Réserve Spéciale D'Ankarana
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Ankarana Special Reserve in northern Madagascar was created in 1956. It is a small, partially vegetated plateau composed of 150-million-year-old middle Jurassic limestone. With an average annual rainfall of about , the underlying rocks have been eroded to produce caves and feed subterranean rivers—a
karst topography Karst is a topography formed from the dissolution of soluble rocks such as limestone, dolomite, and gypsum. It is characterized by underground drainage systems with sinkholes and caves. It has also been documented for more weathering-resistant ro ...
. The rugged relief and the dense vegetation have helped protect the region from human intrusion. The southern entrance of the park is in Mahamasina on the Route nationale 6 some 108 km south-west of Antsiranana and north-east of Ambilobe.


Geology

The plateau slopes gently to the east, but on the west it ends abruptly in the "Wall of Ankarana", a sheer cliff that extends north to south, and rises up to . To the south, the limestone mass breaks up into separate spires known as ''tower
karst Karst is a topography formed from the dissolution of soluble rocks such as limestone, dolomite, and gypsum. It is characterized by underground drainage systems with sinkholes and caves. It has also been documented for more weathering-resistant ro ...
''. In the center of the plateau, seismic activity and eons of rainfall have eroded the limestone, forming deep gorges and ribbons of flowstone. In places where the calcific upper layers have been completely eroded, the harder base rock has been etched into channels and ridges known in malagasy as ''tsingy''. The area is littered with basalt boulders and basalt has also flowed deep into the canyons that dissect the Massif.


Exploration

Beginning in the 1960s, expatriate Frenchman Jean Duflos (who after marriage changed his name to Jean Radofilao) undertook a huge amount of exploration of the cave systems and subterranean rivers of the Massif, much of it on his own or with visiting speleologists. Around of cave passages within the massif have been mapped. La Grotte d'Andrafiabe, one of the most accessible caves, comprises at least of horizontal passages. Indeed, the Massif contains the longest cave systems in Madagascar, and probably in the whole of Africa.


Fauna

Expeditions that first began cataloguing the animals and plants of the Special Reserve created around the Ankarana Massif in the 1980s are described in Dr Jane Wilson-Howarth's travel narrative ''Lemurs of the Lost World'' and in the scientific press. Discoveries included unexpected sub-fossil remains of large extinct lemurs and surviving but previously undescribed species of blind fish, shrimps and other invertebrates. Several expedition members contributed photos to an illustrated introductory guide to Madagascar which features the Crocodile Caves of Ankarana. During the 1986 expedition, Phil Chapman and Jean-Elie Randriamasy collated a bird list for the reserve and recorded 65 species from 32 families representing nearly a third of all bird species that breed in Madagascar. They also noted one interesting aspect of behaviour. They reported that there was an unusual strategy used by many of the small insect-eating songbirds. Species such as the Paradise Flycatcher (''
Terpsiphone mutata The Malagasy paradise flycatcher (''Terpsiphone mutata'') is a species of bird in the family Monarchidae. It is found in Comoros, Madagascar, and Mayotte. Its natural habitats are subtropical or tropical dry forest and subtropical or tropical moi ...
''), the Common Jery (''
Neomixis tenella The common jery (''Neomixis tenella'') is a species of bird in the family Cisticolidae. It is endemic to Madagascar. Its natural habitats are subtropical or tropical dry forest and subtropical or tropical moist lowland forest Tropical and s ...
''), the Greenbuls (''
Phyllastrephus zosterops The spectacled tetraka (''Xanthomixis zosterops'') is a species of Malagasy warbler in the family Bernieridae. It is found only in Madagascar. Its natural habitat is subtropical or tropical moist lowland forests. References Malagasy warb ...
'' and ''
Phyllastrephus madagascariensis The long-billed bernieria (''Bernieria madagascariensis''), formerly known as long-billed greenbul and sometimes as common tetraka or long-billed tetraka, is a songbird species endemic to Madagascar. It was initially considered a greenbul, and l ...
''), the Bulbul (''
Hypsipetes madagascariensis The Malagasy bulbul (''Hypsipetes madagascariensis'') is a species of songbird in the bulbul family, Pycnonotidae. It is found on Madagascar and other regional islands in the south-western Indian Ocean. Taxonomy and systematics The Malagasy b ...
''), the Sunbird (''
Nectarinia souimanga The souimanga sunbird (''Cinnyris sovimanga'') is a small passerine bird of the sunbird family, Nectariniidae. It is native to the islands of the western Indian Ocean where it occurs on Madagascar, the Aldabra Group and the Glorioso Islands. Des ...
'') and the Vagas ('' Lepopterus madagascarinus'' and ''
Xenopirostris polleni Pollen's vanga (''Xenopirostris polleni'') is a species of bird in the family Vangidae. It is endemic to eastern Madagascar. Its natural habitats are subtropical or tropical moist lowland forests and subtropical or tropical moist montane forests ...
'') foraged together in mixed bands. Within each band different species seemed to specialise in where and how they searched out their insect prey. Some species concentrated on the trunk and branches of trees, some on slender boughs, others searched beneath the leaves. By acting together in this way they probably increased foraging efficiency as each species could catch others’ escaped prey. They were also safer from attack by predators, as the group as a whole was more likely to spot approaching danger. The Ankarana Reserve is an important refuge for significant populations of the crowned lemur ''(Eulemur coronatus)'', Sanford's brown lemur ''(Eulemur sanfordi)'' and other mammal species. The following lemurs are also recorded from the area:
northern sportive lemur The northern sportive lemur (''Lepilemur septentrionalis''), also known as the Sahafary sportive lemur or northern weasel lemur, is a species of lemur in the family Lepilemuridae. It is endemic to Madagascar. As a result of severe ecological and ...
(''Lepilemur septentrionalis''), brown mouse lemur (''Microcebus rufus''), fat-tailed dwarf lemur (''Cheirogalus medius''), fork-marked lemur (''Phaner furcifer''), eastern woolly lemur (''Avahi laniger''),
Perrier's sifaka Perrier's sifaka (''Propithecus perrieri'') is a lemur endemic to Madagascar. It was formerly considered to be a subspecies of diademed sifaka It has a very small range in northeastern Madagascar where its habitat is dry deciduous or semihumid ...
(''Propithicus diadema perrieri''), aye-aye (''Daubentonia madagascariensis'') and the western lesser bamboo lemur (''Hapalemur griseus occidentalis''). In addition
subfossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, Seashell, shells, exoskeletons, stone imprints of animals or microbes, objects pre ...
of the following lemurs have been found at Ankarana: greater bamboo lemur (''Prolemur simus''), indri (''Indri indri''), the
sloth lemur The sloth lemurs (Palaeopropithecidae) comprise an extinct family of lemurs that includes four genera. The common name can be misleading, as members of Palaeopropithecidae were not closely related to sloths. This clade has been dubbed the ‘‘s ...
(''
Babakotia radofilai ''Babakotia'' is an extinct genus of medium-sized lemur, or strepsirrhine primate, from Madagascar that contains a single species, ''Babakotia radofilai''. Together with '' Palaeopropithecus'', '' Archaeoindris'', and '' Mesopropithecus'', it fo ...
''), ''Mesopropithicus dolichobrachion'' and ''Palaeopropithicus'' cf ''ingens'' plus '' Pachylemur'' sp., the huge '' Megaladapis'' cf ''madagascariensis/grandidieri'', and the baboon lemur '' Archaeolemur'' sp.


See also

* Madagascar dry deciduous forests * Crowned lemur


References


External links


Madagascar National Parks
{{National parks of Madagascar Special reserves of Madagascar Mountains of Madagascar Geography of Madagascar Plateaus of Africa Protected areas in Diana Region Protected areas established in 1956 Geology of Madagascar Madagascar subhumid forests Madagascar dry deciduous forests Important Bird Areas of Madagascar