''Polistes aurifer'', the golden paper wasp, is a species of

paper wasp Paper wasps are Vespidae, vespid wasps that gather fibers from dead wood and plant stems, which they mix with saliva, and use to construct nests made of gray or brown papery material. Some types of paper wasps are also sometimes called umbrell ...

in the genus ''

Polistes Wasps of the cosmopolitan genus ''Polistes'' (the only genus in the tribe Polistini) are the most familiar of the polistine wasps, and are the most common type of paper wasp in North America. Walter Ebeling coined the vernacular name "umbrella ...

'' of the family

Vespidae The Vespidae are a large (nearly 5000 species), diverse, cosmopolitan family of wasps, including nearly all the known eusocial wasps (such as ''Polistes fuscatus'', ''Vespa orientalis'', and ''Vespula germanica'') and many solitary wasps. Each ...

. It occurs in the western part of North America, from southern

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

through the

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

to northern

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

Taxonomy

It was first described as a new species by

Henri Louis Frédéric de Saussure Henri Louis Frédéric de Saussure (; ; 27 November 1829 – 20 February 1905) was a Swiss mineralogist and entomologist specialising in studies of Hymenoptera and Orthopteroid insects. He also was a prolific taxonomist. Biography Sauss ...

in 1853. Roy Snelling synonymised a number of taxa to this species in 1954.

Owain Richards Owain Westmacott Richards FRS (31 December 1901 – 10 November 1984) was a British entomologist and ecologist who worked as Professor of Zoology and Applied Entomology, Imperial College, London, based at Silwood Park, and an editor of the ''Jou ...

treated this

taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...

as a

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

of ''P. fuscatus'' in his 1978 work, which was followed in subsequent works.

Etymology

The

etymology Etymology ()The New Oxford Dictionary of English (1998) – p. 633 "Etymology /ˌɛtɪˈmɒlədʒi/ the study of the class in words and the way their meanings have changed throughout time". is the study of the history of the Phonological chan ...

of the

specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...

''aurifex'' is from the

Latin language Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...

, where it means a 'goldsmith' or 'one who works with gold'. This word is derived from the Latin word ''aurum'', meaning 'gold', combined with the

suffix In linguistics, a suffix is an affix which is placed after the stem of a word. Common examples are case endings, which indicate the grammatical case of nouns, adjectives, and verb endings, which form the conjugation of verbs. Suffixes can carry ...

''-fer'', derived from the word ''facio'' and meaning '-bearing', '-carrying' or '-bringing', together giving 'gold-bearer'.

Description

The

morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...

of the male

genitals A sex organ (or reproductive organ) is any part of an animal or plant that is involved in sexual reproduction. The reproductive organs together constitute the reproductive system. In animals, the testis in the male, and the ovary in the female, a ...

are very diagnostic for the species. The species has different inter-grading colour patterns depending on geography. Northern specimens are often mostly black, with ample yellow markings but with the

ferruginous The adjective ferruginous may mean: * Containing iron, applied to water, oil, and other non-metals * Having rust on the surface * With the rust (color) See also * Ferrous, containing iron (for metals and alloys) or iron(II) cations * Ferric, conta ...

(rusty red) colour being very restricted. In populations from the southwestern USA forms with an almost completely yellow

metasoma The metasoma is the posterior part of the body, or tagma, of arthropods whose body is composed of three parts, the other two being the prosoma and the mesosoma. In insects, it contains most of the digestive tract, respiratory system, and circula ...

are more common. In some southern US and northwestern Mexican populations the ferruginous colour predominates, with the black colour severely restricted.

Similar species

It has been confused with extremely xanthic specimens of ''P. apachus''. It has also been confused with ''P. fuscatus''. Adults of either sex can be distinguished from this species by the characteristic of having a ferruginous

dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal co ...

surface of the first, and sometimes the following,

flagellomere Antennae ( antenna), sometimes referred to as "feelers", are paired appendages used for sensing in arthropods. Antennae are connected to the first one or two segments of the arthropod head. They vary widely in form but are always made of one o ...

s. In ''P. fuscatus'' this surface is darkly coloured. The wings are a more yellowish colour in ''P. aurifer''. Furthermore ''P. aurifer'' possesses a pair of large yellow patches on the second

tergum A ''tergum'' (Latin for "the back"; plural ''terga'', associated adjective tergal) is the Anatomical terms of location#Dorsal and ventral, dorsal ('upper') portion of an arthropod segment other than the head. The Anatomical terms of location#Ante ...

; these spots are usually disc-shaped, sometimes hidden and confluent with neighbouring yellow in especially xanthic individuals.

Distribution

In Canada it has been recorded in southernmost areas of the provinces of

British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...

(including southern

Vancouver Island Vancouver Island is an island in the northeastern Pacific Ocean and part of the Canadian Provinces and territories of Canada, province of British Columbia. The island is in length, in width at its widest point, and in total area, while are o ...

Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...

and

Saskatchewan Saskatchewan ( ; ) is a Provinces and territories of Canada, province in Western Canada, western Canada, bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, to the northeast by Nunavut, and on t ...

. In the USA it has been recorded in

Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...

, far west

North Dakota North Dakota () is a U.S. state in the Upper Midwest, named after the Native Americans in the United States, indigenous Dakota people, Dakota Sioux. North Dakota is bordered by the Canadian provinces of Saskatchewan and Manitoba to the north a ...

, western

South Dakota South Dakota (; Sioux language, Sioux: , ) is a U.S. state in the West North Central states, North Central region of the United States. It is also part of the Great Plains. South Dakota is named after the Lakota people, Lakota and Dakota peo ...

Idaho Idaho ( ) is a state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Montana and Wyom ...

, westernmost

Kansas Kansas () is a state in the Midwestern United States. Its capital is Topeka, and its largest city is Wichita. Kansas is a landlocked state bordered by Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to the ...

Montana Montana () is a state in the Mountain West division of the Western United States. It is bordered by Idaho to the west, North Dakota and South Dakota to the east, Wyoming to the south, and the Canadian provinces of Alberta, British Columbi ...

, western

Nebraska Nebraska () is a state in the Midwestern region of the United States. It is bordered by South Dakota to the north; Iowa to the east and Missouri to the southeast, both across the Missouri River; Kansas to the south; Colorado to the southwe ...

Nevada Nevada ( ; ) is a U.S. state, state in the Western United States, Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. N ...

New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...

Oregon Oregon () is a U.S. state, state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of it ...

, far west

Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...

Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...

Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered o ...

and

Wyoming Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the south ...

(a single record in the far northwest). In Mexico it has been seen in the states of

Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...

Baja California Sur Baja California Sur (; 'South Lower California'), officially the Free and Sovereign State of Baja California Sur ( es, Estado Libre y Soberano de Baja California Sur), is the least populated state and the 31st admitted state of the 32 federal ent ...

, Chihuahua,

Coahuila Coahuila (), formally Coahuila de Zaragoza (), officially the Free and Sovereign State of Coahuila de Zaragoza ( es, Estado Libre y Soberano de Coahuila de Zaragoza), is one of the 32 states of Mexico. Coahuila borders the Mexican states of N ...

Nuevo León Nuevo León () is a state in the northeast region of Mexico. The state was named after the New Kingdom of León, an administrative territory from the Viceroyalty of New Spain, itself was named after the historic Spanish Kingdom of León. With a ...

and northern

Sonora Sonora (), officially Estado Libre y Soberano de Sonora ( en, Free and Sovereign State of Sonora), is one of the 31 states which, along with Mexico City, comprise the Administrative divisions of Mexico, Federal Entities of Mexico. The state is d ...

. Adventive populations have also been recorded in

Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...

Johnston Atoll Johnston Atoll is an Unincorporated territories of the United States, unincorporated territory of the United States, currently administered by the United States Fish and Wildlife Service (USFWS). Johnston Atoll is a National Wildlife Refuge and ...

Niihau Niihau ( Hawaiian: ), anglicized as Niihau ( ), is the westernmost main and seventh largest inhabited island in Hawaii. It is southwest of Kauaʻi across the Kaulakahi Channel. Its area is . Several intermittent playa lakes provide wetland hab ...

and the

Society Islands The Society Islands (french: Îles de la Société, officially ''Archipel de la Société;'' ty, Tōtaiete mā) are an archipelago located in the South Pacific Ocean. Politically, they are part of French Polynesia, an overseas country of the F ...

. A single specimen was also caught in

Prince Edward Island Prince Edward Island (PEI; ) is one of the thirteen Provinces and territories of Canada, provinces and territories of Canada. It is the smallest province in terms of land area and population, but the most densely populated. The island has seve ...

in 1973.

Ecology

This is an

eusocial Eusociality (from Greek εὖ ''eu'' "good" and social), the highest level of organization of sociality, is defined by the following characteristics: cooperative brood care (including care of offspring from other individuals), overlapping generat ...

wasp which nests in sheltered locations. Adults have been seen from January to November in more southern locations such as California or Arizona, from April to October in British Columbia in the north of the range, from April to November in more higher and

continental Continental may refer to: Places * Continent, the major landmasses of Earth * Continental, Arizona, a small community in Pima County, Arizona, US * Continental, Ohio, a small town in Putnam County, US Arts and entertainment * ''Continental'' ( ...

areas such as Colorado, or having an even more restricted season in northern continental areas such as Alberta or Saskatchewan. It is parasitised upon by the

endoparasitoid In evolutionary ecology, a parasitoid is an organism that lives in close association with its host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionary strategies within parasiti ...

'' Xenos peckii'' and an unidentified '' Gordius'' species, a type of

horsehair worm Nematomorpha (sometimes called Gordiacea, and commonly known as horsehair worms, hairsnakes, or Gordian worms) are a phylum of parasitoid animals superficially similar to nematode worms in morphology, hence the name. Most species range in size ...

Conservation

The

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

has

not evaluated A not evaluated (NE) species is one which has been categorized under the IUCN Red List of threatened species as not yet having been assessed by the International Union for Conservation of Nature. This conservation category is one of nine IUCN th ...

this species'

conservation status The conservation status of a group of organisms (for instance, a species) indicates whether the group still exists and how likely the group is to become extinct in the near future. Many factors are taken into account when assessing conservation ...

References

aurifer Insects described in 1853 {{vespidae-stub