashew family or sumac family, are a family of flowering plants, including about 83 genera with about 860 known species. Members of the Anacardiaceae bear fruits that are drupes and in some cases produce ...
. Human disturbance, has reduced the rainforest to scattered patches. Grasslands are maintained through regular burning, and are used for livestock grazing. Secondary forest, known as ''savoka'', grows in areas which were cleared for
shifting cultivation Shifting cultivation is an agricultural system in which plots of land are cultivated temporarily, then abandoned while post-disturbance fallow vegetation is allowed to freely grow while the cultivator moves on to another plot. The period of cul ...
and then left fallow. the palm-like native plant ''
Ravenala madagascariensis ''Ravenala'' is a genus of monocotyledonous flowering plants. Classically, the genus was considered to include a single species, ''Ravenala madagascariensis'', commonly known as the traveller's tree, traveller's palm or East-West palm, from Mada ...
'' is common in disturbed areas. Introduced species are increasingly common, including Citrus, ''
Psidium ''Psidium'' is a genus of trees and shrubs in the family Myrtaceae. It is native to warmer parts of the Western Hemisphere (Mexico, Central and South America, the West Indies the Galápagos islands). Taxonomy This genus was described first by ...
'' (guava), and the bamboo ''
Cathariostachys capitata ''Cathariostachys'' is a genus of Madagascan bamboo in the grass family. ;Species # '' Cathariostachys capitata'' - eastern Madagascar # ''Cathariostachys madagascariensis'' (A.Camus) S.Dransf. - central Madagascar Madagascar (; mg, ...
''.


Fauna

Several species of
lemur Lemurs ( ) (from Latin ''lemures'' – ghosts or spirits) are Strepsirrhini, wet-nosed primates of the Superfamily (biology), superfamily Lemuroidea (), divided into 8 Family (biology), families and consisting of 15 genera and around 100 exist ...
are endemic or mostly endemic to the Sambirano region, including the black lemur (''Eulemur macaco''), Pariente's fork-marked lemur (''Phaner parienti''), Nosy Be mouse lemur (''Microcebus mamiratra''), and Nosy Be sportive lemur (''Lepilemur tymerlachsoni'').


Conservation

The largest remaining area of lowland rainforest is in Lokobe National Park on Nosy Be.Griveaud, P., Albignac, R. (1972). The Problems of Nature Conservation in Madagascar. In: Battistini, R., Richard-Vindard, G. (eds) ''Biogeography and Ecology in Madagascar''. Monographiae Biologicae, vol 21. Springer, Dordrecht. https://doi-org.wikipedialibrary.idm.oclc.org/10.1007/978-94-015-7159-3_23
Manongarivo Special Reserve Manongarivo Reserve is a wildlife reserve in the North-West of Madagascar in the region of Diana. Manongarivo is home to both the Sambirano mouse lemur and the Sambirano woolly lemur. Geography The reserve has a surface of 64,356 ha and is sit ...
preserves some enclaves of humid forest along with transitional lowland dry forest.


References

{{reflist Madagascar subhumid forests Diana Region