''Rudbeckia hirta'', commonly called black-eyed Susan, is a North American

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...

in the family

Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae ...

native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...

to Eastern and Central

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...

and naturalized in the Western part of the continent as well as in

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...

. It has now been found in all 10

Canadian Provinces Within the geographical areas of Canada, the ten provinces and three territories are sub-national administrative divisions under the jurisdiction of the Canadian Constitution. In the 1867 Canadian Confederation, three provinces of British Nort ...

and all 48 of the states in the

contiguous United States The contiguous United States (officially the conterminous United States) consists of the 48 adjoining U.S. states and the Federal District of the United States of America. The term excludes the only two non-contiguous states, Alaska and Hawaii ...

. ''Rudbeckia hirta'' is the state flower of

Maryland Maryland ( ) is a state in the Mid-Atlantic region of the United States. It shares borders with Virginia, West Virginia, and the District of Columbia to its south and west; Pennsylvania to its north; and Delaware and the Atlantic Ocean t ...

Description

''Rudbeckia hirta'' is an upright

annual Annual may refer to: *Annual publication, periodical publications appearing regularly once per year ** Yearbook ** Literary annual *Annual plant *Annual report *Annual giving *Annual, Morocco, a settlement in northeastern Morocco *Annuals (band), ...

(sometimes biennial or

perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also widel ...

) growing tall by wide. It has alternate, mostly basal

leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...

10–18 cm long, covered by coarse hair, with stout branching stems and daisy-like,

composite Composite or compositing may refer to: Materials * Composite material, a material that is made from several different substances ** Metal matrix composite, composed of metal and other parts ** Cermet, a composite of ceramic and metallic materials ...

flower heads A pseudanthium (Greek for "false flower"; ) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, compos ...

appearing in late summer and early autumn. In the species, the flowers are up to in diameter, with yellow

ray florets The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae w ...

circling conspicuous brown or black, dome-shaped cone of many small

disc floret The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae ...

s. However, extensive breeding has produced a range of sizes and colours, including oranges, reds and browns.

Etymology and common names

The

specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...

''hirta'' is Latin for “hairy”, and refers to the

trichomes Trichomes (); ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a plant ...

occurring on leaves and stems. Other common names for this plant include: brown-eyed Susan, brown betty, gloriosa daisy, golden Jerusalem, English bull's eye, poor-land daisy, yellow daisy, and yellow ox-eye daisy.

Varieties

There are four varieties *''Rudbeckia hirta'' var. ''angustifolia'' - southeastern + south-central United States (South Carolina to Texas) *''Rudbeckia hirta'' var. ''floridana'' -

Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and ...

*''Rudbeckia hirta'' var. ''hirta'' -

Eastern United States The Eastern United States, commonly referred to as the American East, Eastern America, or simply the East, is the region of the United States to the east of the Mississippi River. In some cases the term may refer to a smaller area or the East C ...

(

Maine Maine () is a state in the New England and Northeastern regions of the United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Canadian provinces of New Brunswick and Quebec to the northeast and nor ...

Alabama (We dare defend our rights) , anthem = " Alabama" , image_map = Alabama in United States.svg , seat = Montgomery , LargestCity = Huntsville , LargestCounty = Baldwin County , LargestMetro = Greater Birmingham , area_total_km2 = 135,7 ...

). *''Rudbeckia hirta'' var. ''pulcherrima'' - Widespread in most of

(

Newfoundland Newfoundland and Labrador (; french: Terre-Neuve-et-Labrador; frequently abbreviated as NL) is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region ...

British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, for ...

, south to

and

New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe, New Mexico, Santa Fe , LargestCity = Albuquerque, New Mexico, Albuquerque , LargestMetro = Albuquerque metropolitan area, Tiguex , Offi ...

; naturalized

Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered o ...

California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...

Cultivation

''Rudbeckia hirta'' is widely cultivated in parks and gardens, for summer bedding schemes, borders, containers, wildflower gardens, prairie-style plantings and cut flowers. Numerous

cultivars A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...

have been developed, of which '

Indian Summer An Indian summer is a period of unseasonably warm, dry weather that sometimes occurs in autumn in temperate regions of the northern hemisphere. Several sources describe a true Indian summer as not occurring until after the first frost, or more s ...

' and 'Toto' have gained the

Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...

Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...

. Other popular cultivars include 'Double Gold' and 'Marmalade'. Gloriosa daisies are

tetraploid Polyploidy is a condition in which the cells of an organism have more than one pair of ( homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two sets of chromosomes, where each set contains ...

having much larger flower heads than the wild species, often doubled or with contrasting markings on the

. They were first bred by Alfred Blakeslee of

Smith College Smith College is a private liberal arts women's college in Northampton, Massachusetts. It was chartered in 1871 by Sophia Smith and opened in 1875. It is the largest member of the historic Seven Sisters colleges, a group of elite women's coll ...

by applying

colchicine Colchicine is a medication used to treat gout and Behçet's disease. In gout, it is less preferred to NSAIDs or steroids. Other uses for colchicine include the management of pericarditis and familial Mediterranean fever. Colchicine is taken b ...

to ''R. hirta'' seeds; Blakeslee's stock was further developed by W. Atlee Burpee and introduced to commerce at the 1957 Philadelphia Flower Show. Gloriosa daisies are generally treated as annuals or short-lived

s and are typically grown from seed, though there are some named cultivars.

Symbolism and uses

Maryland state flower

The black-eyed Susan was designated the state flower of

in 1918. In this capacity it is used in gardens and ceremonies to celebrate, memorialize and show affection for the state of Maryland and its people. The

Preakness Stakes The Preakness Stakes is an American thoroughbred horse race held on Armed Forces Day which is also the third Saturday in May each year at Pimlico Race Course in Baltimore, Maryland. It is a Graded stakes race, Grade I race run over a distance of ...

Baltimore, Maryland Baltimore ( , locally: or ) is the most populous city in the U.S. state of Maryland, fourth most populous city in the Mid-Atlantic, and the 30th most populous city in the United States with a population of 585,708 in 2020. Baltimore wa ...

, has been termed "The Run for the Black-Eyed Susans" because a blanket of Viking Poms, a variety of

chrysanthemum Chrysanthemums (), sometimes called mums or chrysanths, are flowering plants of the genus ''Chrysanthemum'' in the family Asteraceae. They are native to East Asia and northeastern Europe. Most species originate from East Asia and the cent ...

s resembling black-eyed Susans, is traditionally placed around the winning horse's neck (actual black-eyed Susans are not in bloom in May during the Preakness).

University of Southern Mississippi

In 1912, the black-eyed Susan became the inspiration for the

University of Southern Mississippi The University of Southern Mississippi (Southern Miss or USM) is a Public university, public research university with its main campus located in Hattiesburg, Mississippi. It is accredited by the Southern Association of Colleges and Schools to a ...

school colors (black and gold), suggested by Florence Burrow Pope, a member of the university's first graduating class. According to Pope: “On a trip home, I saw great masses of Black-Eyed Susans in the pine forests. I decided to encourage my senior class to gather Black-Eyed Susans to spell out the name of the class on sheets to be displayed during exercises on Class Day. I then suggested black and gold as class colors, and my suggestion was adopted."

Butterfly attractant for enhancing gardens

Butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprises ...

are attracted to ''Rudbeckia hirta''. It is a larval host to the

bordered patch ''Chlosyne lacinia'', the bordered patch or sunflower patch, is a North and South American butterfly in the family Nymphalidae.James A. Scott (1986). ''The Butterflies of North America''. Stanford University Press, Stanford, CA. Description ...

, gorgone checkerspot, and

silvery checkerspot ''Chlosyne nycteis'', the silvery checkerspot, is a species of Nymphalinae butterfly that occurs in North America. It is listed as a species of special concern in Connecticut and Maine, and is believed extirpated in Connecticut, Massachusetts, ...

species.

Traditional Native American uses

The plant is thought to be an

herbal medicine Herbal medicine (also herbalism) is the study of pharmacognosy and the use of medicinal plants, which are a basis of traditional medicine. With worldwide research into pharmacology, some herbal medicines have been translated into modern remedie ...

by Native American for various ailments. The roots but not the seedheads of ''Rudbeckia hirta'' can be used much like the related '' Echinacea purpurea'' with unsubstantiated claims to boost

immunity Immunity may refer to: Medicine * Immunity (medical), resistance of an organism to infection or disease * ''Immunity'' (journal), a scientific journal published by Cell Press Biology * Immune system Engineering * Radiofrequence immunity de ...

and fight colds, flu and infections. The

Ojibwa The Ojibwe, Ojibwa, Chippewa, or Saulteaux are an Anishinaabe people in what is currently southern Canada, the northern Midwestern United States, and Northern Plains. According to the U.S. census, in the United States Ojibwe people are one of ...

people used it as a

poultice A poultice, also called a cataplasm, is a soft moist mass, often heated and medicated, that is spread on cloth and placed over the skin to treat an aching, inflamed, or painful part of the body. It can be used on wounds, such as cuts. 'Poultice ...

for

snake bites A snakebite is an injury caused by the bite of a snake, especially a venomous snake. A common sign of a bite from a venomous snake is the presence of two puncture wounds from the animal's fangs. Sometimes venom injection from the bite may occu ...

and to make an

infusion Infusion is the process of extracting chemical compounds or flavors from plant material in a solvent such as water, oil or alcohol, by allowing the material to remain suspended in the solvent over time (a process often called steeping). An in ...

for treating colds and worms in children.

Cautions

The species is toxic to

cats The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members o ...

, when ingested.

Gallery

Rudbeckiahirta1web.jpg,

Inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are forme ...

and

involucral bract In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or o ...

s Rudbeckia hirta Indian Summer.JPG, ''Rudbeckia hirta'' 'Indian Summer' Northern Crescent 3.jpg, Northern Crescent ( Phyciodes cocyta) butterfly

References

External links

Knowlton Foote. 2001. Black-eyed Susan (''Rudbeckia hirta'' L.). New York Flora Association. Newsletter Vol. 13.Florida Native Plant Society: Rudbeckia Hirta
{{Taxonbar, from=Q2532820 Butterfly food plants Flora of North America Plants described in 1753 Plants used in traditional Native American medicine

hirta Hirta ( gd, Hiort) is the largest island in the St Kilda archipelago, on the western edge of Scotland. The names (in Scottish Gaelic) and ''Hirta'' (historically in English) have also been applied to the entire archipelago. Now without a perman ...

Symbols of Maryland Taxa named by Carl Linnaeus