Quatsinoporites
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''Quatsinoporites'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
monotypic genus of agaricomycet fungus in the Agaricomycetes family Hymenochaetaceae. At present it contains the single species ''Quatsinoporites cranhamii''. The genus is solely known from the Barremian aged, Apple Bay deposits of northwestern Vancouver Island. ''Quatsinoporites'' is one of only three known fossil fungus species found on Vancouver Island and the only to be described from the Apple Bay strata. The agaricomycete '' Appianoporites vancouverensis'' was described from an Eocene fossil at the same time as ''Quatsinoporites '', while a third fungus, '' Margaretbarromyces dictyosporus'' was described three years later. __TOC__


History and classification

The genus is known only from the single holotype, a partial bracket fungus fruiting body, or " conk." The specimen, P13021 E, is currently residing in the
paleobotanical Paleobotany, which is also spelled as palaeobotany, is the branch of botany dealing with the recovery and identification of plant remains from geological contexts, and their use for the biological reconstruction of past environments (paleogeogr ...
collections housed by the University of Alberta, Edmonton, Alberta, Canada. The specimen was collected in Apple Bay, near Quatsino Sound on the northwestern shore of Vancouver Island, British Columbia, Canada. The partial conk was preserved in a calcareous nodule recovered from a Greywacke sandstone matrix equivalent in age to the Longarm Formation. The nodules formed in a shallow marine environment along with abundant plant material. It was first studied by a group of researchers consisting of Selena Smith, Randolph Currah and
Ruth Stockey Ruth (or its variants) may refer to: Places France * Château de Ruthie, castle in the commune of Aussurucq in the Pyrénées-Atlantiques département of France Switzerland * Ruth, a hamlet in Cologny United States * Ruth, Alabama * Ruth, Arka ...
, all from the University of Alberta. Smith and colleagues published their 2004
type description A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a clear description of a new species of organism and explain how it differs from species that have ...
in the journal '' Mycologia'' volume 96. The generic epithet ''Quatsinoporites'' coined from a combination of ''Quatsino'' in reference to the nearness of the type locality to Quatsino Sound and "porites" to reflect that it is a polypore fungus. The specific epithet "''cranhamii''" chosen in honor of Gerald Cranham in recognition of his contribution of a number of Vancouver Island plant fossils to the University of Alberta for study. When first described ''Quatsinoporites cranhamii'' was the first fungus species to be described from Vancouver Island and the third from British Columbia. ''
Cryptodidymosphaerites princetonensis ''Cryptodidymosphaerites'' is an extinct monotypic genus of pleosporale fungus of uncertain family placement. When described it contained the single species ''Cryptodidymosphaerites princetonensis''. The genus is solely known from the Early Eoc ...
'' and ''
Palaeoserenomyces allenbyensis ''Paleoserenomyces'' is an extinct monotypic genus of pleosporale fungus of uncertain family placement. When described it contained the single species ''Paleoserenomyces allenbyensis''. The genus is solely known from the Early Eocene, Ypresi ...
'' were the first fossil fungi to be described from British Columbia, known only from the
Early Eocene In the geologic timescale the Ypresian is the oldest age or lowest stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by the Eocene Lutetian Age. The Ypresian i ...
Allenby Formation near Princeton, B.C. '' Appianoporites vancouverensis'', an Eocene age agaricomycete, was described in the same paper from deposits along Appian Way on the eastern shore of Vancouver Island. Three years later the
ascomycete Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The defi ...
'' Margaretbarromyces dictyosporus'' was also described from the Appian Way site.


Description

The holotype of ''Quatsinoporites'' is a lone fragment of fruiting body by and deep, which was abraded by water transport before preservation in a calcareous nodule. The conk section has an average of three, 130 to 540
μm The micrometre ( international spelling as used by the International Bureau of Weights and Measures; SI symbol: μm) or micrometer (American spelling), also commonly known as a micron, is a unit of length in the International System of Unit ...
diameter tubes per millimeter. The fungus is composed of monomitic hyphae. Due to the abraded nature of the specimen the basidia and basidiospores are both unknown at this time. The specimen was studied by cutting the calcareous nodule into slices with a rock saw and using the cellulose acetate peel technique to create slides that were examined under stereo microscope. ''Quatsinoporites'' is placed in Hymenochaetaceae based on the structure of the poroid hymenophore, the presence of setae and monomitic hyphal system lacking clamp connections. This placement is tentative due to limited characters available in the fossil the lack of diagnostic features such as the basidiospores.


References

{{Taxonbar, from=Q7269593 †Quatsinoporites †Quatsinoporites Prehistoric fungi Fossil taxa described in 2004 Flora of British Columbia Cretaceous fungi Monotypic Basidiomycota genera