Phyllodocida is an order of

polychaete Polychaeta () is a paraphyletic class (biology), class of generally marine invertebrate, marine annelid worms, common name, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that ...

worms in the subclass

Aciculata Errantia is a diverse group of marine polychaete worms in the phylum Annelida. Traditionally a subclass of the paraphyletic class "Polychaeta", it is currently regarded as a monophyletic group within the larger Pleistoannelida, composed of Err ...

.World Register of Marine Species
/ref> These worms are mostly marine, though some are found in

brackish water Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estua ...

. Most are active

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...

creatures, moving over the surface or burrowing in sediments, or living in cracks and crevices in bedrock. A few construct tubes in which they live and some are

pelagic The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or wa ...

, swimming through the

water column A water column is a conceptual column of water from the surface of a sea, river or lake to the bottom sediment.Munson, B.H., Axler, R., Hagley C., Host G., Merrick G., Richards C. (2004).Glossary. ''Water on the Web''. University of Minnesota-D ...

. There are estimated to be more than 4,600 accepted

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

in the order.

Characteristics

Phyllodocida are segmented worms and range in size from a few millimetres long to over a metre. Each segment bears a pair of paddle-like

parapodia In invertebrates, the term parapodium ( Gr. ''para'', beyond or beside + ''podia'', feet; plural: parapodia) refers to lateral outgrowths or protrusions from the body. Parapodia are predominantly found in annelids, where they are paired, unjointed ...

. The

prostomium The prostomium (From Ancient Greek, meaning "before the mouth"; plural: prostomia; sometimes also called the "acron") is the cephalized first body segment in an annelid worm's body at the anterior end. It is in front of (but does not include) th ...

generally has one or two pairs of eyes, a dorsal pair of antennae, a ventral pair of sensory

palps Pedipalps (commonly shortened to palps or palpi) are the second pair of appendages of chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to the chelicerae ("jaws") and ...

and a pair of organs on the neck. The

peristomium The peristomium is the first true body segment in an annelid worm's body in the anterior end. It is directly behind the prostomium and contains the mouth, tentacular cirri, and sometimes feeding palps, which may instead occur on the prostomium. If ...

is a ring, often hidden dorsally by the prostomium and the first segment. There is a muscular

proboscis A proboscis () is an elongated appendage from the head of an animal, either a vertebrate or an invertebrate. In invertebrates, the term usually refers to tubular mouthparts used for feeding and sucking. In vertebrates, a proboscis is an elong ...

with one or more pairs of jaws. The next few segments tend to differ from those further back in having enlarged dorsal and ventral cirri (fine appendages) and reduced parapodial lobes and

chaeta A chaeta or cheta (from Greek χαίτη “crest, mane, flowing hair"; plural: chaetae) is a chitinous bristle or seta found in annelid worms, (although the term is also frequently used to describe similar structures in other invertebrates suc ...

e (bristles). Some species have appendages with specialised functions but most have many segments that are similar to each other but which vary in size and shape along the length of the body without abrupt changes in the chaetae and parapodia from one to the next.Tree of Life Web Project
/ref>

Biology

Worms in this order are generally

predators Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...

scavenger Scavengers are animals that consume dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a herbivorous feeding b ...

Phylogenetic relationships

The three main subgroups are Aphroditiformia, Glyceriformia and Nereidiformia. The Aphroditiformia are characterised by the presence of

elytra An elytron (; ; , ) is a modified, hardened forewing of beetles (Coleoptera), though a few of the true bugs (Hemiptera) such as the family Schizopteridae are extremely similar; in true bugs, the forewings are called hemelytra (sometimes alterna ...

e or scales on alternating segments. The Glyceriformia are characterised by the presence of unique cone-shaped and ringed prostomiums. The Nereidiformia are more problematic, having no universally distinguishing common features. There is agreement on the

monophyly In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic grou ...

of a group including the Hesionidae, Nereididae and Chrysopetalidae families but opinions differ about Pilargidae, and molecular and morphological studies continue.Dahlgren, T.G., Lundberg, J., Pleijel, F., and Sundberg, P. 2000. Morphological and molecular evidence of the phylogeny of Nereidiform polychaetes (Annelida). J. zool. Syst. evol. Res. 38:249-253.

Families

Recognised families: Suborder Aphroditiformia :Superfamily Aphroditoidea :: Acoetidae :: Aphroditidae :: Eulepethidae ::

Pholoidae Pholoidae is a family of polychaetes belonging to the order Phyllodocida. Genera: * '' Imajimapholoe'' Pettibone, 1992 * '' Laubierpholoe'' Pettibone, 1992 * '' Metaxypsamma'' Wolf, 1986 * ''Pholoe Foloi ( el, Φολόη, Latin: ''Pholoe'') is a ...

Polynoidae Polynoidae is a family of marine Polychaete worms known as "scale worms" due to the scale-like elytra on the dorsal surface. Almost 900 species are currently recognised belonging to 9 subfamilies and 167 genera. They are active hunters, but gene ...

Sigalionidae Sigalionidae is a family of polychaetes belonging to the order Phyllodocida Phyllodocida is an order of polychaete worms in the subclass Aciculata. These worms are mostly marine, though some are found in brackish water. Most are active benth ...

:Superfamily Chrysopetalacea :: Chrysopetalidae :Superfamily Pisionacea ::

Pisionidae Sigalionidae is a family of polychaetes belonging to the order Phyllodocida Phyllodocida is an order of polychaete worms in the subclass Aciculata. These worms are mostly marine, though some are found in brackish water. Most are active benth ...

Suborder Glyceriformia ::

Glyceridae Glyceridae is a family of polychaete worms. They are commonly referred to as beak-thrower worms or bloodworms. They are bright red, segmented, aquatic worms. The proboscis worm '' Glycera'' is sometimes called bloodworm. The Glyceridae are epi- a ...

Goniadidae Goniadidae is a family of marine worms within the Polychaeta. Goniadids have long, slender bodies and can grow up to 260mm in length, although most are less than 50mm long. Goniadids have an eversible proboscis with a circlet of chitinous jaws ...

Lacydoniidae Lacydoniidae is a family of polychaetes belonging to the order Phyllodocida. Genera: * '' Lacydonia'' Marion, 1874 References Polychaetes {{Annelid-stub ...

Paralacydoniidae Paralacydoniidae is a family of polychaetes belonging to the order Phyllodocida Phyllodocida is an order of polychaete worms in the subclass Aciculata. These worms are mostly marine, though some are found in brackish water. Most are active ben ...

Suborder Nereidiformia :: Antonbruuniidae ::

Hesionidae Hesionidae are a family of phyllodocid "bristle worms" (class Polychaeta). They are (like almost all polychaetes) marine organisms. Most are found on the continental shelf; '' Hesiocaeca methanicola'' is found on methane ice, where it feeds ...

Nereidae Nereididae (formerly spelled Nereidae) are a family of polychaete worms. It contains about 500 – mostly marine – species grouped into 42 genera. They may be commonly called ragworms or clam worms. Characteristics The prostomium of ...

Pilargidae Pilargidae is a family of polychaetes. These marine worms are cylindrical, somewhat flattened, and can be ribbon-like. They can be found free-living on sediment, or shallowly in sediment. Some species within the genera Hermundura and Litocorsa ...

Syllidae Syllidae is a family of small to medium-sized polychaete worms. Syllids are distinguished from other polychaetes by the presence of a muscular region of the anterior digestive tract known as the ''proventricle''. Syllid worms range in size from ...

Suborder

Phyllodocida Phyllodocida is an order of polychaete worms in the subclass Aciculata. These worms are mostly marine, though some are found in brackish water. Most are active benthic creatures, moving over the surface or burrowing in sediments, or living in cr ...

incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...

Iospilidae Iospilidae is a family of polychaetes belonging to the order Phyllodocida. Genera: * ''Iospilopsis'' Augener, 1922 * ''Iospilus'' Viguier, 1886 * ''Paraiospilus'' Viguier, 1911 * ''Phalacrophorus'' Greeff, 1879 References

{{Taxonbar, from= ...

:: Nephtyidae :: Sphaerodoridae ::

Tomopteridae Tomopteridae is a family of holopelagic polychaetes belonging to the order Phyllodocida. The genus Tomopteris consist of about 70 species, and the three other genera of one known species each. Length varies from just 1-2 cm to 30 cm. These are ver ...

:: Typhloscolecidae ::

Yndolaciidae Yndolaciidae is a family of polychaetes belonging to the order Phyllodocida Phyllodocida is an order of polychaete worms in the subclass Aciculata. These worms are mostly marine, though some are found in brackish water. Most are active benthi ...

Suborder Phyllodociformia :: Alciopidae ::

Lopadorhynchidae Lopadorrhynchidae is a family of polychaete worms. References External links * Lopadorrhynchidaeat the World Register of Marine Species The World Register of Marine Species (WoRMS) is a taxonomic database that aims to provide an autho ...

Phyllodocidae Phyllodocidae is a family of polychaete worms. Worms in this family live on the seabed and may burrow under the sediment. Characteristics Members of the Phyllodocidae are characterised by an eversible pharynx and leaf-like dorsal cirri. The hea ...

Pontodoridae Pontodoridae is a family of polychaetes belonging to the order Phyllodocida Phyllodocida is an order of polychaete worms in the subclass Aciculata. These worms are mostly marine, though some are found in brackish water. Most are active benthic ...

Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...

Nautiliniellidae Nautiliniellidae is a family of polychaetes belonging to the order Phyllodocida. Genera: * ''lascarpia'' * ''Iheyomytilidicola'' * ''Laubierus'' References

{{Taxonbar, from=Q5180094 Phyllodocida ...

References

External links

{{Taxonbar, from=Q733080 Protostome orders Annelid orders