bait fish 300px, Feeder Goldfish are common baitfish. Bait fish (or baitfish) are small-sized fish caught and used by anglers as bait to attract larger predatory fish, particularly game fish. Baitfish species are typically those that are common and bree ...

pelagic fish Pelagic fish live in the pelagic zone of ocean or lake waters—being neither close to the bottom nor near the shore—in contrast with demersal fish that do live on or near the bottom, and reef fish that are associated with coral reef ...

predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...

seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...

marine mammal Marine mammals are aquatic mammals that rely on the ocean and other marine ecosystems for their existence. They include animals such as seals, whales, manatees, sea otters and polar bears. They are an informal group, unified only by their reli ...

plankton Plankton are the diverse collection of organisms found in Hydrosphere, water (or atmosphere, air) that are unable to propel themselves against a Ocean current, current (or wind). The individual organisms constituting plankton are called plankt ...

filter feeding Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feedin ...

Clupeiformes Clupeiformes is the order of Actinopterygii, ray-finned fish that includes the herring family, Clupeidae, and the anchovy family, Engraulidae. The group includes many of the most important Forage fish, forage and food fish. Clupeiformes are phy ...

herring Herring are forage fish, mostly belonging to the family of Clupeidae. Herring often move in large schools around fishing banks and near the coast, found particularly in shallow, temperate waters of the North Pacific and North Atlantic Oceans, i ...

sardine "Sardine" and "pilchard" are common names for various species of small, oily forage fish in the herring family Clupeidae. The term "sardine" was first used in English during the early 15th century, a folk etymology says it comes from the Ital ...

shad The Alosinae, or the shads,Alosinae

,

hilsa The ilish (''Tenualosa ilisha'') ( bn, ইলিশ, translit=iliś; also known as the ilishi, hilsa, hilsa herring or hilsa shad, is a species of fish related to the herring, in the family Clupeidae. It is a very popular and sought-after food ...

,

menhaden Menhaden, also known as mossbunker and bunker and "the most important fish in the sea", are forage fish of the genera ''Brevoortia'' and ''Ethmidium'', two genera of marine fish in the family Clupeidae. ''Menhaden'' is a blend of ''poghaden'' ...

,

anchovies An anchovy is a small, common forage fish of the family Engraulidae. Most species are found in marine waters, but several will enter brackish water, and some in South America are restricted to fresh water. More than 140 species are placed in 1 ...

, and

sprat Sprat is the common name applied to a group of forage fish belonging to the genus ''Sprattus'' in the family Clupeidae. The term also is applied to a number of other small sprat-like forage fish (''Clupeoides'', ''Clupeonella'', '' Corica'', ' ...

s), but also other small fish, including

halfbeak Hemiramphidae is a family of fishes that are commonly called halfbeaks, spipe fish or spipefish. They are a geographically widespread and numerically abundant family of epipelagic fish inhabiting warm waters around the world. The halfbeaks are ...

s, silversides,

smelt Smelt may refer to: * Smelting, chemical process * The common name of various fish: ** Smelt (fish), a family of small fish, Osmeridae ** Australian smelt in the family Retropinnidae and species ''Retropinna semoni'' ** Big-scale sand smelt ''At ...

such as

capelin The capelin or caplin (''Mallotus villosus'') is a small forage fish of the smelt family found in the North Atlantic, North Pacific and Arctic oceans. In summer, it grazes on dense swarms of plankton at the edge of the ice shelf. Larger capelin ...

and goldband fusiliers. Forage fish compensate for their small size by forming

schools A school is an educational institution designed to provide learning spaces and learning environments for the teaching of students under the direction of teachers. Most countries have systems of formal education, which is sometimes compulsor ...

. Some swim in synchronised grids with their mouths open so they can efficiently filter plankton. These schools can become immense

shoals In oceanography, geomorphology, and geoscience, a shoal is a natural submerged ridge, bank, or bar that consists of, or is covered by, sand or other unconsolidated material and rises from the bed of a body of water to near the surface. It o ...

which move along coastlines and migrate across open oceans. The shoals are concentrated energy resources for the great marine predators. The predators are keenly focused on the shoals, acutely aware of their numbers and whereabouts, and make migrations themselves that can span thousands of miles to connect, or stay connected, with them.

National Coalition for Marine Conservation National Coalition for Marine Conservation is now operating as "Wild Oceans." Founded in 1973 by fishermen, the National Coalition for Marine Conservation (NCMC) is the United States's oldest public advocacy group dedicated exclusively to conserving ...

Forage fish
The ocean

primary producer Primary or primaries may refer to: Arts, entertainment, and media Music Groups and labels * Primary (band), from Australia * Primary (musician), hip hop musician and record producer from South Korea * Primary Music, Israeli record label Works ...

s, mainly contained in plankton, produce food energy from the sun and are the raw fuel for the ocean food webs. Forage fish transfer this energy by eating the plankton and becoming food themselves for the top predators. In this way, forage fish occupy the central positions in ocean and lake

food web A food web is the natural interconnection of food chains and a graphical representation of what-eats-what in an ecological community. Another name for food web is consumer-resource system. Ecologists can broadly lump all life forms into one ...

s. The

fishing industry The fishing industry includes any industry or activity concerned with taking, culturing, processing, preserving, storing, transporting, marketing or selling fish or fish products. It is defined by the Food and Agriculture Organization as including ...

sometimes catch forage fish for commercial purposes, but primarily for use as

feeder fish Feeder fish is the common name for certain types of small, inexpensive fish commonly fed as live food to other captive animals such as predatory fishes (e.g. sharks in captivity, aquarium sharks, aquaculture of salmonids, farmed salmon and tuna p ...

to farmed piscivorous animals. Some

fisheries scientists Fisheries science is the academic discipline of managing and understanding fisheries. It is a multidisciplinary science, which draws on the disciplines of limnology, oceanography, freshwater biology, marine biology, meteorology, conservation, e ...

are expressing concern that this will affect the populations of

predator fish Predatory fish are hypercarnivorous fish that actively prey upon other fish or aquatic animals, with examples including shark, billfish, barracuda, pike/muskellunge, walleye, perch and salmon. Some omnivorous fish, such as the red-bellied piranha ...

that depend on them.

In the oceans

Typical ocean forage fish are small, silvery schooling

oily fish Oily fish are fish species with oil (fats) in soft tissues and in the coelomic cavity around the gut. Their fillets may contain up to 30% oil, although this figure varies both within and between species. Examples of oily fish include small ...

such as

herring Herring are forage fish, mostly belonging to the family of Clupeidae. Herring often move in large schools around fishing banks and near the coast, found particularly in shallow, temperate waters of the North Pacific and North Atlantic Oceans, i ...

,

anchovies An anchovy is a small, common forage fish of the family Engraulidae. Most species are found in marine waters, but several will enter brackish water, and some in South America are restricted to fresh water. More than 140 species are placed in 1 ...

and

menhaden Menhaden, also known as mossbunker and bunker and "the most important fish in the sea", are forage fish of the genera ''Brevoortia'' and ''Ethmidium'', two genera of marine fish in the family Clupeidae. ''Menhaden'' is a blend of ''poghaden'' ...

, and other small, schooling

baitfish 300px, Feeder Goldfish are common baitfish. Bait fish (or baitfish) are small-sized fish caught and used by anglers as bait to attract larger predatory fish, particularly game fish. Baitfish species are typically those that are common and bree ...

like

capelin The capelin or caplin (''Mallotus villosus'') is a small forage fish of the smelt family found in the North Atlantic, North Pacific and Arctic oceans. In summer, it grazes on dense swarms of plankton at the edge of the ice shelf. Larger capelin ...

,

smelt Smelt may refer to: * Smelting, chemical process * The common name of various fish: ** Smelt (fish), a family of small fish, Osmeridae ** Australian smelt in the family Retropinnidae and species ''Retropinna semoni'' ** Big-scale sand smelt ''At ...

s,

sand lance A sand lance or sandlance is a fish belonging to the family Ammodytidae. Several species of sand lances are commonly known as "sand eels", though they are not related to true eels. Another variant name is launce, and all names of the fish are ...

,

halfbeak Hemiramphidae is a family of fishes that are commonly called halfbeaks, spipe fish or spipefish. They are a geographically widespread and numerically abundant family of epipelagic fish inhabiting warm waters around the world. The halfbeaks are ...

s,

pollock Pollock or pollack (pronounced ) is the common name used for either of the two species of North Atlantic marine fish in the genus ''Pollachius''. ''Pollachius pollachius'' is referred to as pollock in North America, Ireland and the United Kingd ...

, butterfish and juvenile

rockfish Rockfish is a common term for several species of fish, referring to their tendency to hide among rocks. The name rockfish is used for many kinds of fish used for food. This common name belongs to several groups that are not closely related, and ca ...

. Herrings are a preeminent forage fish, often marketed as

sardine "Sardine" and "pilchard" are common names for various species of small, oily forage fish in the herring family Clupeidae. The term "sardine" was first used in English during the early 15th century, a folk etymology says it comes from the Ital ...

s or

pilchard "Sardine" and "pilchard" are common names for various species of small, oily forage fish in the herring family Clupeidae. The term "sardine" was first used in English during the early 15th century, a folk etymology says it comes from the ...

s. The term “forage fish” is a term used in

fisheries Fishery can mean either the enterprise of raising or harvesting fish and other aquatic life; or more commonly, the site where such enterprise takes place ( a.k.a. fishing ground). Commercial fisheries include wild fisheries and fish farms, both ...

, and is applied also to forage species that are not true fish, but play a significant role as prey for predators. Thus invertebrates such as

squid True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fitting t ...

and

shrimp Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are refer ...

are also referred to as "forage fish". Even the tiny shrimp-like creatures called

krill Krill are small crustaceans of the order Euphausiacea, and are found in all the world's oceans. The name "krill" comes from the Norwegian Norwegian, Norwayan, or Norsk may refer to: *Something of, from, or related to Norway, a country in n ...

, small enough to be eaten by other forage fish, yet large enough to eat the same

zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...

as forage fish, are often classified as "forage fish".Marine Fish Conservation Network
Forage fish: The Most Important Fish in the Sea
Forage fish utilise the biomass of

copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...

s,

mysid Mysida is an order of small, shrimp-like crustaceans in the malacostracan superorder Peracarida. Their common name opossum shrimps stems from the presence of a brood pouch or "marsupium" in females. The fact that the larvae are reared in thi ...

s and

krill Krill are small crustaceans of the order Euphausiacea, and are found in all the world's oceans. The name "krill" comes from the Norwegian Norwegian, Norwayan, or Norsk may refer to: *Something of, from, or related to Norway, a country in n ...

in the

pelagic zone The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or wa ...

to become the dominant converters of the enormous ocean production of zooplankton. They are, in turn, central prey items for higher

trophic level The trophic level of an organism is the position it occupies in a food web. A food chain is a succession of organisms that eat other organisms and may, in turn, be eaten themselves. The trophic level of an organism is the number of steps it i ...

s. Forage fish may have achieved their dominance because of the way they live in huge, and often extremely fast cruising schools. Though forage fish are abundant, there are relatively few species. There are more species of primary producers and apex predators in the ocean than there are forage fish.

Ocean food webs

Forage fish occupy central positions in the ocean

food web A food web is the natural interconnection of food chains and a graphical representation of what-eats-what in an ecological community. Another name for food web is consumer-resource system. Ecologists can broadly lump all life forms into one ...

s. The position that a fish occupies in a food web is called its

trophic level The trophic level of an organism is the position it occupies in a food web. A food chain is a succession of organisms that eat other organisms and may, in turn, be eaten themselves. The trophic level of an organism is the number of steps it i ...

(

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

''trophē'' = food). The organisms it eats are at a lower trophic level, and the organisms that eat it are at a higher trophic level. Forage fish occupy middle levels in the food web, serving as a dominant prey to higher level fish, seabirds and mammals.

Ecological pyramid An ecological pyramid (also trophic pyramid, Eltonian pyramid, energy pyramid, or sometimes food pyramid) is a graphical representation designed to show the Biomass (ecology), biomass or Primary production, bioproductivity at each trophic level ...

s are graphical representations, along the lines of the diagram at the right, which show how

biomass Biomass is plant-based material used as a fuel for heat or electricity production. It can be in the form of wood, wood residues, energy crops, agricultural residues, and waste from industry, farms, and households. Some people use the terms bi ...

or

productivity Productivity is the efficiency of production of goods or services expressed by some measure. Measurements of productivity are often expressed as a ratio of an aggregate output to a single input or an aggregate input used in a production proces ...

changes at each trophic level in an

ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syste ...

. The first or bottom level is occupied by

primary producers An autotroph or primary producer is an organism that produces complex organic compounds (such as carbohydrates, fats, and proteins) using carbon from simple substances such as carbon dioxide,Morris, J. et al. (2019). "Biology: How Life Work ...

or

autotroph An autotroph or primary producer is an organism that produces complex organic compounds (such as carbohydrates, fats, and proteins) using carbon from simple substances such as carbon dioxide,Morris, J. et al. (2019). "Biology: How Life Works", ...

s (

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

''autos'' = self and ''trophe'' = food). These are the names given to organisms that do not feed on other organisms, but produce biomass from inorganic compounds, mostly by a

process A process is a series or set of activities that interact to produce a result; it may occur once-only or be recurrent or periodic. Things called a process include: Business and management *Business process, activities that produce a specific se ...

of

photosynthesis Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored i ...

. In oceans, most primary production is performed by

algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...

. This is a contrast to land, where most primary production is performed by

vascular plants Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They al ...

. Algae ranges from single floating cells to attached

seaweeds Seaweed, or macroalgae, refers to thousands of species of macroscopic, multicellular, marine algae. The term includes some types of '' Rhodophyta'' (red), ''Phaeophyta'' (brown) and ''Chlorophyta'' (green) macroalgae. Seaweed species such as ...

, while vascular plants are represented in the ocean by groups such as the

seagrass Seagrasses are the only flowering plants which grow in marine environments. There are about 60 species of fully marine seagrasses which belong to four families (Posidoniaceae, Zosteraceae, Hydrocharitaceae and Cymodoceaceae), all in the orde ...

es. Larger producers, such as seagrasses and seaweeds, are mostly confined to the

littoral The littoral zone or nearshore is the part of a sea, lake, or river that is close to the shore. In coastal ecology, the littoral zone includes the intertidal zone extending from the high water mark (which is rarely inundated), to coastal areas ...

zone and shallow waters, where they attach to the underlying substrate and are still within the

photic zone The photic zone, euphotic zone, epipelagic zone, or sunlight zone is the uppermost layer of a body of water that receives sunlight, allowing phytoplankton to perform photosynthesis. It undergoes a series of physical, chemical, and biological proc ...

. Most primary production in the ocean is performed by

microscopic organisms A microorganism, or microbe,, ''mikros'', "small") and ''organism'' from the el, ὀργανισμός, ''organismós'', "organism"). It is usually written as a single word but is sometimes hyphenated (''micro-organism''), especially in olde ...

, the phytoplankton. Thus, in ocean environments, the first bottom trophic level is occupied principally by

phytoplankton Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater ecosystems. The name comes from the Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'. Ph ...

, microscopic drifting organisms, mostly

one-celled A unicellular organism, also known as a single-celled organism, is an organism that consists of a single cell, unlike a multicellular organism that consists of multiple cells. Organisms fall into two general categories: prokaryotic organisms and ...

algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...

, that float in the sea. Most phytoplankton are too small to be seen individually with the

unaided eye Naked eye, also called bare eye or unaided eye, is the practice of engaging in visual perception unaided by a magnifying, light-collecting optical instrument, such as a telescope or microscope, or eye protection. Vision corrected to normal ...

. They can appear as a green discoloration of the water when they are present in high enough numbers. Since they increase their biomass mostly through photosynthesis they live in the sun-lit surface layer (

euphotic zone The photic zone, euphotic zone, epipelagic zone, or sunlight zone is the uppermost layer of a body of water that receives sunlight, allowing phytoplankton to perform photosynthesis. It undergoes a series of physical, chemical, and biological proc ...

) of the sea. The most important groups of phytoplankton include the

diatom A diatom (Neo-Latin ''diatoma''), "a cutting through, a severance", from el, διάτομος, diátomos, "cut in half, divided equally" from el, διατέμνω, diatémno, "to cut in twain". is any member of a large group comprising sev ...

s and

dinoflagellate The dinoflagellates (Greek δῖνος ''dinos'' "whirling" and Latin ''flagellum'' "whip, scourge") are a monophyletic group of single-celled eukaryotes constituting the phylum Dinoflagellata and are usually considered algae. Dinoflagellates are ...

s. Diatoms are especially important in oceans, where they are estimated to contribute up to 45% of the total ocean's primary production. Diatoms are usually

microscopic The microscopic scale () is the scale of objects and events smaller than those that can easily be seen by the naked eye, requiring a lens (optics), lens or microscope to see them clearly. In physics, the microscopic scale is sometimes regarded a ...

, although some species can reach up to in length. The second trophic level (

primary consumer A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...

s) is occupied by

zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...

which feed off the

phytoplankton Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater ecosystems. The name comes from the Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'. Ph ...

. Together with the phytoplankton, they form the base of the food pyramid that supports most of the world's great fishing grounds. Zooplankton are tiny animals found with the phytoplankton in

ocean The ocean (also the sea or the world ocean) is the body of salt water that covers approximately 70.8% of the surface of Earth and contains 97% of Earth's water. An ocean can also refer to any of the large bodies of water into which the wo ...

ic surface waters, and include tiny

crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

s, and fish larvae and fry (recently hatched fish). Most zooplankton are

filter feeder Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feedin ...

s, and they use appendages to strain the phytoplankton in the water. Some larger zooplankton also feed on smaller zooplankton. Some zooplankton can jump about a bit to avoid predators, but they can't really swim. Like phytoplankton, they float with the currents, tides and winds instead. Zooplanktons can reproduce rapidly, their populations can increase up to thirty percent a day under favourable conditions. Many live short and productive lives and reach maturity quickly. Particularly important groups of zooplankton are the

copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...

s and

krill Krill are small crustaceans of the order Euphausiacea, and are found in all the world's oceans. The name "krill" comes from the Norwegian Norwegian, Norwayan, or Norsk may refer to: *Something of, from, or related to Norway, a country in n ...

. These are not shown in the images above, but are discussed in more detail later.

Copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...

s are a group of small

crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

s found in ocean and

freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...

habitats In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

. They are the biggest source of protein in the sea, and are important prey for forage fish.

Krill Krill are small crustaceans of the order Euphausiacea, and are found in all the world's oceans. The name "krill" comes from the Norwegian Norwegian, Norwayan, or Norsk may refer to: *Something of, from, or related to Norway, a country in n ...

constitute the next biggest source of protein. Krill are particularly large predator zooplankton which feed on smaller zooplankton. This means they really belong to the third trophic level, secondary consumers, along with the forage fish. Together,

phytoplankton Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater ecosystems. The name comes from the Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'. Ph ...

and

zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...

make up most of the

plankton Plankton are the diverse collection of organisms found in Hydrosphere, water (or atmosphere, air) that are unable to propel themselves against a Ocean current, current (or wind). The individual organisms constituting plankton are called plankt ...

in the sea. Plankton is the term applied to any small drifting

organism In biology, an organism () is any living system that functions as an individual entity. All organisms are composed of cells (cell theory). Organisms are classified by taxonomy into groups such as multicellular animals, plants, and ...

s that float in the sea (

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

= wanderer or drifter). By definition, organisms classified as plankton are unable to swim against ocean currents; they cannot resist the ambient current and control their position. In ocean environments, the first two trophic levels are occupied mainly by

plankton Plankton are the diverse collection of organisms found in Hydrosphere, water (or atmosphere, air) that are unable to propel themselves against a Ocean current, current (or wind). The individual organisms constituting plankton are called plankt ...

. Plankton are divided into producers and consumers. The producers are the phytoplankton (Greek ''phyton'' = plant) and the consumers, who eat the phytoplankton, are the zooplankton (Greek ''zoon'' = animal).

Diet

Forage fish feed on

plankton Plankton are the diverse collection of organisms found in Hydrosphere, water (or atmosphere, air) that are unable to propel themselves against a Ocean current, current (or wind). The individual organisms constituting plankton are called plankt ...

. When they are eaten by larger predators, they transfer this energy from the bottom of the food chain to the top and in this way are the central link between

trophic level The trophic level of an organism is the position it occupies in a food web. A food chain is a succession of organisms that eat other organisms and may, in turn, be eaten themselves. The trophic level of an organism is the number of steps it i ...

s. Forage fish are usually

filter feeder Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feedin ...

s, meaning that they feed by straining suspended matter and food particles from water. They usually travel in large, slow moving, tightly packed

schools A school is an educational institution designed to provide learning spaces and learning environments for the teaching of students under the direction of teachers. Most countries have systems of formal education, which is sometimes compulsor ...

with their mouths open. They are typically

omnivorous An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutri ...

. Their diet is usually based primarily on zooplankton, although, since they are omnivorous, they also take in some phytoplankton. Young forage fish, such as herring, mostly feed on

phytoplankton Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater ecosystems. The name comes from the Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'. Ph ...

and as they mature they start to consume larger organisms. Older herrings feed on

zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...

, tiny animals that are found in

ocean The ocean (also the sea or the world ocean) is the body of salt water that covers approximately 70.8% of the surface of Earth and contains 97% of Earth's water. An ocean can also refer to any of the large bodies of water into which the wo ...

ic surface waters, and

fish larvae Ichthyoplankton (from Greek: ἰχθύς, , "fish"; and πλαγκτός, , "drifter") are the eggs and larvae of fish. They are mostly found in the sunlit zone of the water column, less than 200 metres deep, which is sometimes called the epip ...

and fry (recently hatched fish).

Copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...

s and other tiny

crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

s are common zooplankton eaten by forage fish. During

daylight Daylight is the combination of all direct and indirect sunlight during the daytime. This includes direct sunlight, diffuse sky radiation, and (often) both of these reflected by Earth and terrestrial objects, like landforms and buildings. Sunligh ...

, many forage fish stay in the safety of deep water, feeding at the surface only at night when there is less chance of predation. They swim with their mouths open, filtering plankton from the water as it passes through their gills. Ocean

halfbeak Hemiramphidae is a family of fishes that are commonly called halfbeaks, spipe fish or spipefish. They are a geographically widespread and numerically abundant family of epipelagic fish inhabiting warm waters around the world. The halfbeaks are ...

s are

omnivore An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutr ...

s which feed on

algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...

, plankton, marine plants like

seagrass Seagrasses are the only flowering plants which grow in marine environments. There are about 60 species of fully marine seagrasses which belong to four families (Posidoniaceae, Zosteraceae, Hydrocharitaceae and Cymodoceaceae), all in the orde ...

, invertebrates like

pteropods Pteropoda ( common name pteropods, from the Greek meaning "wing-foot") are specialized free-swimming pelagic sea snails and sea slugs, marine opisthobranch gastropods. Most live in the top 10 m of the ocean and are less than 1 cm long. The mo ...

and

crustaceans Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

and smaller fishes. Some tropical species feed on animals during the day and plants at night, while others alternate summer carnivory with winter herbivory. They are in turn eaten by

billfish The term billfish refers to a group of saltwater predatory fish characterised by prominent pointed bills (rostra), and by their large size; some are longer than . Extant billfish include sailfish and marlin, which make up the family Istiophor ...

,

mackerel Mackerel is a common name applied to a number of different species of pelagic fish, mostly from the family Scombridae. They are found in both temperate and tropical seas, mostly living along the coast or offshore in the oceanic environment. ...

, and

shark Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachimo ...

s.

Predators

Forage fish are the food that sustains larger predators above them in the ocean food chain. The superabundance they present in their schools make them ideal food sources for top predator fish such as

tuna A tuna is a saltwater fish that belongs to the tribe Thunnini, a subgrouping of the Scombridae (mackerel) family. The Thunnini comprise 15 species across five genera, the sizes of which vary greatly, ranging from the bullet tuna (max length: ...

,

striped bass The striped bass (''Morone saxatilis''), also called the Atlantic striped bass, striper, linesider, rock, or rockfish, is an anadromous perciform fish of the family Moronidae found primarily along the Atlantic coast of North America. It has al ...

,

cod Cod is the common name for the demersal fish genus '' Gadus'', belonging to the family Gadidae. Cod is also used as part of the common name for a number of other fish species, and one species that belongs to genus ''Gadus'' is commonly not call ...

,

salmon Salmon () is the common name for several list of commercially important fish species, commercially important species of euryhaline ray-finned fish from the family (biology), family Salmonidae, which are native to tributary, tributaries of the ...

,

barracuda A barracuda, or cuda for short, is a large, predatory, ray-finned fish known for its fearsome appearance and ferocious behaviour. The barracuda is a saltwater fish of the genus ''Sphyraena'', the only genus in the family Sphyraenidae, which was ...

and

swordfish Swordfish (''Xiphias gladius''), also known as broadbills in some countries, are large, highly migratory predatory fish characterized by a long, flat, pointed bill. They are a popular sport fish of the billfish category, though elusive. Swordfis ...

, as well as

shark Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachimo ...

s,

whale Whales are a widely distributed and diverse group of fully aquatic placental marine mammals. As an informal and colloquial grouping, they correspond to large members of the infraorder Cetacea, i.e. all cetaceans apart from dolphins and ...

s,

dolphin A dolphin is an aquatic mammal within the infraorder Cetacea. Dolphin species belong to the families Delphinidae (the oceanic dolphins), Platanistidae (the Indian river dolphins), Iniidae (the New World river dolphins), Pontoporiidae (the ...

s,

porpoise Porpoises are a group of fully aquatic marine mammals, all of which are classified under the family Phocoenidae, parvorder Odontoceti (toothed whales). Although similar in appearance to dolphins, they are more closely related to narwhals an ...

s,

seal Seal may refer to any of the following: Common uses * Pinniped, a diverse group of semi-aquatic marine mammals, many of which are commonly called seals, particularly: ** Earless seal, or "true seal" ** Fur seal * Seal (emblem), a device to impr ...

s,

sea lion Sea lions are pinnipeds characterized by external ear flaps, long foreflippers, the ability to walk on all fours, short and thick hair, and a big chest and belly. Together with the fur seals, they make up the family Otariidae, eared seals. ...

s, and

seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...

s.

Schooling

Forage fish compensate for their small size by forming

schools A school is an educational institution designed to provide learning spaces and learning environments for the teaching of students under the direction of teachers. Most countries have systems of formal education, which is sometimes compulsor ...

. These sometimes immense gatherings fuel the ocean food web. Most forage fish are

pelagic fish Pelagic fish live in the pelagic zone of ocean or lake waters—being neither close to the bottom nor near the shore—in contrast with demersal fish that do live on or near the bottom, and reef fish that are associated with coral reef ...

, which means they form their schools in open water, and not on the bottom (

benthic fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They o ...

) or near the bottom (

benthopelagic fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They oc ...

). They are short-lived, and go mostly unnoticed by humans, apart from an occasional support role in a documentary about a great ocean predator. While we may not pay them much attention, the great marine predators are keenly focused on them, acutely aware of their numbers and whereabouts, and make migrations that can span thousands of miles to connect with them. After all, forage fish are their food.

Herring Herring are forage fish, mostly belonging to the family of Clupeidae. Herring often move in large schools around fishing banks and near the coast, found particularly in shallow, temperate waters of the North Pacific and North Atlantic Oceans, i ...

are among the most spectacular schooling fish. They aggregate together in huge numbers. Schools have been measured at over four cubic kilometres in size, containing about four billion fish. These schools move along coastlines and traverse the open oceans. Herring schools in general have very precise arrangements which allow the school to maintain relatively constant cruising speeds. Herrings have excellent hearing, and their schools react very fast to a predator. The herrings keep a certain distance from a moving scuba diver or cruising predator like a killer whale, forming a vacuole which can look like a doughnut from a spotter plane. The intricacies of schooling is far from fully understood, especially the swimming and feeding energetics. Many hypotheses to explain the function of schooling have been suggested, such as better orientation, synchronized hunting, predator confusion and reduced risk of being found. Schooling also has disadvantages, such as excretion buildup in the breathing media and oxygen and food depletion. The way the fish array in the school probably gives energy saving advantages, though this is controversial. On calm days, schools of herring can be detected at the surface a mile away by little waves they form, or from several meters at night when they trigger

bioluminescence Bioluminescence is the production and emission of light by living organisms. It is a form of chemiluminescence. Bioluminescence occurs widely in marine vertebrates and invertebrates, as well as in some fungi, microorganisms including some b ...

in surrounding

plankton Plankton are the diverse collection of organisms found in Hydrosphere, water (or atmosphere, air) that are unable to propel themselves against a Ocean current, current (or wind). The individual organisms constituting plankton are called plankt ...

. Underwater recordings show herring constantly cruising at high speeds up to 108 cm per second, with much higher escape speeds. They are fragile fish, and because of their adaptation to schooling behaviour they are rarely displayed in aquaria. Even with the best facilities aquaria can offer they become sluggish compared to their quivering energy in wild schools. Copepodkils

Juvenile Clupea harengus feeding on copepods macrophotography video

Hunting copepods

Copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...

s are a group of small

crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

s found in ocean and

freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...

habitats In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

. Many species are

plankton Plankton are the diverse collection of organisms found in Hydrosphere, water (or atmosphere, air) that are unable to propel themselves against a Ocean current, current (or wind). The individual organisms constituting plankton are called plankt ...

ic (drifting in the ocean water), while others are

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...

(living on the sea floor). Copepods are typically one millimetre (0.04 in) to two millimetres (0.08 in) long, with a teardrop shaped body. Like other

crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

s they have an armoured

exoskeleton An exoskeleton (from Greek ''éxō'' "outer" and ''skeletós'' "skeleton") is an external skeleton that supports and protects an animal's body, in contrast to an internal skeleton (endoskeleton) in for example, a human. In usage, some of the ...

, but they are so small that this armour, and the entire body, is usually transparent. Copepods are usually the dominant

zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...

. Some scientists say they form the largest animal

biomass Biomass is plant-based material used as a fuel for heat or electricity production. It can be in the form of wood, wood residues, energy crops, agricultural residues, and waste from industry, farms, and households. Some people use the terms bi ...

on the planet. The other contender is the

Antarctic krill Antarctic krill (''Euphausia superba'') is a species of krill found in the Antarctic waters of the Southern Ocean. It is a small, swimming crustacean that lives in large schools, called swarms, sometimes reaching densities of 10,000–30,000 ind ...

. But copepods are smaller than krill, with faster growth rates, and they are more evenly distributed throughout the oceans. This means copepods almost certainly contribute more secondary production to the world's oceans than krill, and perhaps more than all other groups of marine organisms together. They are a major item on the forage fish menu. Copepods are very alert and evasive. They have large antennae. When they spread their antennae they can sense the pressure wave from an approaching fish and jump with great speed over a few centimeters. Herrings are pelagic feeders. Their prey consists of a wide spectrum of

phytoplankton Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater ecosystems. The name comes from the Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'. Ph ...

and

zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...

, amongst which copepods are the dominant prey. Young

herring Herring are forage fish, mostly belonging to the family of Clupeidae. Herring often move in large schools around fishing banks and near the coast, found particularly in shallow, temperate waters of the North Pacific and North Atlantic Oceans, i ...

usually capture small copepods by hunting them individually— they approach them from below. The (half speed) video loop at the left shows a juvenile herring feeding on

copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...

s. In the middle of the image a copepod escapes successfully to the left. The opercula (hard bony flaps covering the

gill A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are ...

s) are spread wide open to compensate the pressure wave which would alert the copepod to trigger a jump.Kils, U (1992) ''The ecoSCOPE and dynIMAGE: Microscale tools for in situ studies of predator–prey interactions''. Arch Hydrobiol Beih 36: 83–96 Herringramkils

If prey concentrations reach very high levels, the herrings adopt a method called "ram feeding". They swim with their mouth wide open and their opercula fully expanded. Every several feet, they close and clean their

gill raker Gill rakers in fish are bony or cartilaginous processes that project from the branchial arch (gill arch) and are involved with suspension feeding tiny prey. They are not to be confused with the gill filaments that compose the fleshy part of the ...

s for a few milliseconds (

filter feeding Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feedin ...

). In the photo on the right, herring ram feed on a school of

copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...

s. The fish all open their mouths and opercula wide at the same time (the red gills are visible—click to enlarge). The fish swim in a grid where the distance between them is the same as the jump length of their prey, as indicated in the animation below. In the animation, juvenile herring hunt the

copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...

s in synchronization: The copepods sense with their antennae the pressure-wave of an approaching herring and react with a fast escape jump. The length of the jump is fairly constant. The fish align themselves in a grid with this characteristic jump length. A copepod can dart about 80 times before it tires out. After a jump, it takes it 60 milliseconds to spread its antennae again, and this time delay becomes its undoing, as the almost endless stream of herrings allows a herring to eventually snap the copepod. A single juvenile herring could never catch a large copepod.

Migrations

Forage fish often make great migrations between their spawning, feeding and nursery grounds. Schools of a particular stock usually travel in a triangle between these grounds. For example, one stock of herrings have their spawning ground in southern

Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and t ...

, their feeding ground in

Iceland Iceland ( is, Ísland; ) is a Nordic island country in the North Atlantic Ocean and in the Arctic Ocean. Iceland is the most sparsely populated country in Europe. Iceland's capital and largest city is Reykjavík, which (along with its s ...

, and their nursery ground in northern Norway. Wide triangular journeys such as these may be important because forage fish, when feeding, cannot distinguish their own offspring. Fertile feeding grounds for forage fish are provided by ocean upwellings. Oceanic gyres are large-scale

ocean currents An ocean current is a continuous, directed movement of sea water generated by a number of forces acting upon the water, including wind, the Coriolis effect, breaking waves, cabbeling, and temperature and salinity differences. Depth contours, s ...

caused by the

Coriolis effect In physics, the Coriolis force is an inertial or fictitious force that acts on objects in motion within a frame of reference that rotates with respect to an inertial frame. In a reference frame with clockwise rotation, the force acts to the ...

. Wind-driven surface currents interact with these gyres and the underwater topography, such as

seamount A seamount is a large geologic landform that rises from the ocean floor that does not reach to the water's surface (sea level), and thus is not an island, islet or cliff-rock. Seamounts are typically formed from extinct volcanoes that rise abru ...

s and the edge of

continental shelves A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...

, to produce downwellings and

upwelling Upwelling is an oceanographic phenomenon that involves wind-driven motion of dense, cooler, and usually nutrient-rich water from deep water towards the ocean surface. It replaces the warmer and usually nutrient-depleted surface water. The nutr ...

s. These can transport nutrients which plankton thrive on. The result can be rich feeding grounds attractive to the plankton feeding forage fish. In turn, the forage fish themselves become a feeding ground for larger predator fish. Most upwellings are coastal, and many of them support some of the most productive fisheries in the world. Regions of notable upwelling include coastal

Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...

,

Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...

,

Arabian Sea The Arabian Sea ( ar, اَلْبَحرْ ٱلْعَرَبِيُّ, Al-Bahr al-ˁArabī) is a region of the northern Indian Ocean bounded on the north by Pakistan, Iran and the Gulf of Oman, on the west by the Gulf of Aden, Guardafui Channel ...

, western

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

, eastern

New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...

and the

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

coast.

Capelin The capelin or caplin (''Mallotus villosus'') is a small forage fish of the smelt family found in the North Atlantic, North Pacific and Arctic oceans. In summer, it grazes on dense swarms of plankton at the edge of the ice shelf. Larger capelin ...

are a forage fish of the

smelt Smelt may refer to: * Smelting, chemical process * The common name of various fish: ** Smelt (fish), a family of small fish, Osmeridae ** Australian smelt in the family Retropinnidae and species ''Retropinna semoni'' ** Big-scale sand smelt ''At ...

family found in the

Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...

and

Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...

oceans. In summer, they graze on dense swarms of

plankton Plankton are the diverse collection of organisms found in Hydrosphere, water (or atmosphere, air) that are unable to propel themselves against a Ocean current, current (or wind). The individual organisms constituting plankton are called plankt ...

at the edge of the ice shelf. Larger capelin also eat

krill Krill are small crustaceans of the order Euphausiacea, and are found in all the world's oceans. The name "krill" comes from the Norwegian Norwegian, Norwayan, or Norsk may refer to: *Something of, from, or related to Norway, a country in n ...

and other

crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

s. The capelin move inshore in large schools to spawn and migrate in spring and summer to feed in plankton rich areas between

Iceland Iceland ( is, Ísland; ) is a Nordic island country in the North Atlantic Ocean and in the Arctic Ocean. Iceland is the most sparsely populated country in Europe. Iceland's capital and largest city is Reykjavík, which (along with its s ...

,

Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is t ...

, and

Jan Mayen Jan Mayen () is a Norwegian volcanic island in the Arctic Ocean with no permanent population. It is long (southwest-northeast) and in area, partly covered by glaciers (an area of around the Beerenberg volcano). It has two parts: larger nort ...

. The migration is affected by

ocean current An ocean current is a continuous, directed movement of sea water generated by a number of forces acting upon the water, including wind, the Coriolis effect, breaking waves, cabbeling, and temperature and salinity differences. Depth contours, s ...

s. Around Iceland maturing capelin make large northward feeding migrations in spring and summer. The return migration takes place in September to November. The spawning migration starts north of Iceland in December or January. The diagram on the right shows the main

spawning Spawn is the eggs and sperm released or deposited into water by aquatic animals. As a verb, ''to spawn'' refers to the process of releasing the eggs and sperm, and the act of both sexes is called spawning. Most aquatic animals, except for aquati ...

grounds and larval drift routes. Capelin on the way to feeding grounds is coloured green, capelin on the way back is blue, and the breeding grounds are red. In a paper published in 2009, researchers from Iceland recount their application of an interacting particle model to the capelin stock around Iceland, successfully predicting the spawning migration route for 2008.

Predator attacks

Schooling forage fish are subject to constant attacks by predators. An example is the attacks that take place during the African

sardine run The KwaZulu-Natal sardine run of southern Africa occurs from May through July when billions of sardines – or more specifically the Southern African pilchard ''Sardinops sagax'' – spawn in the cool waters of the Agulhas Bank and move northwar ...

. The African sardine run is a spectacular migration by millions of silvery sardines along the southern

coastline The coast, also known as the coastline or seashore, is defined as the area where land meets the ocean, or as a line that forms the boundary between the land and the coastline. The Earth has around of coastline. Coasts are important zones in n ...

of Africa. In terms of biomass, the sardine run could rival East Africa's great wildebeest migration. Morus serrator

Sardines have a short life-cycle, living only two or three years. Adult sardines, about two years old, mass on the

Agulhas Bank The Agulhas Bank (, from Portuguese for Cape Agulhas, ''Cabo das Agulhas'', "Cape of Needles") is a broad, shallow part of the southern African continental shelf which extends up to south of Cape Agulhas before falling steeply to the abyssal pl ...

where they spawn during spring and summer, releasing tens of thousands of eggs into the water. The adult sardines then make their way in hundreds of shoals towards the sub-tropical waters of the

Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by th ...

. A larger shoal might be long, wide and deep. Huge numbers of sharks, dolphins, tuna, sailfish,

Cape fur seal The brown fur seal (''Arctocephalus pusillus''), also known as the Cape fur seal, South African fur seal and Australian fur seal, is a species of fur seal. Description The brown fur seal is the largest and most robust member of the fur seals. ...

s and even killer whales congregate and follow the shoals, creating a

feeding frenzy In ecology, a feeding frenzy occurs when predators are overwhelmed by the amount of prey available. The term is also used as an idiom in the English language. Examples in nature For example, a large school of fish can cause nearby sharks, such a ...

along the coastline. When threatened, sardines instinctively group together and create massive

bait ball A bait ball, or baitball, occurs when small fish swarm in a tightly packed spherical formation about a common centre. It is a last-ditch defensive measure adopted by small schooling fish when they are threatened by predators. Small schooling f ...

s. Bait balls can be up to in diameter. They are short lived, seldom lasting longer than 20 minutes. As many as 18,000 dolphins, behaving like sheepdogs, round the sardines into these bait balls, or herd them to shallow water (corralling) where they are easier to catch. Once rounded up, the dolphins and other predators take turns plowing through the bait balls, gorging on the fish as they sweep through. Seabirds also attack them from above, flocks of

gannet Gannets are seabirds comprising the genus ''Morus'' in the family Sulidae, closely related to boobies. Gannets are large white birds with yellowish heads; black-tipped wings; and long bills. Northern gannets are the largest seabirds in the Nor ...

s,

cormorant Phalacrocoracidae is a family of approximately 40 species of aquatic birds commonly known as cormorants and shags. Several different classifications of the family have been proposed, but in 2021 the IOC adopted a consensus taxonomy of seven ge ...

s,

tern Terns are seabirds in the family Laridae that have a worldwide distribution and are normally found near the sea, rivers, or wetlands. Terns are treated as a subgroup of the family Laridae which includes gulls and skimmers and consists of e ...

s and

gull Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century, m ...

s. Some of these seabirds plummet from heights of , plunging through the water leaving vapour-like trails behind like fighter planes. The eggs, left behind at the Agulhas Banks, drift northwest with the current into waters off the west coast, where the larvae develop into juvenile fish. When they are old enough, they aggregate into dense shoals and migrate southwards, returning to the Agulhas banks in order to restart the cycle.

Forage fisheries

History

Herring has been known as a

staple food A staple food, food staple, or simply a staple, is a food that is eaten often and in such quantities that it constitutes a dominant portion of a standard diet for a given person or group of people, supplying a large fraction of energy needs and ...

source since 3000 B.C. In

Roman Roman or Romans most often refers to: *Rome, the capital city of Italy *Ancient Rome, Roman civilization from 8th century BC to 5th century AD *Roman people, the people of ancient Rome *''Epistle to the Romans'', shortened to ''Romans'', a letter ...

times, anchovies were the base for the fermented fish sauce called ''

garum Garum is a fermented fish sauce that was used as a condiment in the cuisines of Phoenicia, ancient Greece, Rome, Carthage and later Byzantium. Liquamen is a similar preparation, and at times they were synonymous. Although garum enjoyed its grea ...

.'' This staple of cuisine was produced in industrial quantities and transported over long distances. Fishing for sardela or sardina (''Sardina pilchardus'') is an ongoing activity on the

Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = "Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capit ...

n

Adriatic The Adriatic Sea () is a body of water separating the Italian Peninsula from the Balkans, Balkan Peninsula. The Adriatic is the northernmost arm of the Mediterranean Sea, extending from the Strait of Otranto (where it connects to the Ionian Sea) ...

coasts of

Dalmatia Dalmatia (; hr, Dalmacija ; it, Dalmazia; see #Name, names in other languages) is one of the four historical region, historical regions of Croatia, alongside Croatia proper, Slavonia, and Istria. Dalmatia is a narrow belt of the east shore of ...

and

Istria Istria ( ; Croatian language, Croatian and Slovene language, Slovene: ; ist, Eîstria; Istro-Romanian language, Istro-Romanian, Italian language, Italian and Venetian language, Venetian: ; formerly in Latin and in Ancient Greek) is the larges ...

. It traces its roots back thousands of years. The region was then largely a Venetian dominion, part of the

Roman Empire The Roman Empire ( la, Imperium Romanum ; grc-gre, Βασιλεία τῶν Ῥωμαίων, Basileía tôn Rhōmaíōn) was the post-Republican period of ancient Rome. As a polity, it included large territorial holdings around the Mediterr ...

. The area has always been sustained through fishing mainly sardines. Along the coast towns still promote the traditional practice of fishing by

lateen A lateen (from French ''latine'', meaning "Latin") or latin-rig is a triangular sail set on a long yard mounted at an angle on the mast, and running in a fore-and-aft direction. The settee can be considered to be an associated type of the same o ...

sail boats for tourism and festivals.

Pilchard "Sardine" and "pilchard" are common names for various species of small, oily forage fish in the herring family Clupeidae. The term "sardine" was first used in English during the early 15th century, a folk etymology says it comes from the ...

fishing and processing thrived in

Cornwall Cornwall (; kw, Kernow ) is a historic county and ceremonial county in South West England. It is recognised as one of the Celtic nations, and is the homeland of the Cornish people. Cornwall is bordered to the north and west by the Atlantic ...

between 1750 and 1880, after which stocks went into an almost terminal decline. Recently (2007) stocks have been improving. The industry has featured in many works of art, including Stanhope Forbes and other

Newlyn School The Newlyn School was an art colony of artists based in or near Newlyn, a fishing village adjacent to Penzance, on the south coast of Cornwall, from the 1880s until the early twentieth century. The establishment of the Newlyn School was reminisc ...

artists.

Contemporary

Traditional

commercial fisheries Commercial fishing is the activity of catching fish and other seafood for commercial profit, mostly from wild fisheries. It provides a large quantity of food to many countries around the world, but those who practice it as an industry must often ...

were directed towards high value ocean predators such as

cod Cod is the common name for the demersal fish genus '' Gadus'', belonging to the family Gadidae. Cod is also used as part of the common name for a number of other fish species, and one species that belongs to genus ''Gadus'' is commonly not call ...

,

rockfish Rockfish is a common term for several species of fish, referring to their tendency to hide among rocks. The name rockfish is used for many kinds of fish used for food. This common name belongs to several groups that are not closely related, and ca ...

and

tuna A tuna is a saltwater fish that belongs to the tribe Thunnini, a subgrouping of the Scombridae (mackerel) family. The Thunnini comprise 15 species across five genera, the sizes of which vary greatly, ranging from the bullet tuna (max length: ...

, rather than forage fish. As technologies developed, fisheries became so effective at locating and catching predator fish that many of the stocks collapsed. The industry compensated by turning to species lower in the food chain. In former times, forage fish were more difficult to fish profitably, and were a small part of the global marine fisheries. But modern industrial fishing technologies have enabled the removal of increasing quantities. Industrial-scale forage fish fisheries need large scale landings of fish to return profits. They are dominated by a small number of corporate fishing and processing companies. Forage fish populations are very vulnerable when faced with modern fishing equipment. They swim near the surface in compacted schools, so they are relatively easy to locate at the surface with sophisticated electronic

fishfinder A fishfinder or sounder (Australia) is an instrument used to locate fish underwater by detecting reflected pulses of sound energy, as in sonar. A modern fishfinder displays measurements of reflected sound on a graphical display, allowing an oper ...

s and from above with

spotter plane A surveillance aircraft is an aircraft used for surveillance. They are operated by military forces and other government agencies in roles such as intelligence gathering, battlefield surveillance, airspace surveillance, reconnaissance, observat ...

s. Once located, they are scooped out of the water using highly efficient nets, such as

purse seine Seine fishing (or seine-haul fishing; ) is a method of fishing that employs a surrounding net, called a seine, that hangs vertically in the water with its bottom edge held down by weights and its top edge buoyed by floats. Seine nets can be dep ...

s, which remove most of the school. Spawning patterns in forage fish are highly predictable. Some fisheries use knowledge of these patterns to harvest the forage species as they come together to spawn, removing the fish before they have actually spawned. Fishing during spawning periods or at other times when forage fish amass in large numbers can also be a blow to predators. Many predators, such as whales, tuna and sharks, have evolved to migrate long distances to specific sites for feeding and breeding. Their survival hinges on their finding these forage schools at their feeding grounds. The great ocean predators find that, no matter how they are adapted for speed, size, endurance or stealth, they are on the losing side when faced with the machinery of contemporary industrial fishing. Altogether, forage fish account for 37 percent (31.5 million tonnes) of all fish taken from the world's oceans each year. However, because there are fewer species of forage fish compared to predator fish, forage species fisheries are the largest in the world. Seven of the top ten fisheries target forage fish. The total world catch of herrings, sardines and anchovies alone in 2005 was 22.4 million tonnes, 24 percent of the total world catch. The Peruvian

anchoveta The Peruvian anchoveta (''Engraulis ringens'') is a species of fish of the anchovy family, Engraulidae, from the Southeast Pacific Ocean. It has yielded greater catches than any other single wild fish species in the world, with annual harvests ...

fishery is now the biggest in the world (10.7 million tonnes in 2004), while the Alaskan

pollock Pollock or pollack (pronounced ) is the common name used for either of the two species of North Atlantic marine fish in the genus ''Pollachius''. ''Pollachius pollachius'' is referred to as pollock in North America, Ireland and the United Kingd ...

fishery in the

Bering Sea The Bering Sea (, ; rus, Бе́рингово мо́ре, r=Béringovo móre) is a marginal sea of the Northern Pacific Ocean. It forms, along with the Bering Strait, the divide between the two largest landmasses on Earth: Eurasia and The Ameri ...

is the largest single species fishery in the world (3 million tonnes). The Alaskan pollock is said to be the largest remaining single species source of palatable fish in the world. However, the biomass of pollock has declined in recent years, perhaps spelling trouble for both the

Bering Sea The Bering Sea (, ; rus, Бе́рингово мо́ре, r=Béringovo móre) is a marginal sea of the Northern Pacific Ocean. It forms, along with the Bering Strait, the divide between the two largest landmasses on Earth: Eurasia and The Ameri ...

ecosystem and the commercial fishery it supports. Acoustic surveys by

NOAA The National Oceanic and Atmospheric Administration (abbreviated as NOAA ) is an United States scientific and regulatory agency within the United States Department of Commerce that forecasts weather, monitors oceanic and atmospheric conditio ...

indicate that the 2008 pollock population is almost 50 percent lower than last year's survey levels. Some scientists think this decline in Alaska pollock could repeat the collapse experienced by

Atlantic cod The Atlantic cod (''Gadus morhua'') is a benthopelagic fish of the family Gadidae, widely consumed by humans. It is also commercially known as cod or codling.omega-3 fatty acid Omega−3 fatty acids, also called Omega-3 oils, ω−3 fatty acids or ''n''−3 fatty acids, are polyunsaturated fatty acids (PUFAs) characterized by the presence of a double bond, three atoms away from the terminal methyl group in their chem ...

s in their flesh. Ninety percent is processed into

fishmeal Fish meal is a commercial product made from whole wild-caught fish, bycatch and fish by-products to feed farm animals, e.g., pigs, poultry, and farmed fish.R. D. Miles and F. A. Chapman.FA122: The Benefits of Fish Meal in Aquaculture DietsFisherie ...

and

fish oil Fish oil is oil derived from the tissues of oily fish. Fish oils contain the omega-3 fatty acids eicosapentaenoic acid (EPA) and docosahexaenoic acid (DHA), precursors of certain eicosanoids that are known to reduce inflammation in the body a ...

. Of this, 46 percent was fed to

farmed fish upright=1.3, Salmon farming in the sea (mariculture) at Loch Ainort, Isle of Skye">mariculture.html" ;"title="Salmon farming in the sea (mariculture">Salmon farming in the sea (mariculture) at Loch Ainort, Isle of Skye, Scotland Fish farming or ...

, 24 percent to pigs, and 22 percent to poultry (2002). Six times the weight of forage fish is fed to pigs and poultry alone than the entire seafood consumption of the U.S. market. One of the most promising alternatives to fish oil as a source of long chain omega-3 fatty acids and certain amino acids is algal oil from

microalgae Microalgae or microphytes are microscopic algae invisible to the naked eye. They are phytoplankton typically found in freshwater and marine systems, living in both the water column and sediment. They are unicellular species which exist indiv ...

, the original source of these fatty acids in forage fish. According to Turchini and De Silva (2008), another 2.5 million tonnes of the annual forage fish catch is consumed by the global cat food industry. In Australia, pet cats eat 13.7 kilograms of fish a year compared to the 11 kilograms eaten by the average Australian. The pet food industry is increasingly marketing premium and super-premium products, when different raw materials, such as the by-products of the fish filleting industry, could be used instead.

Environmental issues

In 2008 the

Sea Around Us Project The Sea Around Us is an international research initiative and a member of the Global Fisheries Cluster at the University of British Columbia. The Sea Around Us assesses the impact of fisheries on the marine ecosystems of the world and offers mitig ...

completed a nine-year study of forage fish led by the fisheries scientists Jacqueline Alder and

Daniel Pauly Daniel Pauly is a French-born marine biologist, well known for his work in studying human impacts on global fisheries and in 2020 was the most cited fisheries scientist in the world. He is a professor and the project leader of the Sea Around Us ...

. They concluded that... In 2015 sardine populations crashed along the west coast of the United States, causing the fishery to close early and remain closed through the 2015–2016 season.Plunging sardine numbers spark quick management response
''Chinook Observer'', 14 April 2015. A key reason for the population crash, was

overfishing Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing fish stock), resulting in th ...

due to the demand of fish meals and fish oil used in feed for aquaculture and for human nutritional supplements. In an effort to provide some relief from the pressure put on forage fish populations, the World Bank along with the University of Arizona, Monterey Bay Aquarium and the New England Aquarium has sponsored a competition called the F3 (Fish-Free Feed) Challenge, which will award $200,000 to the most successful fish feed manufacturer who develops aquaculture feeds not made from fish.

In lakes and rivers

Forage fish also inhabit freshwater habitats, such as lakes and rivers, where they serve as food for larger freshwater predators. Usually smaller than 15 centimetres (6 in) in length, these small

bait fish 300px, Feeder Goldfish are common baitfish. Bait fish (or baitfish) are small-sized fish caught and used by anglers as bait to attract larger predatory fish, particularly game fish. Baitfish species are typically those that are common and bree ...

make up most of the fish found in lakes and rivers. The

minnow Minnow is the common name for a number of species of small freshwater fish, belonging to several genera of the families Cyprinidae and Leuciscidae. They are also known in Ireland as pinkeens. Smaller fish in the subfamily Leusciscidae are co ...

family alone, consisting of

minnow Minnow is the common name for a number of species of small freshwater fish, belonging to several genera of the families Cyprinidae and Leuciscidae. They are also known in Ireland as pinkeens. Smaller fish in the subfamily Leusciscidae are co ...

s, chubs, shiners and

dace A dace is a small fish that can be one of many different species. The unmodified name is usually a reference to the common dace (''Leuciscus leuciscus''). This, like most fish called "daces", belongs to the family Cyprinidae, mostly in subfamily ...

s, consists of more than fifty species.Kleber, John E (1992)
''The Kentucky Encyclopedia''
University Press of Kentucky, Page 320. Other freshwater forage fish include suckers,

killifish A killifish is any of various oviparous (egg-laying) cyprinodontiform fish (including families Aplocheilidae, Cyprinodontidae, Fundulidae, Profundulidae and Valenciidae). All together, there are 1,270 species of killifish, the biggest family ...

,

shad The Alosinae, or the shads,Alosinae

,

bony fish Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartilag ...

as well as fish of the sunfish family, excluding

black bass Black is a color which results from the absence or complete absorption of visible light. It is an achromatic color, without hue, like white and grey. It is often used symbolically or figuratively to represent darkness. Black and white have of ...

es and

crappie Crappies () are two species of North American freshwater fish of the genus ''Pomoxis'' in the family Centrarchidae (sunfishes). Both species of crappies are popular game fish among recreational anglers. Etymology The genus name ''Pomoxis'' ...

, and smaller species of the

carp Carp are various species of oily freshwater fish from the family Cyprinidae, a very large group of fish native to Europe and Asia. While carp is consumed in many parts of the world, they are generally considered an invasive species in parts of ...

family. There are also

anadromous Fish migration is mass relocation by fish from one area or body of water to another. Many types of fish migrate on a regular basis, on time scales ranging from daily to annually or longer, and over distances ranging from a few metres to thousan ...

forage fish, such as

eulachon The eulacheon ( (''Thaleichthys pacificus''), also spelled oolichan , ooligan , hooligan ), also called the candlefish, is a small anadromous species of smelt that spawns in some of the major river systems along the Pacific coast of North Americ ...

. In the context of human activity, within any fresh or

saltwater Saline water (more commonly known as salt water) is water that contains a high concentration of dissolved salts (mainly sodium chloride). On the United States Geological Survey (USGS) salinity scale, saline water is saltier than brackish water, ...

ecosystem, there will always be both desirable and undesirable fishes, and this varies from country to country, and often from region to region within a country. Sport fishermen divide freshwater predators of forage fish into those: * which have a good fighting ability and are good to eat, called sport (or game) fish. * the other less desirable fish, called

rough fish Rough fish (or the slang trash fish or dirt fish) is a term used by some United States state agencies and anglers to describe fish that are less desirable to sport anglers within a defined region. The term usually refers to larger game fish species ...

in North America and coarse fish in Britain ''Rough or coarse fish'' usually refers to fish that are not commonly eaten, not sought after for sporting reasons, or have become invasive species reducing the populations of desirable fish. They compete for forage fish with the more popular sport fish. They are often regarded as a nuisance, and are not usually protected by game laws. Forage fish generally are not considered rough or coarse fish because of their usefulness as bait. The term ''rough fish'' is used by U.S. state agencies and anglers to describe undesirable predator fish. In North America, anglers fish for

salmon Salmon () is the common name for several list of commercially important fish species, commercially important species of euryhaline ray-finned fish from the family (biology), family Salmonidae, which are native to tributary, tributaries of the ...

,

trout Trout are species of freshwater fish belonging to the genera '' Oncorhynchus'', ''Salmo'' and ''Salvelinus'', all of the subfamily Salmoninae of the family Salmonidae. The word ''trout'' is also used as part of the name of some non-salmoni ...

, bass,

pike Pike, Pikes or The Pike may refer to: Fish * Blue pike or blue walleye, an extinct color morph of the yellow walleye ''Sander vitreus'' * Ctenoluciidae, the "pike characins", some species of which are commonly known as pikes * ''Esox'', genus of ...

,

catfish Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive, ...

,

walleye The walleye (''Sander vitreus'', synonym ''Stizostedion vitreum''), also called the yellow pike or yellow pickerel, is a freshwater perciform fish native to most of Canada and to the Northern United States. It is a North American close relat ...

and

muskellunge The muskellunge ''(Esox masquinongy)'', often shortened to muskie, musky or lunge is a species of large freshwater predatory fish native to North America. It is the largest member of the pike family, Esocidae. Origin of name The name "muskellun ...

. The smallest fish are called

panfish The word panfish, also spelled pan-fish or pan fish, is an American English term describing any edible freshwater fish that usually do not outgrow the size of an average frying pan. It is also commonly used by recreational anglers to refer to any ...

, because they can fit in a standard cooking pan. Some examples are

crappie Crappies () are two species of North American freshwater fish of the genus ''Pomoxis'' in the family Centrarchidae (sunfishes). Both species of crappies are popular game fish among recreational anglers. Etymology The genus name ''Pomoxis'' ...

s,

rock bass The rock bass (''Ambloplites rupestris''), also known as the rock perch, goggle-eye, red eye, and black perch, is a freshwater fish native to east-central North America. This red eyed creature is a species of freshwater fish in the sunfish fa ...

,

perch Perch is a common name for fish of the genus ''Perca'', freshwater gamefish belonging to the family Percidae. The perch, of which three species occur in different geographical areas, lend their name to a large order of vertebrates: the Percif ...

,

bluegill The bluegill (''Lepomis macrochirus''), sometimes referred to as "bream", "brim", "sunny", or "copper nose" as is common in Texas, is a species of North American freshwater fish, native to and commonly found in streams, rivers, lakes, ponds and ...

and sunfish. The term ''coarse fish'' originated in the United Kingdom in the early 19th century. Prior to that time, recreational fishing was the sport of the

gentry Gentry (from Old French ''genterie'', from ''gentil'', "high-born, noble") are "well-born, genteel and well-bred people" of high social class, especially in the past. Word similar to gentle imple and decentfamilies ''Gentry'', in its widest ...

, who angled for

trout Trout are species of freshwater fish belonging to the genera '' Oncorhynchus'', ''Salmo'' and ''Salvelinus'', all of the subfamily Salmoninae of the family Salmonidae. The word ''trout'' is also used as part of the name of some non-salmoni ...

and

salmon Salmon () is the common name for several list of commercially important fish species, commercially important species of euryhaline ray-finned fish from the family (biology), family Salmonidae, which are native to tributary, tributaries of the ...

which they called "game fish". Fish other than game fish were disdained as "coarse fish". These days, "game fish" refers to

Salmonid Salmonidae is a family of ray-finned fish that constitutes the only currently extant family in the order Salmoniformes . It includes salmon (both Atlantic and Pacific species), trout (both ocean-going and landlocked), chars, freshwater whitefis ...

s (other than grayling) — that is,

salmon Salmon () is the common name for several list of commercially important fish species, commercially important species of euryhaline ray-finned fish from the family (biology), family Salmonidae, which are native to tributary, tributaries of the ...

,

trout Trout are species of freshwater fish belonging to the genera '' Oncorhynchus'', ''Salmo'' and ''Salvelinus'', all of the subfamily Salmoninae of the family Salmonidae. The word ''trout'' is also used as part of the name of some non-salmoni ...

and

char Char may refer to: People *Char Fontane, American actress *Char Margolis, American spiritualist * René Char (1907–1988), French poet *The Char family of Colombia: ** Fuad Char, Colombian senator ** Alejandro Char Chaljub, mayor of Barranquilla ...

. Coarse fish are made up mostly of the larger species of

Cyprinid Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true minnows, and relatives like the barbs and barbels. Cyprinidae is the largest and most diverse fish family and the largest verte ...

s (

carp Carp are various species of oily freshwater fish from the family Cyprinidae, a very large group of fish native to Europe and Asia. While carp is consumed in many parts of the world, they are generally considered an invasive species in parts of ...

,

roach Roach may refer to: Animals * Cockroach, various insect species of the order Blattodea * Common roach (''Rutilus rutilus''), a fresh and brackish water fish of the family Cyprinidae ** ''Rutilus'' or roaches, a genus of fishes * California roach ...

,

bream Bream ( ) are species of freshwater and marine fish belonging to a variety of genera including ''Abramis'' (e.g., ''A. brama'', the common bream), ''Acanthopagrus'', '' Argyrops'', ''Blicca'', '' Brama'', ''Chilotilapia'', '' Etelis'', ''Lepo ...

) as well as

pike Pike, Pikes or The Pike may refer to: Fish * Blue pike or blue walleye, an extinct color morph of the yellow walleye ''Sander vitreus'' * Ctenoluciidae, the "pike characins", some species of which are commonly known as pikes * ''Esox'', genus of ...

,

catfish Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive, ...

,

gar Gars are members of the family Lepisosteidae, which are the only surviving members of the Ginglymodi, an ancient holosteian group of ray-finned fish, which first appeared during the Triassic, over 240 million years ago. Gars comprise seven livin ...

and

lamprey Lampreys (sometimes inaccurately called lamprey eels) are an ancient extant lineage of jawless fish of the order Petromyzontiformes , placed in the superclass Cyclostomata. The adult lamprey may be characterized by a toothed, funnel-like s ...

. Coarse fish are no longer disdained; indeed, fishing for coarse fish has become a popular pastime.

Bait and feeder fish

Forage fish are sometimes referred to as ''bait fish'' or ''feeder fish''.

Bait fish 300px, Feeder Goldfish are common baitfish. Bait fish (or baitfish) are small-sized fish caught and used by anglers as bait to attract larger predatory fish, particularly game fish. Baitfish species are typically those that are common and bree ...

is a term used particularly by recreational fishermen, although commercial fisherman also catch fish to bait longlines and traps. Forage fish is a

fisheries Fishery can mean either the enterprise of raising or harvesting fish and other aquatic life; or more commonly, the site where such enterprise takes place ( a.k.a. fishing ground). Commercial fisheries include wild fisheries and fish farms, both ...

term, and is used in the context of fisheries. Bait fish, by contrast, are fish that are caught by humans to use as bait for other fish. The terms overlap in the sense that most bait fish are also forage fish, and vice versa.

Feeder fish Feeder fish is the common name for certain types of small, inexpensive fish commonly fed as live food to other captive animals such as predatory fishes (e.g. sharks in captivity, aquarium sharks, aquaculture of salmonids, farmed salmon and tuna p ...

is a term used particularly in the context of fish

aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...

s. It refers essentially to the same concept as forage fish, small fish that are eaten by larger fish, but the term is adapted to the particular requirements of working with fish in aquariums.

Timeline

* 2006: The U.S.

National Coalition for Marine Conservation National Coalition for Marine Conservation is now operating as "Wild Oceans." Founded in 1973 by fishermen, the National Coalition for Marine Conservation (NCMC) is the United States's oldest public advocacy group dedicated exclusively to conserving ...

asks U.S. fishery managers to put "Forage First!". Their campaign was launched with the publication of their report, ''Taking the Bait: Are America’s Fisheries Out-competing Predators for their Prey?'', available at cost to the U. S. fishing industry, encouraging fishery managers to protect predator–prey relationships as a first step toward an ecosystem based approach to fishery management. * 2009: The international Lenfest Forage Fish Task Force is established to develop workable management plans for tackling the depletion of forage fish. * 2015: sardine populations crashed along the west coast of the United States.

Recent reports

* Pikitch E and 12 others (2012
Little Fish, Big Impact: Managing a Crucial Link in Ocean Food Webs
''Lenfest Ocean Program'', Washington, DC. Summary and other materials available on th
Lenfest Ocean Program website

Notes

References

*

External links

Herring Research: Using Acoustics to Count Fish
* Sanders, Michael (1995

FAO The Food and Agriculture Organization of the United Nations (FAO)french: link=no, Organisation des Nations unies pour l'alimentation et l'agriculture; it, Organizzazione delle Nazioni Unite per l'Alimentazione e l'Agricoltura is an intern ...

. * Skjoldal, Hein Rune (2011
"Ecological effects of fisheries on small pelagics"
Background document for CBD Expert meeting, Bergen, 7–9 December 2011.
Daniel Pauly at Fish Forever
– YouTube: Pauly talks about his "epiphany" about directly eating anchovies. {{DEFAULTSORT:Forage Fish Fishing industry Ichthyology

In the oceans

Ocean food webs

Diet

Predators

Schooling

Hunting copepods

Migrations

Predator attacks

Forage fisheries

History

Contemporary

Environmental issues

In lakes and rivers

Bait and feeder fish

Timeline

Recent reports

See also

Notes

References

External links