Pasiphila Muscosata
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''Pasiphila muscosata'', the emerald pug moth, is a
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
in the family
Geometridae The geometer moths are moths belonging to the family Geometridae of the insect order Lepidoptera, the moths and butterflies. Their scientific name derives from the Ancient Greek ''geo'' γεω (derivative form of or "the earth"), and ''met ...
. It is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
and has been found in the
North North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating Direction (geometry), direction or geography. Etymology T ...
,
South South is one of the cardinal directions or Points of the compass, compass points. The direction is the opposite of north and is perpendicular to both east and west. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Pro ...
and
Stewart Islands Sikaiana (formerly called the Stewart Islands) is a small atoll NE of Malaita in Solomon Islands in the south Pacific Ocean. It is almost in length and its lagoon, known as Te Moana, is totally enclosed by the coral reef. Its total land s ...
. The larvae of this species are known to be present in January and live on ''
Muehlenbeckia ''Muehlenbeckia'' or maidenhair is a genus of flowering plants in the family Polygonaceae. It is native to the borders of the Pacific, including South and North America, Papua New Guinea and Australasia. It has been introduced elsewhere, includin ...
'' species including ''
Muehlenbeckia australis ''Muehlenbeckia australis'', the large-leaved muehlenbeckia or pohuehue, is a prostrate or climbing plant native to New Zealand. Description The species grows up to tall with grey bark. The leaves are on stiff petioles and are long. Lamina is ...
.'' The larvae tends to be brown but it is variable in both colour and markings. The larvae of this species pupates in a loose cocoon on the ground and adult moths emerge in September. Adults are commonly on the wing until the following May. Although adults tend to be a deep emerald green colour this species is again variable and there is also an orange-yellow variety as well as intermediate forms. Adults are attracted to light and often rest during the day on tree trunks.


Taxonomy

This species was first described by Francis Walker in 1862 and named ''Eupithecia muscosata''. Walker used a specimen collected by T. R. Oxley in
Nelson Nelson may refer to: Arts and entertainment * ''Nelson'' (1918 film), a historical film directed by Maurice Elvey * ''Nelson'' (1926 film), a historical film directed by Walter Summers * ''Nelson'' (opera), an opera by Lennox Berkeley to a lib ...
. In 1868
Achille Guenée Achille Guenée (sometimes M.A. Guenée; 1 January 1809 – 30 December 1880) was a French lawyer and entomologist. Biography Achille Guenée was born in Chartres and died in Châteaudun. He was educated in Chartres, where he showed a very earl ...
, thinking he was describing a new species, named this species ''Eupithecia cidariaria''. In 1875
Cajetan von Felder Baron Cajetan von Felder (german: link=no, Cajetan Freiherr von Felder; 19 September 1814 – 30 November 1894) was an Austrian lawyer, entomologist and liberal politician. He served as mayor of Vienna from 1868 to 1878. Life and career Felder ...
and
Alois Friedrich Rogenhofer Alois Friedrich Rogenhofer (22 December 1831, in Vienna – 15 January 1897, in Vienna) was an Austrian entomologist. He was a curator at the Naturhistorisches Museum in Vienna, where he was the first keeper of the Lepidoptera. Rogenhofer was ma ...
, also thinking they were describing a new species, named this moth ''Cidaria aquossata''. In 1888
Edward Meyrick Edward Meyrick (25 November 1854, in Ramsbury – 31 March 1938, at Thornhanger, Marlborough) was an English schoolmaster and amateur entomologist. He was an expert on microlepidoptera and some consider him one of the founders of modern m ...
placed this species in the genus ''Pasiphila'' and synonymised the names ''Eupithecia cidariaria'' and ''Cidaria aquosata''. In 1898
George Hudson George Hudson (probably 10 March 1800 – 14 December 1871) was an English railway financier and politician who, because he controlled a significant part of the railway network in the 1840s, became known as "The Railway King"—a title conferr ...
discussed this species as a synonym of ''Chloroclystis bilineolata''. In 1928 Hudson again discussed as well as illustrated this species under the name ''Chloroclystis muscosata''. In 1971 John S. Dugdale placed this species again in the genus ''Pasiphila''. In 1988 John S. Dugdale discussed this species under the name ''Pasiphila muscosata'' and in 2010 Robert Hoare in the New Zealand Inventory of Biodiversity followed this placement. The male
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen is held at the
Natural History Museum, London The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum an ...
.


Description

Walker originally described the adult male of this species as follows: This species is variable in appearance with Hudson describing a variety of this moth which has a ground colour of orange-yellow. He also stated that intermediate forms could be found but that these were much rarer than the more typical green form or the orange-yellow variety. Hudson described the larva of this species as follows: The larvae of this species, similar to the adults, is also variable in colour and markings. Hudson hypothesised that the appearance of the larvae imitates the dried flowers of its host plants.


Distribution

This species is endemic to New Zealand and can be found in the North, South and Stewart Islands.


Life cycle and behaviour

From late December to mid January adults lay their oval, flattened eggs, usually singly, on the leaves of their host species. These eggs hatch after about fifteen days. They turn a dark colour in the last few days of maturing. The larvae of this species tend to be present in January. They are known to rest with their body held in a twisted shape and when walking have been observed to sway from side to side. This species pupates in a loose cocoon on the ground. There appears to be only one brood per year for this species. The adults of this species are on the wing all year round but more commonly from September until May. They are attracted to light. They have been observed resting during the day on tree trunks where their appearance resembles moss making the species extremely difficult to find.


Habitat and hosts

Larvae of this species live on species in the genus ''
Muehlenbeckia ''Muehlenbeckia'' or maidenhair is a genus of flowering plants in the family Polygonaceae. It is native to the borders of the Pacific, including South and North America, Papua New Guinea and Australasia. It has been introduced elsewhere, includin ...
'' as well as on '' Lophomyrtus bullata''. They have been successfully reared in captivity on ''
Muehlenbeckia australis ''Muehlenbeckia australis'', the large-leaved muehlenbeckia or pohuehue, is a prostrate or climbing plant native to New Zealand. Description The species grows up to tall with grey bark. The leaves are on stiff petioles and are long. Lamina is ...
''.


References

{{Taxonbar , from=Q13559847 Moths described in 1862 muscosata Moths of New Zealand Endemic fauna of New Zealand Taxa named by Francis Walker (entomologist) Endemic moths of New Zealand