Plagiaulacida
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Plagiaulacida is a group of extinct
multituberculate Multituberculata (commonly known as multituberculates, named for the multiple tubercles of their teeth) is an extinct order of rodent-like mammals with a fossil record spanning over 130 million years. They first appeared in the Middle Jurassic, a ...
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s. Multituberculates were among the most common mammals of the
Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceo ...
, "the age of the
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s". Plagiaulacids are a paraphyletic grouping, containing all multituberculates that lie outside of the advanced group
Cimolodonta Cimolodonta is a taxon of extinct mammals that lived from the Cretaceous to the Eocene. They were some of the more derived members of the extinct order Multituberculata. They probably lived something of a rodent-like existence until their ecolo ...
. They ranged from the Middle
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
Period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...
to the early Late Cretaceous of the
northern hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
. During the
Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
, they were replaced by the more advanced cimolodontans. Kielan-Jaworowska and Hurum (2001) divides “Plagiaulacida” into three informal lineages, the paulchoffatiids, the plagiaulicids, and the allodontids.


Allodontid line

The Allodontid line may be a superfamily, Allodontoidea. Both allodontids and paulchoffatiids (below) were among the most basal of the plagiaulacids. The Allodontid line contains: The family
Allodontidae Allodontidae (from ancient Greek "ἄλλος" "ὀδούς", "different tooth") is a family of extinct multituberculate mammal that lived in what is now North America during the Upper Jurassic period. They were relatively early mammals and ar ...
is known from two genera from the Upper Jurassic
Morrison Formation The Morrison Formation is a distinctive sequence of Late Jurassic, Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandsto ...
of
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. The family Zofiabaataridae contains a single genus, '' Zofiabaatar'' and is also from the Morrison Formation. The affinities of a further Morrison Formation genus, ''
Glirodon ''Glirodon'' is a genus of extinct mammal from the Upper Jurassic. It was a relatively early member of the also-extinct order of Multituberculata, suborder "Plagiaulacida". These mammals lived in North America during the Mesozoic, also known a ...
'', are unclear, but it's also within the Allodontid line.


Paulchoffatiid line

The Paulchoffatiid line may be a superfamily, Paulchoffatioidea. Some remains from the Middle Jurassic of England might belong within this group. Representatives are best known from the Upper Jurassic, (especially from Guimarota, Portugal), though some were still extant during the Lower Cretaceous. The genera of the family
Paulchoffatiidae Paulchoffatiidae is a family of extinct mammals that lived predominantly during the Upper Jurassic period, though a couple of Genus, genera are known from the Early Cretaceous. Fossils have been reported from Europe (Portugal, Spain, Germany and ...
are divided into two of subfamilies, plus a couple of harder-to-place individuals: *Subfamily Paulchoffatiinae includes ''
Paulchoffatia ''Paulchoffatia'' is a genus of extinct mammal of the Upper Jurassic - Lower Cretaceous. It was a relatively early member of the also extinct order Multituberculata, within the suborder "Plagiaulacida" and family Paulchoffatiidae. It lived in ...
'' and its relatives. This taxon contains nine genera. *Subfamily Kuehneodontinae consists solely of the genus ''
Kuehneodon ''Kuehneodon'' is a genus of extinct mammal of the Upper Jurassic - Lower Cretaceous of Europe. It was a relatively early member of the also extinct order of Multituberculata. Members of this genus lived alongside such dinosaurs as Allosaurus. ...
'', though there are half-a-dozen named species. Other genera include '' Galveodon'' and '' Sunnyodon'', both based on teeth from the Lower Cretaceous of Spain and England respectively. Also referable to the paulchoffatiid line, but not the family itself, is the family
Pinheirodontidae Pinheirodontidae is a poorly known family of fossil mammals which belong to the informal suborder "Plagiaulacida" within the order Multituberculata. Remains are known from the Late Jurassic to Early Cretaceous of Europe, (predominantly Portugal ...
, which is known from Early Cretaceous teeth found in
Iberia The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
and England. As well as ''
Rugosodon ''Rugosodon'' is an extinct genus of multituberculate (rodent-like) mammals from eastern China that lived 160 million years ago during the Jurassic period. The discovery of its type species and currently only known species ''Rugosodon eurasiaticu ...
'' from the Middle-Late Jurassic of China.


Plagiaulacid line (possibly Superfamily Plagiaulacoidea)

Family
Plagiaulacidae Plagiaulacidae is a family of fossil mammals within the order Multituberculata. Remains are known from the Upper Jurassic and earliest Cretaceous of North America and Europe. They were among the more derived representatives of the informal subo ...
is known from the Upper Jurassic (North America) to Lower Cretaceous (Europe), being represented by ''
Plagiaulax ''Plagiaulax'' is a genus of mammal from the Lower Cretaceous of Europe. It was a member of the also extinct order Multituberculata, and shared the world with dinosaurs. It is of the suborder "Plagiaulacida" and family Plagiaulacidae. The genus ...
'', ''
Bolodon ''Bolodon'' ("lump tooth" from Ancient Greek βῶλος (bôlos), “clod, lump” + ὀδών (odṓn), "tooth") is a genus of extinct mammal from the Lower Cretaceous of Europe and North America. It was a member of the extinct order of Multi ...
'', and '' Morrisonodon''. Family Albionbaataridae is known from the Upper Jurassic to Lower Cretaceous of Europe and Asia, (China – undescribed, 2001). These were shrew-sized Multituberculates, with some similarities to the paulchoffis. Members of the family
Eobaataridae Eobaataridae is a family of fossil mammals within the order Multituberculata. Remains are known from the Lower Cretaceous of Europe and Asia. They are among the most derived representatives of the informal suborder "Plagiaulacida", and closely re ...
display dental similarities with members of ''Paracimexomys'' group, (
Cimolodonta Cimolodonta is a taxon of extinct mammals that lived from the Cretaceous to the Eocene. They were some of the more derived members of the extinct order Multituberculata. They probably lived something of a rodent-like existence until their ecolo ...
). They are known from the Upper Jurassic to Lower Cretaceous of Europe and Asia. '' Sinobaatar'' was described after the study by Kielan-Jaworowska and Hurum (2001). The
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
n word ‘baatar’ is frequently employed in the nomenclature of Multituberculates. This reflects the fact that many of the most complete fossils have been recovered from sites in Mongolia, though this more applies to members of the more derived
Cimolodonta Cimolodonta is a taxon of extinct mammals that lived from the Cretaceous to the Eocene. They were some of the more derived members of the extinct order Multituberculata. They probably lived something of a rodent-like existence until their ecolo ...
. A couple of further genera possibly fit somewhere within “Plagiaulacida”. This has been tentatively proposed for ''
Janumys ''Janumys'' is a genus of extinct mammal of the middle Cretaceous. It was a member of the order of Multituberculata (also extinct). It lived in North America during the Mesozoic era, also known as the "age of the reptiles." It has been provi ...
'' of the Middle Cretaceous. Its contemporary, '' Ameribaatar'', is of uncertain affinities. Both were first described late in 2001.


Taxonomy

Subclass †
Allotheria Allotheria (meaning "other beasts", from the Greek , '–other and , '–wild animal) is an extinct branch of successful Mesozoic mammals. The most important characteristic was the presence of lower molariform teeth equipped with two longitudin ...
* Order †
Multituberculata Multituberculata (commonly known as multituberculates, named for the multiple tubercles of their teeth) is an extinct order of rodent-like mammals with a fossil record spanning over 130 million years. They first appeared in the Middle Jurassic, a ...
: ** suborder †Plagiaulacida *** Family †
Paulchoffatiidae Paulchoffatiidae is a family of extinct mammals that lived predominantly during the Upper Jurassic period, though a couple of Genus, genera are known from the Early Cretaceous. Fossils have been reported from Europe (Portugal, Spain, Germany and ...
**** subfamily †
Paulchoffatiinae Paulchoffatiidae is a family of extinct mammals that lived predominantly during the Upper Jurassic period, though a couple of Genus, genera are known from the Early Cretaceous. Fossils have been reported from Europe (Portugal, Spain, Germany and ...
***** Genus †''
Paulchoffatia ''Paulchoffatia'' is a genus of extinct mammal of the Upper Jurassic - Lower Cretaceous. It was a relatively early member of the also extinct order Multituberculata, within the suborder "Plagiaulacida" and family Paulchoffatiidae. It lived in ...
'' ****** Species †'' P. delgador'' ***** Genus †'' Pseudobolodon'' ****** Species †'' P. oreas'' ****** Species †'' P. krebsi'' ***** Genus †'' Henkelodon'' ****** Species †'' H. naias'' ***** Genus †'' Guimarotodon'' ****** Species †'' G. leiriensis'' ***** Genus †''
Meketibolodon ''Meketibolodon'' is a genus of extinct mammal from the Kimmeridgian (Upper Jurassic) Camadas de Guimarota of Guimarota, Portugal. It was a relatively early member of the also extinct order Multituberculata, suborder Plagiaulacida, family ...
'' ****** Species †'' M. robustus'' ***** Genus †''
Plesiochoffatia ''Plesiochoffatia'' is an extinct mammal of the Upper Jurassic. It was a relatively early member of the also extinct order Multituberculata. It was a resident of Portugal during the "age of the dinosaurs." It's in the suborder "Plagiaulacida" ...
'' ****** Species †'' P. thoas'' ****** Species †'' P. peparethos'' ****** Species †'' P. staphylos'' ***** Genus †''
Xenachoffatia ''Xenachoffatia'' is a small Jurassic mammal from Portugal. It was a relatively early member of the also extinct order of Multituberculata. It lived during "the age of the dinosaurs" and belongs to the suborder Plagiaulacida, family Paulchoffa ...
'' ****** Species †'' X. oinopion'' ***** Genus †''
Bathmochoffatia ''Bathmochoffatia'' is an extinct mammal of the Upper Jurassic. It was a relatively early member of the also extinct order Multituberculata. It lived in Portugal at about the same time as the far more famous dinosaur, Allosaurus. It is in the s ...
'' ****** Species †'' B. hapax'' ***** Genus †''
Kielanodon ''Kielanodon'' is an extinct mammal of the Portuguese Upper Jurassic. It was a relatively early member of the also extinct order of Multituberculata. It eked out a living during the Mesozoic era, also known as the "Age of the Dinosaurs." It i ...
'' ****** Species †'' K. hopsoni'' ***** Genus †''
Meketichoffatia ''Meketichoffatia'' was a small mammal from the Upper Jurassic of Portugal. It was a relatively early member of the extinct order Multituberculata. It lived at the same time as dinosaurs such as Allosaurus. It's within the suborder "Plagiaul ...
'' ****** Species †'' M. krausei'' ***** Genus †'' Galveodon'' ****** Species †'' G. nannothus'' ***** Genus †'' Sunnyodon'' ****** Species †'' S. notleyi'' **** Subfamily †Kuehneodontinae ***** Genus †''
Kuehneodon ''Kuehneodon'' is a genus of extinct mammal of the Upper Jurassic - Lower Cretaceous of Europe. It was a relatively early member of the also extinct order of Multituberculata. Members of this genus lived alongside such dinosaurs as Allosaurus. ...
'' ****** Species †'' K. dietrichi'' ****** Species †'' K. barcasensis'' ****** Species †'' K. dryas'' ****** Species †'' K. guimarotensis'' ****** Species †'' K. hahni'' ****** Species †'' K. simpsoni'' ****** Species †'' K. uniradiculatus'' **** Family †
Pinheirodontidae Pinheirodontidae is a poorly known family of fossil mammals which belong to the informal suborder "Plagiaulacida" within the order Multituberculata. Remains are known from the Late Jurassic to Early Cretaceous of Europe, (predominantly Portugal ...
***** Genus †'' Pinheirodon'' ****** Species †'' P. pygmaeus'' ****** Species †'' P. vastus'' ***** Genus †''
Bernardodon ''Bernardodon'' was a small, Lower Cretaceous mammal from Portugal. It is part of the extinct order Multituberculata, living at the same time as the dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first ...
'' ****** Species †'' B. atlantica'' ***** Genus †'' Gerhardodon'' ****** Species †'' G. purbeckensis'' ***** Genus †''
Iberodon ''Iberodon'' is a small, extinct mammal of the Lower Cretaceous from Portugal. It was a member of the also extinct order Multituberculata, and led its obscure and plant-eating existence in the company of dinosaurs. It lies within the suborder ...
'' ****** Species †'' I. quadrituberculatus'' ***** Genus †'' Lavocatia'' ****** Species †'' L. alfambrensis'' ***** Genus †'' Ecprepaulax'' ****** Species †'' E. anomala'' **** Family †
Allodontidae Allodontidae (from ancient Greek "ἄλλος" "ὀδούς", "different tooth") is a family of extinct multituberculate mammal that lived in what is now North America during the Upper Jurassic period. They were relatively early mammals and ar ...
***** Genus †''
Ctenacodon ''Ctenacodon'' ("comb needle tooth" from Ancient Greek κτενός (ktenós), "comb" + Latin acus, "needle" + Ancient Greek ὀδών (odṓn), "tooth") is a genus of extinct mammal that lived in what is now North America during the Upper Jura ...
'' ****** Species †'' C. serratus'' ****** Species †'' C. nanus'' ****** Species †'' C. laticeps'' ****** Species †'' C. scindens'' ***** Genus †'' Psalodon'' ****** Species †'' P. potens'' ****** Species †'' P. fortis'' ****** Species †'' P. marshi'' **** Family † Zofiabaataridae ***** Genus †'' Zofiabaatar'' ****** Species †'' Z. pulcher'' **** Family ''
Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...
'' ***** Genus †''
Glirodon ''Glirodon'' is a genus of extinct mammal from the Upper Jurassic. It was a relatively early member of the also-extinct order of Multituberculata, suborder "Plagiaulacida". These mammals lived in North America during the Mesozoic, also known a ...
'' ***** Species †'' G. grandis'' **** Family †
Plagiaulacidae Plagiaulacidae is a family of fossil mammals within the order Multituberculata. Remains are known from the Upper Jurassic and earliest Cretaceous of North America and Europe. They were among the more derived representatives of the informal subo ...
***** Genus †'' Morrisonodon'' ****** Species †'' M. brentbaatar'' ***** Genus †''
Plagiaulax ''Plagiaulax'' is a genus of mammal from the Lower Cretaceous of Europe. It was a member of the also extinct order Multituberculata, and shared the world with dinosaurs. It is of the suborder "Plagiaulacida" and family Plagiaulacidae. The genus ...
'' ****** Species †'' P. becklesii'' ***** Genus †''
Bolodon ''Bolodon'' ("lump tooth" from Ancient Greek βῶλος (bôlos), “clod, lump” + ὀδών (odṓn), "tooth") is a genus of extinct mammal from the Lower Cretaceous of Europe and North America. It was a member of the extinct order of Multi ...
'' ****** Species †'' B. crassidens'' ****** Species †'' B. falconeri'' ****** Species †'' B. minor'' ****** Species †'' B. osborni'' ****** Species †'' B. elongatus'' **** Family †
Eobaataridae Eobaataridae is a family of fossil mammals within the order Multituberculata. Remains are known from the Lower Cretaceous of Europe and Asia. They are among the most derived representatives of the informal suborder "Plagiaulacida", and closely re ...
***** Genus †''
Eobaatar ''Eobaatar'' is a genus of extinct mammal from the Lower Cretaceous of Mongolia, Spain and England. A member of the also extinct order Multituberculata, it lies within the suborder Plagiaulacida and family Eobaataridae. The genus ''Eobaatar'' w ...
'' ****** Species †'' E. magnus'' ****** Species †'' E. minor'' ****** Species †'' E. hispanicus'' ****** Species †'' E. pajaronensis'' ***** Genus †''
Loxaulax ''Loxaulax'' ("slanting groove" from Ancient Greek λοξός (loxós), “slanting, crosswise” + αὖλαξ (aûlax), "furrow, groove") is a genus of extinct mammal from the Lower Cretaceous of southern England. It was a member of the also e ...
'' ****** Species †'' L. valdensis'' ***** Genus †''
Monobaatar ''Monobaatar'' is a genus of extinct mammal from the Lower Cretaceous of Mongolia. It was within the also extinct order Multituberculata, and lived during the "age of the dinosaurs". It is also within the suborder " Plagiaulacida" and has been t ...
'' ****** Species †'' M. mimicus'' ***** Genus †''
Parendotherium ''Loxaulax'' ("slanting groove" from Ancient Greek λοξός (loxós), “slanting, crosswise” + αὖλαξ (aûlax), "furrow, groove") is a genus of extinct mammal from the Lower Cretaceous of southern England. It was a member of the also e ...
'' ****** Species †'' Parendotherium herreroi'' ***** Genus †'' Sinobaatar'' ****** Species †'' Sinobaatar lingyuanensis'' ***** Genus †''
Heishanobaatar ''Heishanobaatar'' ("Heishan hero" from Chinese 黑山 (Hēishān), " Heishan" + Mongolian baatar, "hero") is an extinct genus of eobaatarid multituberculate which existed in Shahai and Fuxin formations, northeastern China, during the early Cre ...
'' ****** Species †'' H. triangulus'' ***** Genus †'' Teutonodon'' ****** Species †'' Teutonodon langenbergensis'' **** Family † Albionbaataridae ***** Genus †'' Albionbaatar'' ****** Species †'' A. denisae'' ***** Genus †'' Proalbionbaatar'' ****** Species †'' P. plagiocyrtus'' ***** Genus † Kielanobaatar ****** Species †'' K. badaohaoensis'' **** Family †
Arginbaataridae ''Arginbaatar'' is a genus of extinct mammal from the Lower Cretaceous of Mongolia. It was a member of the Multituberculata, an order which is also extinct. It belongs to the family Arginbaataridae (Hahn & Hahn 1983). The genus ''Arginbaatar'' w ...
***** Genus †'' Arginbaatar'' ****** Species †'' Arginbaatar dmitrievae''


References

* Hahn G & Hahn R (2000), ''Multituberculates from the Guimarota mine'', p. 97-107 in * Martin T & Krebs B (eds), ''Guimarota - A Jurassic Ecosystem'', Published by Dr Friedrich Pfeil, Münich, Germany. * Kielan-Jaworowska Z & Hurum JH (2001), Phylogeny and Systematics of multituberculate mammals. Paleontology 44, p. 389-429. * Much of this information has been derived fro

Multituberculata Cope, 1884. {{Taxonbar, from=Q3906163 Multituberculates Jurassic mammals Cretaceous mammals Paraphyletic groups