''Propanoplosaurus'' is a genus of herbivorous

nodosaurid Nodosauridae is a family of ankylosaurian dinosaurs, from the Late Jurassic to the Late Cretaceous period in what is now North America, South America, Europe, and Asia. Description Nodosaurids, like their close relatives the ankylosaurids, we ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

from the Early Cretaceous Patuxent Formation of

Maryland Maryland ( ) is a state in the Mid-Atlantic region of the United States. It shares borders with Virginia, West Virginia, and the District of Columbia to its south and west; Pennsylvania to its north; and Delaware and the Atlantic Ocean to ...

, USA. Its

type specimen In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes th ...

is a natural cast and partial natural mold of a

hatchling In oviparous biology, a hatchling is a newly hatched fish, amphibian, reptile, or bird. A group of mammals called monotremes lay eggs, and their young are hatchlings as well. Fish Fish hatchlings generally do not receive parental care, similar to ...

Discovery and Naming

From 1994 onwards Ray Stanford uncovered an ichnofauna in

near the border with

Washington D.C. ) , image_skyline = , image_caption = Clockwise from top left: the Washington Monument and Lincoln Memorial on the National Mall, United States Capitol, Logan Circle, Jefferson Memorial, White House, Adams Morgan, Na ...

Along with dinosaur footprints the impressions of a neonate nodosaurid were found by Stanford in 1997. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...

''Propanoplosaurus marylandicus'' was named and described by Stanford,

David Weishampel Professor David Bruce Weishampel (born November 16, 1952) is an American palaeontologist in the Center for Functional Anatomy and Evolution at Johns Hopkins University School of Medicine. Weishampel received his Ph.D. in Geology from the Univers ...

and Valerie DeLeon in 2011. The generic name combines the

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

prefix ''pro~'' with the name of the genus ''

Panoplosaurus ''Panoplosaurus'' is a genus of armoured dinosaur from the Late Cretaceous of Alberta, Canada. Few specimens of the genus are known, all from the middle Campanian of the Dinosaur Park Formation, roughly 76 to 75 million years ago. It was first d ...

'' because the new species lived earlier than — but resembled much — this already described nodosaurid. The specific name refers to Maryland. The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...

, USNM 540686, was found in the Patuxent Formation, dating from the late Aptian. It consists of the impressions of the back of the head together with a natural cast of the ribcage, some vertebrae, the right forelimb, the right femur and a right foot. The animal is shown lying on its back. The authors rejected the possibility the specimen represented a

pseudofossil Pseudofossils are inorganic objects, markings, or impressions that might be mistaken for fossils. Pseudofossils may be misleading, as some types of mineral deposits can mimic lifeforms by forming what appear to be highly detailed or organized str ...

or an embryo. The specimen is the first nodosaurid skeleton from the eastern seaboard from which previously only nodosaurid teeth named as ''

Priconodon ''Priconodon'' (meaning "saw cone tooth") is an extinct genus of dinosaur (perhaps nodosaurid), known from its large teeth. Its remains have been found in the Aptian-Albian age Lower Cretaceous Arundel Formation of Muirkirk, Prince George's Count ...

'' were known, and the first neonate dinosaur from that region.

Description

The specimen has a preserved length of thirteen centimetres. The total length of the individual was estimated as between twenty-four and twenty-eight centimetres. Only the skull shows

osteoderm Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinc ...

s and the authors suggest this was a common developmental stage of all nodosaurids, together with a long middle section of the snout which is characterised by a unique cross-pattern of the bone plates, probably formed by the triangular osteoderms of the

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. T ...

Classification

''Propanoplosaurus'' was by the describers assigned to the

Nodosauridae Nodosauridae is a family of ankylosaurian dinosaurs, from the Late Jurassic to the Late Cretaceous period in what is now North America, South America, Europe, and Asia. Description Nodosaurids, like their close relatives the ankylosaurids, we ...

References

Nodosaurids Early Cretaceous dinosaurs of North America Fossil taxa described in 2011 Taxa named by David B. Weishampel Ornithischian genera {{Ankylosaurian stub

Discovery and Naming

Description

Classification

See also

References