Philochortus
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''Philochortus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of lizards of the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Lacertidae The Lacertidae are the family (biology), family of the wall lizards, true lizards, or sometimes simply lacertas, which are native to Afro-Eurasia. It is a diverse family with at least 300 species in 39 genera. They represent the dominant group o ...
.
Species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of this genus are distributed in
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Mediter ...
,
Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...
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Libya Libya (; ar, ليبيا, Lībiyā), officially the State of Libya ( ar, دولة ليبيا, Dawlat Lībiyā), is a country in the Maghreb region in North Africa. It is bordered by the Mediterranean Sea to the north, Egypt to Egypt–Libya bo ...
,
Mali Mali (; ), officially the Republic of Mali,, , ff, 𞤈𞤫𞤲𞥆𞤣𞤢𞥄𞤲𞤣𞤭 𞤃𞤢𞥄𞤤𞤭, Renndaandi Maali, italics=no, ar, جمهورية مالي, Jumhūriyyāt Mālī is a landlocked country in West Africa. Mali ...
,
Niger ) , official_languages = , languages_type = National languagesEthiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
,
Djibouti Djibouti, ar, جيبوتي ', french: link=no, Djibouti, so, Jabuuti officially the Republic of Djibouti, is a country in the Horn of Africa, bordered by Somalia to the south, Ethiopia to the southwest, Eritrea in the north, and the Red ...
,
Eritrea Eritrea ( ; ti, ኤርትራ, Ertra, ; ar, إرتريا, ʾIritriyā), officially the State of Eritrea, is a country in the Horn of Africa region of Eastern Africa, with its capital and largest city at Asmara. It is bordered by Ethiopia ...
,
Somalia Somalia, , Osmanya script: 𐒈𐒝𐒑𐒛𐒐𐒘𐒕𐒖; ar, الصومال, aṣ-Ṣūmāl officially the Federal Republic of SomaliaThe ''Federal Republic of Somalia'' is the country's name per Article 1 of thProvisional Constituti ...
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Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
,
Yemen Yemen (; ar, ٱلْيَمَن, al-Yaman), officially the Republic of Yemen,, ) is a country in Western Asia. It is situated on the southern end of the Arabian Peninsula, and borders Saudi Arabia to the Saudi Arabia–Yemen border, north and ...
, and
Saudi Arabia Saudi Arabia, officially the Kingdom of Saudi Arabia (KSA), is a country in Western Asia. It covers the bulk of the Arabian Peninsula, and has a land area of about , making it the fifth-largest country in Asia, the second-largest in the A ...
.


Etymology

''Philochortus'' means grass-loving (Greek: ''philos'' = friend, ''chortos'' = grass). The common name of ''Philochortus'' species is therefore grass-loving lizards, or shield-backed ground lizards because of their typical back scalation and terrestrial habitat.


Diagnosis

Species of ''Philochortus'' are medium to large-sized lacertids with long cylindrical tails. The unregenerated tail is up to 3.25 times longer than head and body. Eyes with movable lids. The
nostril A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called turbi ...
is pierced between two shields and usually bordered by the first
supralabial In reptiles, the supralabial scales, also called upper-labials, are those scales that border the mouth opening along the upper jaw. They do not include the median scaleWright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates ( ...
or narrowly separated from it. The collar is well marked. The ventral plates are smooth, feebly imbricate and arranged in 6 longitudinal series. The dorsal scales are smooth or keeled. Back with 2 to 6 longitudinal series of enlarged plate-like scales along the dorsal mid-line. This is the main difference to ''
Latastia ''Latastia'' is a genus of lizards of the family Lacertidae. Species of this genus are distributed in Africa (Egypt, Ethiopia, Djibouti, Eritrea, Guinea-Bissau, Cameroon, Kenya, Malawi, Mali, Mozambique, Niger, Zambia, Senegal, Zimbabwe, Soma ...
''. Femoral pores are present on the inside of the thighs, more prominent in males. Tail base in males much broader than in females.Boulenger 1921


Habitat and natural history

The natural history of all
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of ''Philochortus'' is only poorly known. Many of the species inhabit
semiarid A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of semi-ar ...
,
arid A region is arid when it severely lacks available water, to the extent of hindering or preventing the growth and development of plant and animal life. Regions with arid climates tend to lack vegetation and are called xeric or desertic. Most ar ...
or
hyperarid An aridity index (AI) is a numerical indicator of the degree of dryness of the climate at a given location. A number of aridity indices have been proposed (see below); these indicators serve to identify, locate or delimit regions that suffer from ...
regions, mountainous areas as well as plains from sea level up to 1500 m altitude (''P. neumanni'' ). Most species prefer sparsely vegetated, open and sandy or fairly rocky localities (''P. neumanni'' ) and can be found in ''
Acacia ''Acacia'', commonly known as the wattles or acacias, is a large genus of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa and Australasia. The genus na ...
''-''
Commiphora The genus of the myrrhs, ''Commiphora'', is the most species-rich genus of flowering plants in the frankincense and myrrh family, Burseraceae. The genus contains approximately 190 species of shrubs and trees, which are distributed throughout th ...
'' deciduous bushland and
semidesert A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of semi-ar ...
bushland vegetation types. But ''P. zolii'' and ''P. neumanni'' can be found also in marginal unattended cultivated areas with ''
Desmostachya bipinnata ''Desmostachya bipinnata'', commonly known as halfa grass, big cordgrass, and salt reed-grass, is an Old World perennial grass, long known and used in human history. The grass is tall, tufted, leafy, perennial grass, branching from the base, erec ...
'' grass and other scrub and well vegetated natural habitats.Baha El Din 2006 The species of ''Philochortus'' are diurnal, sun-loving active predators on
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s and other
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s, egg-laying and terrestrial but ''P. zolii'' is an avid climber. The long stiff tail is aiding in balance on vegetation. They dig burrows in the soil below clumps of grasses or small bushes.


Species

The following seven
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
are recognized as being valid."''Philochortus'' ". The Reptile Database. www.reptile-database.org. *'' Philochortus hardeggeri'' - Hardegger's orangetail lizard, Hardegger's shield-backed lizard *'' Philochortus intermedius'' - southern orangetail lizard, Boulenger's shield-backed lizard *'' Philochortus neumanni'' - Neumann's orangetail lizard *'' Philochortus phillipsi'' - Phillips's shield-backed lizard *'' Philochortus rudolfensis'' - southern shield-backed lizard *'' Philochortus spinalis'' - Peters's shield-backed lizard, Eritrea orangetail lizard *''
Philochortus zolii ''Philochortus zolii'' is a species of lizard in the family Lacertidae. The species is native to northern Africa. Taxonomy Trape ''et al''. (2012) considered West African ''P. lhotei'' a junior synonym of ''P. zolii'' as its morphological chara ...
'' ''
Nota bene (, or ; plural form ) is a Latin phrase meaning "note well". It is often abbreviated as NB, n.b., or with the ligature and first appeared in English writing . In Modern English, it is used, particularly in legal papers, to draw the atten ...
'': A
binomial authority In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
in parentheses indicates that the species was originally described in a genus other than ''Philochortus''.


References


Further reading

*Baha El Din S (2006). ''A Guide to the Reptiles and Amphibians of Egypt''. Cairo and New York: The American University in Cairo Press. * Boulenger GA (1898)
"On a Second Collection of Reptiles made by Mr. E. Lort-Phillips in Somaliland"
''The Annals and Magazine of Natural History, Seventh Series'' 2: 130-133. (''Latastia phillipsii'', new species, pp. 131–132). *Boulenger GA (1917)
"On the lizards of the genus ''Philochortus'' Matschie"
''Proceedings of the Zoological Society of London'' 1917: 145-157 + Plates I-II.
''Philochortus intermedius''
new species, pp. 152–154 + Plate II, figures 3, 3a, 3b, 3c, 4). *Boulenger GA (1921)
''Monograph of the Lacertidae. Volume II''
London: Trustees of the
British Museum (Natural History) The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum an ...
. viii + 451 pp. (Genus ''Philochortus'', pp. 1-2; species of ''Philochortus'', pp. 3-14). *Largen MJ, Spawls S (2006). "Lizards of Ethiopia (Reptilia Sauria): an annotated checklist, bibliography, gazetteer and identification key". ''Tropical Zoology'' 19: 21-109. * Matschie P (1893)
"''Über einige von Herrn Oscar Neumann bei Aden gesammelte u. beobachtete Säugethiere, Reptilien und Amphibien'' "
''Sitzungs-Berichte der Gesellschaft Naturforschender Freunde zu Berlin'' 1893: 24-31. (''Philochortus'', new genus, p. 30; ''P. neumanni'', new species, pp. 30–31). (in
German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ger ...
). * Parker HW (1932)
"Scientific results of the Cambridge Expedition to the East African Lakes, 1930-1.—5. Reptiles and Amphibians"
''The Journal of the Linnean Society of London. Zoology'' 38: 213-229. (''Philocortus intermedius rudolfensis'', new subspecies, p. 226). *Parker HW (1942)
"The Lizards of British Somaliland"
''Bulletin of the
Museum of Comparative Zoology A museum ( ; plural museums or, rarely, musea) is a building or institution that cares for and displays a collection of artifacts and other objects of artistic, cultural, historical, or scientific importance. Many public museums make these ...
at Harvard College'' 91 (1): 1-101. * Peters W (1874)
"''Über einige neue Reptilien'' (Lacerta, Eremias, Diploglossus, Euprepes, Lygosoma, Sepsina, Ablepharus, Simotes, Onychocephalus)"
''Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin'' 1874: 368-377 + Plate. (''Lacerta spinalis'', new species, pp. 369–370 + Plate, figure 2). (in German). *Schleich HH, Kästle W, Kabisch K (1996). ''Amphibians and Reptiles of North Africa''. Königstein: Koeltz Scientific Books. 630 pp. . * Scortecci G (1934)
"''Descrizione preliminare di una nuova specie del genere'' Philochortus (Philochortus zolii) ''della zona Gat (Missione della Reale Società Geografica)''"
''Atti Soc. Italiana Sci. Nat.'' 73: 305-308. (''
Philochortus zolii ''Philochortus zolii'' is a species of lizard in the family Lacertidae. The species is native to northern Africa. Taxonomy Trape ''et al''. (2012) considered West African ''P. lhotei'' a junior synonym of ''P. zolii'' as its morphological chara ...
'', new species). (in
Italian Italian(s) may refer to: * Anything of, from, or related to the people of Italy over the centuries ** Italians, an ethnic group or simply a citizen of the Italian Republic or Italian Kingdom ** Italian language, a Romance language *** Regional Ita ...
). *Spawls S, Howell KM, Drewes RC, Ashe J (2002). ''A Field Guide to the Reptiles of East Africa''. San Diego, San Francisco, New York, Boston, London: Academic Press, Elsevier Science. 544 pp. * Steindachner F (1891)
"''Ueber neue und seltene Lacertiden aus der herpetologischen Sammlung des k. k. naturhistorischen Hofmuseums'' "
''Annalen des Naturhistorischen Museums in Wien'' 6: 371-378 + Plates XI-XII. (''Latastia hardeggeri'', new species, pp. 371–373 + Plate XI, figures 1-3). (in German). {{Taxonbar, from=Q2070539 Lizard genera Taxa named by Paul Matschie