Actinopterygii (; ), members of which are known as ray-finned fishes, is a

class Class or The Class may refer to: Common uses not otherwise categorized * Class (biology), a taxonomic rank * Class (knowledge representation), a collection of individuals or objects * Class (philosophy), an analytical concept used differentl ...

bony fish Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartilag ...

. They comprise over 50% of living

vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

. The ray-finned

fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...

es are so called because their fins are webs of skin supported by bony or horny spines (rays), as opposed to the fleshy, lobed fins that characterize the class Sarcopterygii (lobe-finned fish). These actinopterygian fin rays attach directly to the proximal or basal skeletal elements, the radials, which represent the link or connection between these fins and the internal skeleton (e.g., pelvic and pectoral girdles). By species count, actinopterygians dominate the

s, and they constitute nearly 99% of the over 30,000 species of fish. They are ubiquitous throughout

freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...

and

marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...

environments from the deep sea to the highest mountain streams. Extant species can range in size from '' Paedocypris'', at , to the massive

ocean sunfish The ocean sunfish or common mola (''Mola mola'') is one of the largest bony fish in the world. It was misidentified as the heaviest bony fish, which was actually a different species, ''Mola alexandrini''. Adults typically weigh between . The spe ...

, at , and the long-bodied

oarfish Oarfish are huge, greatly elongated, pelagic lampriform fish belonging to the small family Regalecidae. Found in areas spanning from temperate ocean zones to tropical ones, yet rarely seen, the oarfish family contains three species in two gene ...

, at . The vast majority of Actinopterygii (~99%) are teleosts.

Characteristics

Ray-finned fishes occur in many variant forms. The main features of typical ray-finned fish are shown in the adjacent diagram. The swim bladder is the more derived structure. Ray-finned fishes have many different types of scales; but all teleosts have leptoid scales. The outer part of these scales fan out with bony ridges, while the inner part is crossed with fibrous connective tissue. Leptoid scales are thinner and more transparent than other types of scales, and lack the hardened enamel or dentine-like layers found in the scales of many other fish. Unlike ganoid scales, which are found in non-teleost actinopterygians, new scales are added in concentric layers as the fish grows. Ray-finned and lobe-finned fishes sometimes possesses lungs used for aerial respiration. Only bichirs retain ventrally budding lungs.

Body shapes and fin arrangements

Ray-finned fish vary in size and shape, in their feeding specializations, and in the number and arrangement of their ray-fins.

Reproduction

In nearly all ray-finned fish, the sexes are separate, and in most species the females spawn eggs that are fertilized externally, typically with the male inseminating the eggs after they are laid. Development then proceeds with a free-swimming larval stage. However other patterns of

ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...

exist, with one of the commonest being sequential hermaphroditism. In most cases this involves protogyny, fish starting life as females and converting to males at some stage, triggered by some internal or external factor.

Protandry Sequential hermaphroditism (called dichogamy in botany) is a type of hermaphroditism that occurs in many fish, gastropods, and plants. Sequential hermaphroditism occurs when the individual changes its sex at some point in its life. In particular, ...

, where a fish converts from male to female, is much less common than protogyny. Most families use external rather than internal fertilization. Of the

oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...

teleosts, most (79%) do not provide parental care.

Viviparity Among animals, viviparity is development of the embryo inside the body of the parent. This is opposed to oviparity which is a reproductive mode in which females lay developing eggs that complete their development and hatch externally from the m ...

ovoviviparity Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a term used as a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop insi ...

, or some form of parental care for eggs, whether by the male, the female, or both parents is seen in a significant fraction (21%) of the 422 teleost families; no care is likely the ancestral condition. The oldest case of viviparity in ray-finned fish is found in

Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma and ...

species of ''

Saurichthys ''Saurichthys'' (from el, σαῦρος , 'lizard' and el, ἰχθῦς 'fish') is an extinct genus of predatory ray-finned fish from the Triassic period. It type genus family Saurichthyidae (Changhsingian- Middle Jurassic), and the larges ...

''. Viviparity is relatively rare and is found in about 6% of living teleost species; male care is far more common than female care. Male territoriality "preadapts" a species for evolving male parental care. There are a few examples of fish that self-fertilise. The mangrove rivulus is an amphibious, simultaneous hermaphrodite, producing both eggs and spawn and having internal fertilisation. This mode of reproduction may be related to the fish's habit of spending long periods out of water in the mangrove forests it inhabits. Males are occasionally produced at temperatures below and can fertilise eggs that are then spawned by the female. This maintains genetic variability in a species that is otherwise highly inbred.

Classification and fossil record

Actinopterygii is divided into the classes Cladistia and Actinopteri. The latter comprises the subclasses Chondrostei and Neopterygii. The Neopterygii, in turn, is divided into the infraclasses Holostei and

Teleostei Teleostei (; Ancient Greek, Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts ), is, by far, the largest class (biology), infraclass in the class Actinopterygii, the ray-finned fishes, containing 96% of a ...

. During the

Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceo ...

(

Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...

Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

) and

Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...

the teleosts in particular diversified widely. As a result, 96% of living fish species are teleosts (40% of all fish species belong to the teleost subgroup

Acanthomorpha Acanthomorpha (meaning "thorn-shaped") is an extraordinarily diverse taxon of teleost fishes with spiny rays. The clade contains about one third of the world's modern species of vertebrates: over 14,000 species. A key anatomical innovation in ...

), while all other groups of actinopterygians represent depauperate lineages. The classification of ray-finned fishes can be summarized as follows: * Cladistia, which include bichirs and reedfish * Actinopteri, which include: ** Chondrostei, which include Acipenseriformes (paddlefishes and sturgeons) ** Neopterygii, which include: ***Teleostei (most living fishes) ***Holostei, which include: ****Lepisosteiformes (gars) ****Amiiformes (bowfin) The cladogram below shows the main

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

s of living actinopterygians and their evolutionary relationships to other

extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...

groups of

fishes Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...

and the four-limbed vertebrates ( tetrapods). The latter include mostly terrestrial

but also groups that became secondarily aquatic (e.g. Whales and Dolphins). Tetrapods

evolved Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation t ...

from a group of

during the

Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...

period. Approximate

divergence In vector calculus, divergence is a vector operator that operates on a vector field, producing a scalar field giving the quantity of the vector field's source at each point. More technically, the divergence represents the volume density of the ...

dates for the different actinopterygian clades (in millions of years, mya) are from Near et al., 2012. The polypterids (bichirs and reedfish) are the

sister lineage In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...

of all other actinopterygians, the Acipenseriformes (sturgeons and paddlefishes) are the sister lineage of Neopterygii, and Holostei (bowfin and gars) are the sister lineage of teleosts. The Elopomorpha (

eel Eels are ray-finned fish belonging to the order Anguilliformes (), which consists of eight suborders, 19 families, 111 genera, and about 800 species. Eels undergo considerable development from the early larval stage to the eventual adult stage ...

s and tarpons) appear to be the most basal teleosts. The earliest known

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

actinopterygian is ''

Andreolepis hedei ''Andreolepis'' is an extinct genus of prehistoric fish, which lived around 420 million years ago. It was described by Walter Gross in 1968 based on scales found in the Hemse Formation in Gotland, Sweden. It is placed in the monogeneric f ...

'', dating back 420 million years (

Late Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozo ...

), remains of which have been found in

Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...

Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...

, and

Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, a ...

. Crown group actinopterygians most likely originated near the Devonian-Carboniferous boundary. The earliest fossil relatives of modern teleosts are from the

period (''

Prohalecites ''Prohalecites'' is an extinct genus of ray-finned fish from the Ladinian and possibly Carnian (Triassic) of Italy. It is the oldest known teleosteomorph, a group that includes extant teleosts and their close fossil relatives. The type and on ...

'', ''

Pholidophorus ''Pholidophorus'' (from el, φολῐ́ς , 'horny scale' and el, φέρω , 'to bear') is an extinct genus of stem-teleost fish. Numerous species were assigned to this genus in the past, but only the type species ''Pholidophorus latiusculus'', ...

''), although it is suspected that teleosts originated already during the

Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...

Era An era is a span of time defined for the purposes of chronology or historiography, as in the regnal eras in the history of a given monarchy, a calendar era used for a given calendar, or the geological eras defined for the history of Earth. Compa ...

PalaeoniscusFreieslebenensis-NaturalHistoryMuseum-August23-08

Taxonomy

The listing below is a summary of all

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

(indicated by a

dagger A dagger is a fighting knife with a very sharp point and usually two sharp edges, typically designed or capable of being used as a thrusting or stabbing weapon.State v. Martin, 633 S.W.2d 80 (Mo. 1982): This is the dictionary or popular-use de ...

, †) and living groups of Actinopterygii with their respective

taxonomic rank In biological classification, taxonomic rank is the relative level of a group of organisms (a taxon) in an ancestral or hereditary hierarchy. A common system consists of species, genus, family (biology), family, order (biology), order, class (b ...

. The

taxonomy Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...

follows Phylogenetic Classification of Bony Fishes with notes when this differs from Nelson, ITIS and

FishBase FishBase is a global species database of fish species (specifically finfish). It is the largest and most extensively accessed online database on adult finfish on the web.

and extinct groups from Van der Laan 2016 and Xu 2021. * Order †?

Asarotiformes ''Asarotus'' is an extinct genus of prehistoric bony fish that lived during the Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). ...

Schaeffer 1968 * Order †? Discordichthyiformes Minikh 1998 * Order †? Paphosisciformes Grogan & Lund 2015 * Order †? Scanilepiformes Selezneya 1985 * Order †

Cheirolepidiformes ''Cheirolepis'' (from el, χείρ , 'hand' and el, λεπίς 'scale') is an extinct genus of ray-finned fish that lived in the Devonian period of Europe and North America. It is the only genus yet known within the family Cheirolepidae and t ...

Kazantseva-Selezneva 1977 * Order † Paramblypteriformes Heyler 1969 * Order † Rhadinichthyiformes * Order †

Palaeonisciformes The Palaeonisciformes (Palaeoniscida) are an extinct order of early ray-finned fishes ( Actinopterygii). Palaeonisciformes ''sensu lato'' first appeared in the fossil record in the Late Silurian and last appeared in the Late Cretaceous. The na ...

Hay 1902 * Order †

Tarrasiiformes Tarasiiformes is an extinct order of prehistoric ray-finned fish. Taxonomy * Order †Tarrasiiformes sensu Lund & Poplin 2002 aplistiaref name="mikko"> ** Family †Tarrasiidae Traquair 1881 emend. Woodward 1891 *** Genus †'' Apholidotos'' ...

sensu Lund & Poplin 2002 * Order †

Ptycholepiformes Ptycholepiformes are an extinct order of prehistoric ray-finned fish that existed during the Triassic period and the Early Jurassic epoch. The order includes the genera '' Acrorhabdus'', ''Ardoreosomus'', ''Boreosomus'', '' Chungkingichthys'', '' ...

Andrews et al. 1967 * Order † Haplolepidiformes Westoll 1944 * Order † Aeduelliformes Heyler 1969 * Order † Platysomiformes Aldinger 1937 * Order † Dorypteriformes Cope 1871 * Order † Eurynotiformes Sallan & Coates 2013 * Class Cladistia Pander 1860 ** Order †

Guildayichthyiformes Guildayichthyidae is a prehistoric family of marine fish from the Mississippian Bear Gulch Limestone of Montana. It is the only family in the order Guildayichthyiformes. Guildayichthyids possess an uncommon mixture of primitive and modern charac ...

Lund 2000 ** Order Polypteriformes Bleeker 1859 ( bichirs and

reedfish The reedfish, ropefish (more commonly used in the United States), or snakefish, ''Erpetoichthys calabaricus'', is a species of fish in the bichir family and order. It is the only member of the genus ''Erpetoichthys''. It is native to fresh and ...

es) * Class Actinopteri Cope 1972 s.s. ** Order † Elonichthyiformes Kazantseva-Selezneva 1977 ** Order † Phanerorhynchiformes ** Order †

Bobasatraniiformes Bobasatraniiformes is an extinct Order (biology), order of durophagous ray-finned fish that existed from the Lopingian, late Permian to the Middle Triassic in both marine life, marine and freshwater environments. The order includes two family ...

Berg 1940 ** Order † Saurichthyiformes Aldinger 1937 ** Subclass Chondrostei Müller, 1844 *** Order † Birgeriiformes Heyler 1969 *** Order † Chondrosteiformes Aldinger, 1937 *** Order

Acipenseriformes Acipenseriformes is an order of basal ray-finned fishes that includes living and fossil sturgeons and paddlefishes (Acipenseroidei), as well as the extinct families Chondrosteidae and Peipiaosteidae. They are the second earliest div ...

Berg 1940 (includes

sturgeon Sturgeon is the common name for the 27 species of fish belonging to the family Acipenseridae. The earliest sturgeon fossils date to the Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretace ...

s and

paddlefish Paddlefish (family Polyodontidae) are a family of ray-finned fish belonging to order Acipenseriformes, and one of two living groups of the order alongside sturgeons (Acipenseridae). They are distinguished from other fish by their titular elongla ...

es) ** Subclass Neopterygii Regan 1923 sensu Xu & Wu 2012 *** Order †

Pholidopleuriformes Pholidopleuriformes is an extinct order of ray-finned fish. Classification ** Family †Pholidopleuridae Abel 1919/Wade 1932 *** Genus †'' Gracilignathichthys'' Bürgin 1992 **** †'' Gracilignathichthys microlepis'' Bürgin 1992 *** Genus � ...

Berg 1937 ***Order †

Redfieldiiformes Redfieldiiformes is an extinct Order (biology), order of ray-finned fish (actinopterygians) which lived from the Middle Triassic to Early Jurassic. Redfieldiiforms were fairly typical Triassic fish in overall anatomy. They had a fusiform (streaml ...

Berg 1940 ***Order †

Platysiagiformes Platysiagidae is an extinct family of stem-neopterygian ray-finned fish which lived from the Early Triassic to the Early Jurassic. It includes the genera '' Helmolepis'', '' Platysiagum'' and possibly '' Caelatichthys''. The family was formerly p ...

Brough 1939 ***Order † Polzbergiiformes Griffith 1977 ***Order †

Perleidiformes Perleidiformes are an extinct order of prehistoric ray-finned fish from the Triassic period Although numerous Triassic taxa have been referred to Perleidiformes, which ones should be included for it to form a monophyletic group is a matter of on ...

Berg 1937 ***Order † Louwoichthyiformes Xu 2021 *** Order †

Peltopleuriformes Peltopleuriformes is an extinct order of ray-finned fish. Classification * Order †Peltopleuriformes Lehman 1966 eltopleuroidei Lehman 1966** Family †Peltopleuridae Bough 1939 *** Genus †''Marcopoloichthys'' Tintori et al. 2008 *** Genu ...

Lehman 1966 *** Order †

Luganoiiformes ''Luganoia'' is an extinct genus of prehistoric bony fish that lived during the early Ladinian stage of the Middle Triassic epoch. See also * Prehistoric fish * List of prehistoric bony fish A ''list'' is any set of items in a row. List or lis ...

Lehman 1958 *** Order †

Pycnodontiformes Pycnodontiformes is an extinct order of primarily marine bony fish. The group first appeared during the Late Triassic and disappeared during the Eocene. The group has been found in rock formations in Africa, Asia, Europe, North and South America ...

Berg 1937 *** Infraclass Holostei Müller 1844 **** Division

Halecomorphi Halecomorphi is a taxon of ray-finned bony fish in the clade Neopterygii. The sole living Halecomorph is the bowfin (''Amia calva''), but the group contains many extinct species in several families (including Amiidae, Caturidae, Liodesmidae, S ...

Cope 1872 sensu Grande & Bemis 1998 ***** Order †

Parasemionotiformes Parasemionotiformes is an extinct order of neopterygian ray-finned fish that existed globally during the Triassic period. It comprises the families Parasemionotidae and Promecosominidae. Many of the included genera are monotypic and most speci ...

Lehman 1966 ***** Order † Ionoscopiformes Grande & Bemis 1998 ***** Order Amiiformes Huxley 1861 sensu Grande & Bemis 1998 (

bowfin The bowfin (''Amia calva'') is a bony fish, native to North America. Common names include mudfish, mud pike, dogfish, grindle, grinnel, swamp trout, and choupique. It is regarded as a relict, being the sole surviving species of the Halecomorphi ...

s) **** Division Ginglymodi Cope 1871 ***** Order † Dapediiformes Thies & Waschkewitz 2015 ***** Order † Semionotiformes Arambourg & Bertin 1958 ***** Order Lepisosteiformes Hay 1929 (

gar Gars are members of the family Lepisosteidae, which are the only surviving members of the Ginglymodi, an ancient holosteian group of ray-finned fish, which first appeared during the Triassic, over 240 million years ago. Gars comprise seven livin ...

s) *** Clade Teleosteomorpha Arratia 2000 sensu Arratia 2013 **** Order †

Prohaleciteiformes ''Prohalecites'' is an extinct genus of ray-finned fish from the Ladinian and possibly Carnian (Triassic) of Italy. It is the oldest known teleosteomorph, a group that includes extant teleosts and their close fossil relatives. The type and o ...

Arratia 2017 **** Division Aspidorhynchei Nelson, Grand & Wilson 2016 ***** Order †

Aspidorhynchiformes Aspidorhynchiformes (from New Latin "shield-snout forms") is an extinct order of ray-finned fish. It contains only a single family, the Aspidorhynchidae. Members of the group are noted for their elongated, conical rostrums, of varying length, fo ...

Bleeker 1859 ***** Order †

Pachycormiformes Pachycormiformes is an extinct order of marine ray-finned fish known from the Early Jurassic to the end of the Cretaceous. It only includes a single family, Pachycormidae. They were characterized by having serrated pectoral fins (though more rece ...

Berg 1937 **** Infraclass

Müller 1844 sensu Arratia 2013 ***** Order †?

Araripichthyiformes ''Araripichthys'' is an extinct genus of ray-finned fish that lived from the Aptian to Coniacian stages of the Cretaceous period. The genus is named after the Araripe Basin, where it was found in the Crato Formation, Crato and Santana Formations. ...

***** Order †? Ligulelliiformes Taverne 2011 ***** Order †?

Tselfatiiformes Tselfatiiformes is an extinct order of bony fishes from the infraclass Teleostei. The order represents the most important radiation of marine teleosts during the Cretaceous period. Fossils of tselfatiiforms are known from Europe, North America, c ...

Nelson 1994 ***** Order † Pholidophoriformes Berg 1940 ***** Order †

Dorsetichthyiformes ''Dorsetichthys'' is an extinct genus of stem-teleost ray-finned fish from the Early Jurassic period of Europe. Description ''Dorsetichthys'' was a herring-like fish about long, although it was not closely related to modern herring. Like them, ...

Nelson, Grand & Wilson 2016 ***** Order †

Leptolepidiformes Leptolepidae (also spelt as Leptolepididae) is an extinct family of herring-like stem-teleost fish found throughout the world during the Jurassic. They were among the first fish to possess certain teleost synapomorphies, such as cycloid scale ...

***** Order † Crossognathiformes Taverne 1989 ***** Order †

Ichthyodectiformes Ichthyodectiformes is an extinct order of marine stem-teleost ray-finned fish. The order is named after the genus '' Ichthyodectes'', established by Edward Drinker Cope in 1870. Ichthyodectiforms are usually considered to be some of the closest ...

Bardeck & Sprinkle 1969 ***** Teleocephala de Pinna 1996 s.s. ******Megacohort Elopocephalai Patterson 1977 sensu Arratia 1999 ( Elopomorpha Greenwood et al. 1966) ******* Order Elopiformes Gosline 1960 ( ladyfishes and tarpon) ******* Order Albuliformes Greenwood et al. 1966 sensu Forey et al. 1996 (bonefishes) ******* Order Notacanthiformes Goodrich 1909 (

halosaurs Halosaurs are eel-shaped fishes found only at great ocean depths. As the family Halosauridae, halosaurs are one of two families within the order Notacanthiformes; the other being the deep-sea spiny eels, Notacanthidae. Halosaurs are thought to ha ...

and spiny eels) ******* Order

Anguilliformes Eels are ray-finned fish belonging to the order Anguilliformes (), which consists of eight suborders, 19 families, 111 genera, and about 800 species. Eels undergo considerable development from the early larval stage to the eventual adult stage ...

Jarocki 1822 sensu Goodrich 1909 (true

s) ****** Megacohort Osteoglossocephalai sensu Arratia 1999 *******Supercohort Osteoglossocephala sensu Arratia 1999 (

Osteoglossomorpha Osteoglossomorpha is a group of bony fish in the Teleostei. Notable members A notable member is the arapaima (''Arapaima gigas''), the largest freshwater fish in South America and one of the largest bony fishes alive. Other notable members incl ...

Greenwood et al. 1966) ******** Order † Lycopteriformes Chang & Chou 1977 ******** Order

Hiodontiformes Hiodontiformes is an order of ray-finned fish consisting of the two living species of the mooneye family, Hiodontidae, and three extinct genera. These are traditionally classified within the order Osteoglossiformes, a placement some authorit ...

McAllister 1968 sensu Taverne 1979 ( mooneye and

goldeye The goldeye (''Hiodon alosoides'') is a freshwater fish found in Canada and the northern United States. It is one of only two extant species in the family Hiodontidae, the other species being ''Hiodon tergisus''. The species name ''alosoides'' ...

) ******** Order Osteoglossiformes Regan 1909 sensu Zhang 2004 (bony-tongued fishes) ******* Supercohort Clupeocephala Patterson & Rosen 1977 sensu Arratia 2010 ********Cohort Otomorpha Wiley & Johnson 2010 (

Otocephala Otocephala is a clade of ray-finned fishes within the infraclass Teleostei that evolved some 230 million years ago. It is named for the presence of a hearing (otophysic) link from the swimbladder to the inner ear. Other names proposed for the gro ...

; Ostarioclupeomorpha) ********* Subcohort Clupei Wiley & Johnson 2010 ( Clupeomorpha Greenwood et al. 1966) ********** Order † Ellimmichthyiformes Grande 1982 ********** Order Clupeiformes Bleeker 1859 (

herring Herring are forage fish, mostly belonging to the family of Clupeidae. Herring often move in large schools around fishing banks and near the coast, found particularly in shallow, temperate waters of the North Pacific and North Atlantic Oceans, i ...

s and anchovies) ********* Subcohort Alepocephali ********** Order

Alepocephaliformes Alepocephaliformes is an order of ray-finned fish. It was previously classified as the suborder Alepocephaloidei of the order Argentiniformes The Argentiniformes are an order of ray-finned fish whose distinctness was recognized only fairly ...

Marshall 1962 ********* Subcohort

Ostariophysi Ostariophysi is the second-largest superorder of fish. Members of this superorder are called ostariophysians. This diverse group contains 10,758 species, about 28% of known fish species in the world and 68% of freshwater species, and are present ...

Sagemehl 1885 ********** Section Anotophysa (Rosen & Greenwood 1970) Sagemehl 1885 *********** Order † Sorbininardiformes Taverne 1999 *********** Order

Gonorynchiformes The Gonorynchiformes are an order of ray-finned fish that includes the important food source, the milkfish (''Chanos chanos'', family Chanidae), and a number of lesser-known types, both marine and freshwater. The alternate spelling "Gonorhyn ...

Regan 1909 ( milkfishes) ********** Section Otophysa Garstang 1931 *********** Order

Cypriniformes Cypriniformes is an order of ray-finned fish, including the carps, minnows, loaches, and relatives. Cypriniformes is an Order within the Superorder Ostariophysi consisting of "Carp-like" Ostariophysins. This order contains 11-12 families, ...

Bleeker 1859 sensu Goodrich 1909 ( barbs,

carp Carp are various species of oily freshwater fish from the family Cyprinidae, a very large group of fish native to Europe and Asia. While carp is consumed in many parts of the world, they are generally considered an invasive species in parts of ...

danios ''Danio'' is a genus of small freshwater fish in the family Cyprinidae found in South and Southeast Asia, commonly kept in aquaria. They are generally characterised by a pattern of horizontal stripes, rows of spots or vertical bars. Some species ...

, goldfishes,

loaches Loaches are fish of the superfamily Cobitoidea. They are freshwater, benthic (bottom-dwelling) fish found in rivers and creeks throughout Eurasia and northern Africa. Loaches are among the most diverse groups of fish; the 1249 known species of ...

, minnows, rasboras) *********** Order Characiformes Goodrich 1909 (

characin Characiformes is an order of ray-finned fish, comprising the characins and their allies. Grouped in 18 recognized families, more than 2000 different species are described, including the well-known piranha and tetras.; Buckup P.A.: "Relationshi ...

pencilfish ''Nannostomus'', (from the Greek: ''nanos'' = small, and the Latin: ''stomus'' = relating to the mouth), is a genus of fish belonging to the characin family Lebiasinidae. All of the species in this genus are known as pencil fish or pencilfish, a ...

es, hatchetfishes,

piranha A piranha or piraña (, , or ; or , ) is one of a number of freshwater fish in the family Serrasalmidae, or the subfamily Serrasalminae within the tetra family, Characidae in order Characiformes. These fish inhabit South American rivers, ...

s, tetras, dourado / golden (genus ''Salminus'') and

pacu Pacu () is a common name used to refer to several species of omnivorous South American freshwater serrasalmid fish that are related to the piranha. Pacu and piranha do not have similar teeth, the main difference being jaw alignment; piranha hav ...

) *********** Order

Gymnotiformes The Gymnotiformes are an order of teleost bony fishes commonly known as Neotropical knifefish or South American knifefish. They have long bodies and swim using undulations of their elongated anal fin. Found almost exclusively in fresh water (the ...

Berg 1940 ( electric eels and knifefishes) *********** Order Siluriformes Cuvier 1817 sensu Hay 1929 (

catfish Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive, ...

es) ******** Cohort Euteleosteomorpha (Greenwood et al. 1966) (

Euteleostei Euteleostei, whose members are known as euteleosts, is a clade of bony fishes within Teleostei that evolved some 240 million years ago. It is divided into Protacanthopterygii (including the salmon and dragonfish) and Neoteleostei (including t ...

Greenwood 1967 sensu Johnson & Patterson 1996) *********Subcohort Lepidogalaxii ********** Lepidogalaxiiformes Betancur-Rodriguez et al. 2013 (

salamanderfish ''Lepidogalaxias salamandroides'' is a species of small fish of Western Australia. It is the monotypic, only member of the family Lepidogalaxiidae and genus ''Lepidogalaxias''. Common names for this fish include salamanderfish and Shannon mudminn ...

) ********* Subcohort Protacanthopterygii Greenwood et al. 1966 sensu Johnson & Patterson 1996 ********** Order Argentiniformes ( barreleyes and

slickhead Slickheads, also known as nakedheads or smoothheads, are deep water fishes that belong to the family Alepocephalidae. They are most commonly found in the bathypelagic layer, which is approximately 3000m below the surface. They get their name from ...

s) (formerly in Osmeriformes) ********** Order Galaxiiformes ********** Order Salmoniformes Bleeker 1859 sensu Nelson 1994 (

salmon Salmon () is the common name for several list of commercially important fish species, commercially important species of euryhaline ray-finned fish from the family (biology), family Salmonidae, which are native to tributary, tributaries of the ...

and

trout Trout are species of freshwater fish belonging to the genera '' Oncorhynchus'', ''Salmo'' and ''Salvelinus'', all of the subfamily Salmoninae of the family Salmonidae. The word ''trout'' is also used as part of the name of some non-salmoni ...

) ********** Order

Esociformes The Esociformes () are a small order of ray-finned fish, with two families, Umbridae and Esocidae. The pikes of genus '' Esox'' give the order its name. This order is closely related to the Salmoniformes, the two comprising the superorder Prot ...

Bleeker 1859 (

pike Pike, Pikes or The Pike may refer to: Fish * Blue pike or blue walleye, an extinct color morph of the yellow walleye ''Sander vitreus'' * Ctenoluciidae, the "pike characins", some species of which are commonly known as pikes * ''Esox'', genus of ...

) ********* Subcohort Stomiati ********** Order

Osmeriformes The Osmeriformes are an order of ray-finned fish that includes the true or freshwater smelts and allies, such as the galaxiids and noodlefishes; they are also collectively called osmeriforms. They belong to the teleost superorder Protacanthopte ...

( smelts) ********** Order

Stomiatiformes Stomiiformes is an order of deep-sea ray-finned fishes of very diverse morphology. It includes, for example, dragonfishes, lightfishes (Gonostomatidae and Phosichthyidae), loosejaws, marine hatchetfishes and viperfishes. The order contains 4 ...

Regan 1909 (

bristlemouth The Gonostomatidae are a family of mesopelagic marine fish, commonly named bristlemouths, lightfishes, or anglemouths. It is a relatively small family, containing only eight known genera and 32 species. However, bristlemouths make up for their l ...

s and

marine hatchetfish Marine hatchetfishes or deep-sea hatchetfishes are small deep-sea mesopelagic ray-finned fish of the stomiiform subfamily Sternoptychinae. They should not be confused with the freshwater hatchetfishes, which are not particularly closely relate ...

es) ********* Subcohort

Neoteleostei The Neoteleostei is a large clade of bony fish that includes the Ateleopodidae (jellynoses), Aulopiformes (lizardfish), Myctophiformes (lanternfish), Polymixiiformes (beardfish), Percopsiformes (Troutperches), Gadiformes (cods), Zeiformes (dorie ...

Nelson 1969 **********Infracohort Ateleopodia *********** Order Ateleopodiformes (

jellynose fish The jellynose fishes or tadpole fishes are the small order Ateleopodiformes. This group of ray-finned fish is monotypic, containing a single family Ateleopodidae. It has about a dozen species in four genera, but these enigmatic fishes are in n ...

) ********** Infracohort Eurypterygia Rosen 1973 ***********Section Aulopa yclosquamata Rosen 1973************ Order Aulopiformes Rosen 1973 ( Bombay duck and lancetfishes) *********** Section Ctenosquamata Rosen 1973 ************Subsection Myctophata copelomorpha************* Order

Myctophiformes The Myctophiformes are an order of ray-finned fishes consisting of two families of deep-sea marine fish, most notably the highly abundant lanternfishes (Myctophidae). The blackchins (Neoscopelidae) contain six species in three genera, while t ...

Regan 1911 (

lanternfish Lanternfishes (or myctophids, from the Greek μυκτήρ ''myktḗr'', "nose" and ''ophis'', "serpent") are small mesopelagic fish of the large family Myctophidae. One of two families in the order Myctophiformes, the Myctophidae are represente ...

es) ************ Subsection

Betancur-Rodriguez et al. 2013 *************Division Lampridacea Betancur-Rodriguez et al. 2013 ampridomorpha; Lampripterygii************** Order Lampriformes Regan 1909 (

opah Opahs, also commonly known as moonfish, sunfish (not to be confused with Molidae), kingfish, redfin ocean pan are large, colorful, deep-bodied pelagic lampriform fishes comprising the small family Lampridae (also spelled Lamprididae). The famil ...

and

ribbonfishes The ribbonfish are any lampriform fishes in the family Trachipteridae. These pelagic fish are named for their slim, ribbon-like appearance. They are rarely seen alive, as they typically live in deep waters, though are not bottom feeders. The p ...

) ************* Division Paracanthomorphacea sensu Grande et al. 2013 (

Paracanthopterygii Paracanthopterygii is a superorder of fishes. Members of this group are called paracanthopterygians. It includes five orders: * † Sphenocephaliformes * Percopsiformes ( trout-perches & allies) * Zeiformes (dories) * Stylephoriformes ( Tube ...

Greenwood 1937) ************** Order

Percopsiformes The Percopsiformes are a small order of ray-finned fishes, comprising the trout-perch and its allies. It contains just ten extant species, grouped into seven genera and three families. Five of these genera are monotypic In biology, a mono ...

Berg 1937 ( cavefishes and

trout-perch ''Percopsis omiscomaycus'', also known as the trout-perch, the grounder or the sand minnow, is one of two species in the family Percopsidae. Its name comes from the Greek root words ''perc'', meaning perch and ''opsi'' meaning appearance. The sp ...

es) ************** Order † Sphenocephaliformes Rosen & Patterson 1969 ************** Order

Zeiformes The Zeiformes are a small order of marine ray-finned fishes most notable for the dories, a group of common food fish. The order consists of about 33 species in seven families, mostly deep-sea types. Zeiform bodies are usually thin and deep. Mo ...

Regan 1909 ( dories) ************** Order

Stylephoriformes ''Stylephorus chordatus'', the tube-eye or thread-tail, is a deep-sea fish, the only fish in the genus ''Stylephorus'' and family Stylephoridae. It is found in deep subtropical and tropical waters around the world, living at depths during the da ...

Miya et al. 2007 ************** Order

Gadiformes Gadiformes are an order of ray-finned fish, also called the Anacanthini, that includes the cod. Many major food fish are in this order. They are found in marine waters throughout the world and the vast majority of the species are found in temper ...

Goodrich 1909 (

cod Cod is the common name for the demersal fish genus '' Gadus'', belonging to the family Gadidae. Cod is also used as part of the common name for a number of other fish species, and one species that belongs to genus ''Gadus'' is commonly not call ...

s) ************* Division Polymixiacea Betancur-Rodriguez et al. 2013 (Polymyxiomorpha; Polymixiipterygii) ************** Order † Pattersonichthyiformes Gaudant 1976 ************** Order † Ctenothrissiformes Berg 1937 ************** Order

Polymixiiformes The beardfishes consist of a single extant genus, ''Polymixia'', of deep-sea marine ray-finned fish named for their pair of long hyoid barbels. They are classified in their own order Polymixiiformes . But as Nelson says, "few groups have been sh ...

Lowe 1838 (

beardfish The beardfishes consist of a single extant genus, '' Polymixia'', of deep-sea marine ray-finned fish named for their pair of long hyoid barbels. They are classified in their own order Polymixiiformes . But as Nelson says, "few groups have been s ...

es) ************* Division Euacanthomorphacea Betancur-Rodriguez et al. 2013 (Euacanthomorpha sensu Johnson & Patterson 1993; Acanthopterygii Gouan 1770 sensu]) **************Subdivision Berycimorphaceae Betancur-Rodriguez et al. 2013 *************** Order Beryciformes (

fangtooth Fangtooths are beryciform fish of the family (biology), family Anoplogastridae (sometimes spelled "Anoplogasteridae") that live in the deep sea. The name is from the Greek language, Greek , meaning "unarmed", and (), meaning "stomach". With a ci ...

s and

pineconefish Pinecone fishes are small and unusual marine fish of the family ''Monocentridae''. The family contains just four species in two genera, one of which is monotypic. Their distribution is limited to tropical and subtropical waters of the Indo-Pac ...

es) (incl. Stephanoberyciformes; Cetomimiformes) ************** Subdivision Holocentrimorphaceae Betancur-Rodriguez et al. 2013 *************** Order Holocentriformes (

Soldierfish Myripristinae is a subfamily of the Holocentridae Holocentridae is a family of ray-finned fish, the only family of the order Holocentriformes. The members of the subfamily Holocentrinae are typically known as squirrelfish, while the members of ...

es) ************** Subdivision Percomorphaceae Betancur-Rodriguez et al. 2013 ( Percomorpha sensu Miya et al. 2003;

Acanthopteri Perciformes (), also called the Percomorpha or Acanthopteri, is an order or superorder of ray-finned fish. If considered a single order, they are the most numerous order of vertebrates, containing about 41% of all bony fish. Perciformes means ...

) ***************Series Ophidiimopharia Betancur-Rodriguez et al. 2013 **************** Order

Ophidiiformes Ophidiiformes is an order of ray-finned fish that includes the cusk-eels (family Ophidiidae), pearlfishes (family Carapidae), viviparous brotulas (family Bythitidae), and others. Members of this order have small heads and long slender bodies. Th ...

( pearlfishes) *************** Series Batrachoidimopharia Betancur-Rodriguez et al. 2013 **************** Order

Batrachoidiformes Batrachoididae is the only family in the ray-finned fish order Batrachoidiformes . Members of this family are usually called toadfish, or "frogfish": both the English common name and scientific name refer to their toad-like appearance (''batrakh ...

( toadfishes) *************** Series Gobiomopharia Betancur-Rodriguez et al. 2013 **************** Order

Kurtiformes The Kurtiformes consist of two extant families of ray-finned fish, the Indo-Pacific Kurtidae (consisting solely of two species in the genus '' Kurtus'') and the much more diverse and widespread Apogonidae (the cardinalfishes). The order is p ...

(

Nurseryfish ''Kurtus'' is a genus of percomorph fishes, called the nurseryfishes, forehead brooders, or incubator fish, native to fresh, brackish and coastal marine waters ranging from India, through southeast Asia to New Guinea and northern Australia. ''Ku ...

es and cardinalfishes) **************** Order

Gobiiformes The Gobiiformes are an order of fish that includes the gobies and their relatives. The order, which was previously considered a suborder of Perciformes, is made up of about 2,211 species that are divided between seven families. Phylogenetic rel ...

(Sleepers and gobies) *************** Series Scombrimopharia Betancur-Rodriguez et al. 2013 **************** Order Syngnathiformes (

seahorses A seahorse (also written ''sea-horse'' and ''sea horse'') is any of 46 species of small marine fish in the genus ''Hippocampus''. "Hippocampus" comes from the Ancient Greek (), itself from () meaning "horse" and () meaning "sea monster" or " ...

, pipefishes,

sea moth The seamoths make up a family of fishes, the Pegasidae, within the order Syngnathiformes. They are named for Pegasus, a creature from Greek mythology. Seamoths are notable for their unusual appearance, including flattened bodies, the presence of ...

cornetfish The cornetfishes or flutemouths are a small family, the Fistulariidae, of extremely elongated fishes in the order Syngnathiformes. The family consists of a single genus, ''Fistularia'', with four species, found worldwide in tropical and subtropic ...

es and

flying gurnard The flying gurnard (''Dactylopterus volitans''), also known as the helmet gurnard, is a bottom-dwelling fish of tropical to warm temperate waters on both sides of the Atlantic. On the American side, it is found as far north as Massachusetts (ex ...

sIn Nelson and ITIS, Syngnathiformes is placed as the suborder Syngnathoidei of the order

Gasterosteiformes Gasterosteoidei is a suborder of ray-finned fishes that includes the sticklebacks and relatives, the 5th edition of ''Fishes of the World'' classifies this suborder within the order Scorpaeniformes. Systematics Gasterosteoidei is treated as a s ...

.) **************** Order

Scombriformes Scombriformes is an Order (biology), order of bony fish containing nine Family (Biology), families which were classified under the suborders Scombroidei and Stromateoidei, of the wider grouping known as Perciformes, ''Fishes of the World'', 5th e ...

(

Tuna A tuna is a saltwater fish that belongs to the tribe Thunnini, a subgrouping of the Scombridae (mackerel) family. The Thunnini comprise 15 species across five genera, the sizes of which vary greatly, ranging from the bullet tuna (max length: ...

s and (

mackerel Mackerel is a common name applied to a number of different species of pelagic fish, mostly from the family Scombridae. They are found in both temperate and tropical seas, mostly living along the coast or offshore in the oceanic environment. ...

s) *************** Series Carangimopharia Betancur-Rodriguez et al. 2013 **************** Subseries Anabantaria Betancur-Rodriguez et al. 2014 ***************** Order

Synbranchiformes Synbranchiformes, often called swamp eels, is an order of ray-finned fishes that are eel-like but have spiny rays, indicating that they belong to the superorder Acanthopterygii. Taxonomy No synbrachiform fossil is known. The Mastacembeloidei were ...

( swamp eels) ***************** Order Anabantiformes (Labyrinthici) ( gouramies, snakeheads, ) **************** Subseries Carangaria Betancur-Rodriguez et al. 2014 ***************** Carangaria incertae sedis ***************** Order

Istiophoriformes Istiophoriformes are an order of bony fish which is not fully recognized by some taxonomists, with some including the two extant families Xiphiidae and Istiophoridae, and others including the family Sphyraenidae. Families The following familie ...

Betancur-Rodriguez 2013 ( Marlins,

swordfishes Swordfish (''Xiphias gladius''), also known as broadbills in some countries, are large, highly migratory predatory fish characterized by a long, flat, pointed bill. They are a popular sport fish of the billfish category, though elusive. Swordfis ...

, billfishes) ***************** Order

Carangiformes Carangiformes is an order of the ray-finned fishes. The order is part of a clade which is a sister clade to the Ovalentaria, the other orders in the clade being Synbranchiformes, Anabantiformes, Istiophoriformes and Pleuronectiformes. The Carang ...

(

Jack mackerel Jack mackerels or saurels are marine fish in the genus ''Trachurus'' of the family Carangidae. The name of the genus derives from the Greek words ''trachys'' ("rough") and ''oura'' ("tail"). Some species, such as ''T. murphyi'', are harvested in ...

s, pompanos) ***************** Order

Pleuronectiformes A flatfish is a member of the ray-finned demersal fish order Pleuronectiformes, also called the Heterosomata, sometimes classified as a suborder of Perciformes. In many species, both eyes lie on one side of the head, one or the other migratin ...

Bleeker 1859 (

flatfish A flatfish is a member of the Ray-finned fish, ray-finned demersal fish order (biology), order Pleuronectiformes, also called the Heterosomata, sometimes classified as a suborder of Perciformes. In many species, both eyes lie on one side of the ...

es) **************** Subseries Ovalentaria Smith & Near 2012 (

Stiassnyiformes Stiassnyiformes is an order of bony fish (Teleostei) proposed in 2009.Blaise Li, Agnès Dettaï, Corinne Cruaud, Arnaud Couloux, Martine Desoutter-Meniger, Guillaume Lecointre: ''RNF213, a new nuclear marker for acanthomorph phylogeny.'' Molecul ...

sensu Li et al. 2009) ***************** Ovalentaria incertae sedis ***************** Order

Cichliformes Cichliformes is an order of fishes. Its members were previously classified under the order Perciformes, but now many authorities consider it to be an independent order within the subseries Ovalentaria. Families There are two families within th ...

Betancur-Rodriguez et al. 2013 (

Cichlid Cichlids are fish from the family Cichlidae in the order Cichliformes. Cichlids were traditionally classed in a suborder, the Labroidei, along with the wrasses ( Labridae), in the order Perciformes, but molecular studies have contradicted this ...

Convict blenny ''Pholidichthys leucotaenia'', commonly known as the convict blenny/goby or the engineer blenny/goby, is a marine fish from the west-central Pacific Ocean. Despite its common names, it is neither a blenny nor a goby, but is in fact one of two spe ...

, leaf fishes) ***************** Order

Atheriniformes The Atheriniformes, also known as the silversides, are an order (biology), order of ray-finned fishes that includes the Atherinidae, Old World silversides and several less-familiar families, including the unusual Phallostethidae. The order inclu ...

Rosen 1964 ( silversides and

rainbowfish The rainbowfish or Melanotaeniidae is a family of small, colourful freshwater fish found in northern and eastern Australia, New Guinea (including islands in Cenderawasih Bay and Raja Ampat Islands in Indonesia), Sulawesi and Madagascar. The lar ...

es) ***************** Order Cyprinodontiformes Berg 1940 ( livebearers, killifishes) ***************** Order Beloniformes Berg 1940 (

flyingfish The Exocoetidae are a family of marine fish in the order Beloniformes class Actinopterygii, known colloquially as flying fish or flying cod. About 64 species are grouped in seven to nine genera. While they cannot fly in the same way a bird ...

es and

ricefish The ricefishes are a family (Adrianichthyidae) of small ray-finned fish that are found in fresh and brackish waters from India to Japan and out into the Malay Archipelago, most notably Sulawesi (where the Lake Poso and Lore Lindu species are kn ...

es) ***************** Order

Mugiliformes The mullets or grey mullets are a family (Mugilidae) of ray-finned fish found worldwide in coastal temperate and tropical waters, and some species in fresh water. Mullets have served as an important source of food in Mediterranean Europe since R ...

Berg 1940 ( mullets) ***************** Order

Blenniiformes Blenny (from the Greek and , mucus, slime) is a common name for many types of fish, including several families of percomorph marine, brackish, and some freshwater fish sharing similar morphology and behaviour. Six families are considered "true ...

Springer 1993 ( Blennies) ***************** Order

Gobiesociformes Clingfishes are fishes of the family Gobiesocidae, the only family in the order Gobiesociformes. These fairly small to very small fishes are widespread in tropical and temperate regions, mostly near the coast, but a few species in deeper seas or ...

Gill 1872 (

Clingfish Clingfishes are fishes of the family Gobiesocidae, the only family in the order Gobiesociformes. These fairly small to very small fishes are widespread in tropical and temperate regions, mostly near the coast, but a few species in deeper seas or ...

es) *************** Series Eupercaria Betancur-Rodriguez et al. 2014 (Percomorpharia Betancur-Rodriguez et al. 2013) ****************

Eupercaria incertae sedis Percomorpha () is a large clade of ray-finned fish that includes the tuna, seahorses, gobies, cichlids, flatfish, wrasse, perches, anglerfish, and pufferfish. Evolution Percomorpha are the most diverse group of teleost fish today. Teleosts, a ...

**************** Order

Gerreiformes The mojarras are a family, Gerreidae, of fish in the order Perciformes. The family includes about 53 species found worldwide in tropical and warm temperate regions. They mostly inhabit coastal salt and brackish waters, although some occur in fre ...

( Mojarras) **************** Order

Labriformes Labriformes is an order of ray-finned fishes which includes the wrasse, cales and parrotfishes, within the clade Percomorpha. Some authors include the Labroformes as the clade Labroidei within the Perciformes while others include more families ...

(

Wrasse The wrasses are a family, Labridae, of marine fish, many of which are brightly colored. The family is large and diverse, with over 600 species in 81 genera, which are divided into 9 subgroups or tribes. They are typically small, most of them le ...

s and

Parrotfish Parrotfishes are a group of about 90 fish species regarded as a Family (biology), family (Scaridae), or a subfamily (Scarinae) of the wrasses. With about 95 species, this group's largest species richness is in the Indo-Pacific. They are found ...

es) **************** Order

Caproiformes Caproidae, or boarfishes, are a small family of marine fishes comprising two genera and 12 species. They were formerly placed in the order Zeiformes with the dories, but are now placed with the Perciformes since they have many perciform charact ...

( Boarfishes) **************** Order

Lophiiformes The anglerfish are fish of the teleost order Lophiiformes (). They are bony fish named for their characteristic mode of predation, in which a modified luminescent fin ray (the esca or illicium) acts as a lure for other fish. The luminescence c ...

Garman 1899 (

Anglerfish The anglerfish are fish of the teleost order Lophiiformes (). They are bony fish named for their characteristic mode of predation, in which a modified luminescent fin ray (the esca or illicium) acts as a lure for other fish. The luminescence ...

es) **************** Order Tetraodontiformes Regan 1929 (

Filefish The filefish (Monacanthidae) are a diverse family of tropical to subtropical tetraodontiform marine fish, which are also known as foolfish, leatherjackets or shingles. They live in the Atlantic, Pacific and Indian Oceans. Filefish are closely rel ...

es and

pufferfish Tetraodontidae is a family of primarily marine and estuarine fish of the order Tetraodontiformes. The family includes many familiar species variously called pufferfish, puffers, balloonfish, blowfish, blowies, bubblefish, globefish, swellfis ...

) **************** Order

Centrarchiformes Centrarchiformes is an obsolete order of ray-finned fish, now included amongst the perciformes, with 17 previously included families. This order first appeared about 55.8 million years ago in the Eocene Era, and is composed primarily of omnivore ...

Bleeker 1859 ( Sunfishes and mandarin fishes) ****************Order

(

Stickleback The sticklebacks are a family of ray-finned fishes, the Gasterosteidae which have a Holarctic distribution in fresh, brackish and marine waters. They were thought to be related to the pipefish and seahorses but are now thought to be more closel ...

s and relatives) ****************Order Scorpaeniformes ( Lionfishes and relatives) **************** Order Perciformes Bleeker 1859

References

External links

* * {{Authority control

Ray-finned fishes Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...

Fish classes Silurian bony fish Extant Silurian first appearances