Red-throated Caracara
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The red-throated caracara (''Ibycter americanus'') is a social species of
bird of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators h ...
in the family
Falconidae The falcons and caracaras are around 60 species of diurnal birds of prey that make up the family Falconidae (representing all extant species in the order Falconiformes). The family is divided into three subfamilies, Herpetotherinae, which inclu ...
. It is placed in the
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Ibycter'', or sometimes united in ''Daptrius'' with the
black caracara The black caracara (''Daptrius ater'') is a species of bird of prey in the family Falconidae found in Amazonian and French Guiana lowlands, commonly along rivers. They are locally referred to as ''Ger' futu busikaka'' in the Republic of Suriname ...
. Unique among caracaras, it mainly feeds on the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e of
bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyly, monophyletic lineage within the ...
s and
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. Th ...
s, but also takes the adult insects and
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
s and
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
. It is found from Mexico south to Venezuela in most of Central and South America. Its natural
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s are
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
or tropical moist lowland
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s and subtropical or tropical moist
montane forest Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
s.


Taxonomy

The red-throated caracara was described by the French polymath
Georges-Louis Leclerc, Comte de Buffon Georges-Louis Leclerc, Comte de Buffon (; 7 September 1707 – 16 April 1788) was a French naturalist, mathematician, cosmologist, and encyclopédiste. His works influenced the next two generations of naturalists, including two prominent Fr ...
in 1770 in his ''Histoire Naturelle des Oiseaux'' from a specimen collected in
Cayenne Cayenne (; ; gcr, Kayenn) is the capital city of French Guiana, an overseas region and Overseas department, department of France located in South America. The city stands on a former island at the mouth of the Cayenne River on the Atlantic Oc ...
,
French Guiana French Guiana ( or ; french: link=no, Guyane ; gcr, label=French Guianese Creole, Lagwiyann ) is an overseas departments and regions of France, overseas department/region and single territorial collectivity of France on the northern Atlantic ...
. The bird was also illustrated in a hand-coloured plate engraved by
François-Nicolas Martinet François-Nicolas Martinet (1731 - 1800) was a French engineer, engraver and naturalist. Martinet engraved the plates for numerous works on natural history, especially ornithology. Notable in particular are those for ''l'Ornithologia, sive Synop ...
in the ''Planches Enluminées D'Histoire Naturelle'' which was produced under the supervision of
Edme-Louis Daubenton Edme-Louis Daubenton (12 August 1730 – 12 December 1785) was a French naturalist. Daubenton was the cousin of another French naturalist, Louis Jean-Marie Daubenton. Georges-Louis Leclerc, the Comte de Buffon engaged Edme-Louis Daubenton to su ...
to accompany Buffon's text. Neither the plate caption nor Buffon's description included a scientific name but in 1783 the Dutch naturalist
Pieter Boddaert Pieter Boddaert (1730 – 6 May 1795) was a Dutch physician and natural history, naturalist. Early life, family and education Boddaert was the son of a Middelburg jurist and poet by the same name (1694–1760). The younger Pieter obtained his M.D ...
coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Falco americanus'' in his catalogue of the ''Planches Enluminées''. The red-throated caracara was for many years placed with the
black caracara The black caracara (''Daptrius ater'') is a species of bird of prey in the family Falconidae found in Amazonian and French Guiana lowlands, commonly along rivers. They are locally referred to as ''Ger' futu busikaka'' in the Republic of Suriname ...
in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
'' Daptrius'' but based on a molecular genetic study published in 1999 it was moved to be the only species in the resurrected genus ''Ibycter'' that had been introduced by the French ornithologist
Louis Jean Pierre Vieillot Louis Pierre Vieillot (10 May 1748, Yvetot – 24 August 1830, Sotteville-lès-Rouen) was a French ornithologist. Vieillot is the author of the first scientific descriptions and Linnaean names of a number of birds, including species he collecte ...
in 1816. The species is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
. The genus name ''Ibycter '' is from the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''ibuktēr'' meaning "singer of war-songs".


Description

Males average 20.1 in (51 cm) long, while females average 22.1 in (56 cm); they are distinguished from the
black caracara The black caracara (''Daptrius ater'') is a species of bird of prey in the family Falconidae found in Amazonian and French Guiana lowlands, commonly along rivers. They are locally referred to as ''Ger' futu busikaka'' in the Republic of Suriname ...
by larger size and
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
that is mainly black, with the belly, tail feathers, and undertail feathers being white. Both their faces and throats are bare with a few black feathers scattered on the throat; the exposed skin is red. Both male and female red-throated caracaras are similar in appearance. Males have a wing length of 35.55 cm, a tail length of 24.96 cm, a bill length of 2.5 cm, and a tarsus length of 5.41 cm. Females have a wing length of 35.93 cm, a tail length of 25.31 cm, a bill length of 2.58 cm, and a tarsus length of 5.62 cm.


Distribution and habitat

This species inhabits the humid lowland forests of
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
,
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
,
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
,
Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...
,
El Salvador El Salvador (; , meaning " The Saviour"), officially the Republic of El Salvador ( es, República de El Salvador), is a country in Central America. It is bordered on the northeast by Honduras, on the northwest by Guatemala, and on the south b ...
,
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
,
French Guiana French Guiana ( or ; french: link=no, Guyane ; gcr, label=French Guianese Creole, Lagwiyann ) is an overseas departments and regions of France, overseas department/region and single territorial collectivity of France on the northern Atlantic ...
,
Guatemala Guatemala ( ; ), officially the Republic of Guatemala ( es, República de Guatemala, links=no), is a country in Central America. It is bordered to the north and west by Mexico; to the northeast by Belize and the Caribbean; to the east by H ...
,
Guyana Guyana ( or ), officially the Cooperative Republic of Guyana, is a country on the northern mainland of South America. Guyana is an indigenous word which means "Land of Many Waters". The capital city is Georgetown. Guyana is bordered by the ...
,
Honduras Honduras, officially the Republic of Honduras, is a country in Central America. The republic of Honduras is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Oce ...
,
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
,
Nicaragua Nicaragua (; ), officially the Republic of Nicaragua (), is the largest country in Central America, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west. Managua is the cou ...
,
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...
,
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
, and
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
. The slow flight of the red-throated caracara makes it suited it to fly in the
understory In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but abov ...
of the forest where the vegetation is thin. The sparse vegetation gives the red-throated caracara greater visibility to spot food and predators. The
ornate hawk-eagle The ornate hawk-eagle (''Spizaetus ornatus'') is a fairly large bird of prey from the tropical Americas. Formerly, some authorities referred to this species as the crested hawk-eagle, a name that may cause some confusion as it is more commonly us ...
and the black-and-white hawk-eagle are predators of the red-throated caracara.


Behaviour and ecology


Food and feeding

The red-throated caracara hunts in the
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an a ...
and the understory of the lowland jungle, foraging mainly for insect nests. Most red-throated caracaras hunt silently, but occasionally make soft caws and sometimes hunt in groups. When hunting in groups, one or two individuals scout for predators in the canopy, while the remaining flock hunts in the understory. The red-throated caracara is highly territorial, with four to eight individuals in a group.Life History - Red-throated Caracara (Ibycter americanus) - Neotropical Birds
/ref> The diet consists mainly of wasp and bee larvae, though it will eat mature insects and also forage on fruits and berries found in the humid subtropical and tropical lowlands, and mountainous regions of its Central and South American habitat. Biodiversity of the forest
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syste ...
is paramount for the birds' special diet, since wasps and bees often make their nests in hollows or amongst branches of mature trees found in old-growth forests. Deforestation and intensive agriculture practices severely hamper the red-throated caracara's population, likely accounting for its rare sightings today. After the 1950s, both its population and range rapidly declined in Costa Rica, Honduras, Panama, Ecuador, and French Guiana, causing the species to be placed on the World Wildlife endangered list. Until 2013, very little was known of the red-throated caracara's feeding behavior until a team of Canadian biologists from the University of Simon Fraser spent months researching the birds using camera surveillance at the Nouragues Field Station in French Guiana. The scientific footage shows the birds using a rapid-fire “fly-by” aerial-diving attack strategy to knock nests down onto the forest floor, while skillfully evading most wasp stings. The birds use air squadron precision, repeatedly diving then scooping upward, to drive off or confuse angry defender swarms around the hive.Birds outsmart wasps to feed young - SFU News - Simon Fraser University
/ref> Researchers also found that neotropical defender wasps eventually abandon their damaged hives and retreat, alongside smaller worker wasps, to rebuild a new nest site. All predators evolve ways of hunting or trapping prey. Biologist Sean McCann observed that these intelligent birds have a highly specialized predation trait in response the wasps’ behavior to cut losses and rebuild elsewhere. The predation impact on the numbers of prey populations is undetermined. Furthermore, it is not clear how much the red-throated caracara's primary food source, wasp larvae, places constraints on the birds' ability to survive since their complex predation is interlinked with neotropical wasp behavior. Knowledge of the birds' chemical resistance to stings is also unknown. Chemical traces found on the birds’ feet are similar to those secreted from Azteca ants, likely contacted along tree branches and nest sites which both species inhabit.


See also

*
Falco Lombardi is a fictional anthropomorphic bird from the ''Star Fox'' series of video games. He was created by Shigeru Miyamoto and designed by Takaya Imamura. Falco acts as the wingman and best friend of the titular Fox McCloud for the majority of the se ...


References


Further reading

*


External links


Red-throated caracara videos
on the Internet Bird Collection

VIREO * ttp://www.pinkmoose.ic24.net/images/red-faced-caracara.jpg Photo-Medium Res ttp://www.pinkmoose.ic24.net/peru/amazonia.htm Articlepinkmoose–"Amazonia Lodge"
Photo-Medium Res
https://www.nhlstenden.com/;"Suriname Birds" {{Taxonbar, from=Q517757 Polyborinae, red-throated caracara Birds of Central America Birds of Brazil Birds of Colombia Birds of Venezuela Birds of Ecuador Birds of the Guianas Birds of the Amazon Basin
red-throated caracara The red-throated caracara (''Ibycter americanus'') is a social species of bird of prey in the family Falconidae. It is placed in the monotypic genus ''Ibycter'', or sometimes united in ''Daptrius'' with the black caracara. Unique among caracaras ...
Taxonomy articles created by Polbot