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''Monotropa hypopitys'', the so-called Dutchman's pipe, false beech-drops, pinesap, or yellow bird's-nest, is a
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
perennial plant A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
, formerly classified in the families Monotropaceae or Pyrolaceae, but now included within the subfamily Monotropoideae of the family
Ericaceae The Ericaceae are a family of flowering plants, commonly known as the heath or heather family, found most commonly in acidic and infertile growing conditions. The family is large, with c.4250 known species spread across 124 genera, making it th ...
. It is native to temperate regions of the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
, and is scarce or rare in many areas. However, it is still the most widespread member of the subfamily. While currently included in the genus ''
Monotropa ''Monotropa'' is a genus of three species of herbaceous perennial flowering plants that were formerly classified in the family Monotropaceae and presently are classified in Ericaceae. They are native to temperate regions of the Northern Hemisph ...
'', recent genetic evidence strongly suggests that ''Monotropa hypopitys'' should be placed in its own genus, ''Hypopitys'', with the single species ''Hypopitys monotropa'' Crantz, but possibly containing several other species. Unlike most plants, it does not contain
chlorophyll Chlorophyll (also chlorophyl) is any of several related green pigments found in cyanobacteria and in the chloroplasts of algae and plants. Its name is derived from the Greek words , ("pale green") and , ("leaf"). Chlorophyll allow plants to a ...
; it is a
myco-heterotroph Myco-heterotrophy (from Ancient Greek, Greek μύκης , "fungus", ἕτερος ', "another", "different" and τροφή ', "nutrition") is a symbiosis, symbiotic relationship between certain kinds of plants and fungi, in which the plant gets a ...
, getting its food through
parasitism Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted structurally to this way of lif ...
upon
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
rather than
photosynthesis Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored i ...
. These fungi form a
mycorrhiza   A mycorrhiza (from Greek μύκης ', "fungus", and ῥίζα ', "root"; pl. mycorrhizae, mycorrhiza or mycorrhizas) is a symbiotic association between a fungus and a plant. The term mycorrhiza refers to the role of the fungus in the plant ...
with nearby tree species. Plants are fleshy and grow 10–35 cm tall. True stems are nonexistent. Instead, the only part which emerges from the soil are unbranched adventitious
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s which are developmentally similar to
adventitious roots Important structures in plant development are buds, shoots, roots, leaves, and flowers; plants produce these tissues and structures throughout their life from meristems located at the tips of organs, or between mature tissues. Thus, a living plant ...
. All parts of the plant are pale yellowish white to reddish-tinged. The
bract In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
s are 5–10 mm long scale-like structures, which cover most of the inflorescence. Plants flower from April to December depending on the geographic region (May to October in North America). The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s are
pendulous This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
when young, but become erect when they begin to mature into the
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
which is a capsule. The flowers are 9–12 mm long and produced in a cluster of 1–11 together at the apex of the inflorescence, which is a
raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
. It flowers between early summer and mid autumn; plants that flower in summer are yellow and sparsely hairy, while those that flower in autumn are red and densely hairy. These two color "forms" overlap in flowering time. It has been suggested that yellow individuals are largely self-pollinating. The species name is from
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
ized
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
''hypo-'', "under", and ''pitys'', "
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...
", referring to where pinesap often grows. However,
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
misspelled it ''hypopithys''. Many authorities have followed his spelling. On the other hand, it has been stated that the spelling "is generally treated as correctable to 'hypopitys'", as followed by other authorities. The variability of the species in colour, hairiness, and in the number of flowers, has led to an extensive
synonymy A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
for the species. The entire list is over 80 synonyms; some of which include ''Hypopitys americana'', ''H. fimbriata'', ''H. insignata'', ''H. lanuginosa'', ''H. latisquama'', ''H. monotropa'', ''Monotropa chinensis'', ''M. hypophegea'', ''M. lanuginosa'', ''M. latisquama'', ''M. multiflora'' and ''M. taiwaniana''. In California, ''Monotropa hypopitys'' may be confused for the much less common ''
Pityopus ''Pityopus'' is a monotypic genus of flowering plants in the family Ericaceae containing the single species ''Pityopus californicus'', which is known by the common name pinefoot. Distribution The plant is native to the mountains of the West Coas ...
'' or ''
Hemitomes ''Hemitomes'' is a monotypic genus of plants containing the single species ''Hemitomes congestum'', which is known as gnome plant and cone plant. This rare and unusual plant is native to the west coast of North America from British Columbia to Ca ...
'' which are superficially similar.


References


External links

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Monotropa hypopitys
{{Taxonbar, from=Q164043 Monotropoideae Parasitic plants Flora of Europe Flora of Asia Flora of North America Plants described in 1753 Taxa named by Carl Linnaeus