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The paradise fish, paradise-fish, paradisefish, or paradise gourami (''Macropodus opercularis'') is a species of
gourami Gouramis, or gouramies , are a group of freshwater anabantiform fishes that comprise the family Osphronemidae. The fish are native to Asia—from the Indian Subcontinent to Southeast Asia and northeasterly towards Korea. The name "gourami", of ...
found in most types of fresh water in
East Asia East Asia is the eastern region of Asia, which is defined in both geographical and ethno-cultural terms. The modern states of East Asia include China, Japan, Mongolia, North Korea, South Korea, and Taiwan. China, North Korea, South Korea and ...
, ranging from the
Korean Peninsula Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic o ...
to northern
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
. This species can reach a
standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...
of , though most are only about . Paradise gouramis were one of the first
ornamental fish Lists of aquarium life include lists of fish, amphibians, invertebrates and plants in freshwater, brackish and marine aquariums. In fishkeeping, suitable species of aquarium fish, plants and other organisms vary with the size, water chemistry and ...
available to western
aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
keepers, having been imported 1869 to France by the French aquarium fish importer
Pierre Carbonnier Pierre Carbonnier, (7 August 1828 – 8 April 1883) was a French scientist, ichthyologist, fish breeder and public Aquarium director. Member of Imperial Society of acclimatization (Société Impériale d'Acclimatation). Biography Pierre Carb ...
in
Paris Paris () is the capital and most populous city of France, with an estimated population of 2,165,423 residents in 2019 in an area of more than 105 km² (41 sq mi), making it the 30th most densely populated city in the world in 2020. S ...
. The paradise fish is one of the more aggressive members of its family. It is more aggressive than the
three spot gourami The three spot gourami (''Trichopodus trichopterus''), also known as the opaline gourami, blue gourami, and gold gourami, is a species of fish native to southeastern Asia, but also introduced elsewhere. This gourami gets its name from the two sp ...
, yet less pugnacious in nature than the less commonly kept
combtail ''Belontia'', sometimes referred to combtail gouramies, is a genus of gouramis, the only genus within the subfamily Belontiinae, native to freshwater habitats in Southeast Asia and Sri Lanka. These are medium sized to large gouramies that are sel ...
.


Behaviour

Paradise fish are fairly combative, harassing and attacking each other, as well as potentially assaulting and killing small fish. During a fight, the paradise fish will often change its color, usually displaying dark blue lateral lines on the sides of their bodies; extend its fins; and spread out its operculum. Paradise fish are more likely to show aggressive behavior towards other paradise fish than to fish of a different species. Acts of aggression tend to increase as the distance to the fish's home in Asia increases. In the wild, they are
predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
s, eating
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s,
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s, and
fish fry A fish fry is a meal containing battered or breaded fried fish. It usually also includes french fries, coleslaw, macaroni salad, lemon slices, tartar sauce, hot sauce, malt vinegar and dessert. Some Native American versions are cooked by coati ...
. The popularity of this species has waned in recent decades as much more colorful (and often less abusive) species of
gourami Gouramis, or gouramies , are a group of freshwater anabantiform fishes that comprise the family Osphronemidae. The fish are native to Asia—from the Indian Subcontinent to Southeast Asia and northeasterly towards Korea. The name "gourami", of ...
s have become widely available to hobbyists. This species is one of the few fish that can change its color (lighter or darker) in response to stimuli. It also appears that paradise fish are capable of learning through a type of restrictive process. Most forms of active teaching seem to hinder the paradise fish's ability to learn the movements of its owner via careful observation.


Habitat and diet

Paradise fish are tolerant of a wide range of water conditions, surviving in cool and warm waters alike. In the wild, they are most commonly found in shallow water containing dense vegetation, such as a marsh or rice field. However they can be kept in outdoor
pond A pond is an area filled with water, either natural or artificial, that is smaller than a lake. Defining them to be less than in area, less than deep, and with less than 30% emergent vegetation helps in distinguishing their ecology from th ...
s, or even the simplest of unheated aquaria. They will accept virtually any food, but should be given a reasonably high-protein diet (as opposed to vegetable-based foods.) They also eat mosquito larvae, black worms,
brine shrimp ''Artemia'' is a genus of aquatic crustaceans also known as brine shrimp. It is the only genus in the family Artemiidae. The first historical record of the existence of ''Artemia'' dates back to the first half of the 10th century AD from Urmia La ...
, and small flies.


Disease control

In Taiwan, the native populations of paradise fish have been reduced to low levels by pollution in the rivers, and are now listed as a threatened species. The local population of yellow fever mosquitoes (''
Aedes aegypti ''Aedes aegypti'', the yellow fever mosquito, is a mosquito that can spread dengue fever, chikungunya, Zika fever, Mayaro and yellow fever viruses, and other disease agents. The mosquito can be recognized by black and white markings on its legs ...
'') has since increased in the absence of one of its main predators. The infection rate for
dengue fever Dengue fever is a mosquito-borne tropical disease caused by the dengue virus. Symptoms typically begin three to fourteen days after infection. These may include a high fever, headache, vomiting, muscle and joint pains, and a characterist ...
has subsequently increased in the human population, caused in part by to the lack of natural mosquito predators. Paradise fish are also considered to be an ideal subject for behavioral genetic studies and have been used to study
Iridoviridae ''Iridoviridae'' is a family of viruses with double-stranded DNA genomes. Amphibians, fish, and invertebrates such as arthropods serve as natural hosts. There are currently 22 species in this family, divided among two subfamilies and seven gener ...
type viruses.Xu, Liwen, Juan Feng, and Youhua Huang. "Identification Of Lymphocystis Disease Virus From Paradise Fish Macropodus Opercularis (LCDV-PF)." Archives Of Virology 159.9 (2014): 2445–2449. MEDLINE. Web. 19 Feb. 2015.


In home aquaria

Male paradise fish should be kept apart, since they will fight aggressively by locking jaws. A male can be kept with females; females may also be kept together in groups. A tank that includes paradise fish should be at least in size for a single male or for a community tank. The tank should be well planted and covered; bogwood and rockwork may be included. Paradise fish tankmates must be chosen with care. Suitable ones include
giant danio The giant danio (''Devario aequipinnatus'') is a tropical fish belonging to the minnow family Cyprinidae. Originating in Sri Lanka, Nepal, and the west coast of India, this species grows to a maximum length of 4-6 inches (10-15 cm), making i ...
s, large
tetra Terrestrial Trunked Radio (TETRA; formerly known as Trans-European Trunked Radio), a European standard for a trunked radio system, is a professional mobile radio and two-way transceiver specification. TETRA was specifically designed for use by go ...
s, most smaller
catfish Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive, ...
es, and even some of the less aggressive
cichlid Cichlids are fish from the family Cichlidae in the order Cichliformes. Cichlids were traditionally classed in a suborder, the Labroidei, along with the wrasses ( Labridae), in the order Perciformes, but molecular studies have contradicted this ...
s, such as
firemouth cichlid The firemouth cichlid (''Thorichthys meeki'') is a species of cichlid fish native to Central America. They occur in rivers of the Yucatán Peninsula, Mexico, south through Belize and into northern Guatemala. Their natural habitat is typically sh ...
s. Slow-moving or long-finned fish such as fancy goldfish and
freshwater angelfish ''Pterophyllum'' is a small genus of freshwater fish from the family Cichlidae known to most aquarists as angelfish. All ''Pterophyllum'' species originate from the Amazon Basin, Orinoco Basin and various rivers in the Guiana Shield in tropical ...
are likely to be attacked by males;
betta ''Betta'' is a large genus of small, active, often colorful, freshwater ray-finned fishes, in the gourami family (Osphronemidae). The best known ''Betta'' species is ''B. splendens,'' commonly known as the Siamese fighting fish and often kept ...
s and
gourami Gouramis, or gouramies , are a group of freshwater anabantiform fishes that comprise the family Osphronemidae. The fish are native to Asia—from the Indian Subcontinent to Southeast Asia and northeasterly towards Korea. The name "gourami", of ...
s may also be victimized due to their resemblance to paradise fish. Male paradise fish may also attempt to court female bettas and gouramis. Fish less than are likely to be consumed violently. If kept with significantly larger but non-aggressive fish, such as ''
Geophagus ''Geophagus'' is a genus of cichlids that mainly live in South America as far south as Argentina and Uruguay, but a single species, ''G. crassilabris'' is from Panama.Mattos, J.L.O., Costa, W.J.E.M. & Santos, A.C.A. (2015): ''Geophagus diamantin ...
'' cichlids, large ''
Synodontis ''Synodontis'' is the largest genus of mochokid catfishes. It is the biggest genus within the 10 genera and 190 different species in the family Mochokidae. ''Synodontis'' has over 131 different species within the genus. ''Synodontis'' are also k ...
'' catfishes, or larger gouramis, they are usually submissive and do not act nearly as abusively as when they are the dominant species in the aquarium. However, they themselves can be bullied by similar sized or even smaller fish if that fish has already established territory in the tank which it is not willing to share or give up. If this is the case they will not even attempt to fight and will take to hiding behind filters, plants, or in décor, and will succumb to stress eating and diarrhoea.


Reproduction

As is typical of most bettas and gouramis,
spawning Spawn is the eggs and sperm released or deposited into water by aquatic animals. As a verb, ''to spawn'' refers to the process of releasing the eggs and sperm, and the act of both sexes is called spawning. Most aquatic animals, except for aquati ...
involves a male building a
bubble nest Bubble nests, also called foam nests, are created by some fish and frog species as floating masses of bubbles blown with an oral secretion, saliva bubbles, and occasionally aquatic plants. Fish that build and guard bubble nests are known as aph ...
(a floating mat of
saliva Saliva (commonly referred to as spit) is an extracellular fluid produced and secreted by salivary glands in the mouth. In humans, saliva is around 99% water, plus electrolytes, mucus, white blood cells, epithelial cells (from which DNA can be ...
-coated air bubbles, often incorporating plant matter) and attracting a female to it. If the female accepts the male's advances, the fish will 'embrace' in open water, releasing both eggs and sperm into the water. The male gathers the fertilized
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...
s after each embrace, spitting them up into the bubble nest. After spawning, the male may violently attack his mate or any other fish that might approach the new fertilized eggs or hatched fry, which are both a common source of food in the natural habitat. A breeder usually chooses to move the female to a separate tank to improve the chances of survival of both the female and the hatched fry. After the fry have begun to swim freely, the male's protective behavior subsides, so the breeder removes the male for the protection of the fry, and they are raised on
infusoria Infusoria are minute freshwater life forms including ciliates, euglenoids, protozoa, unicellular algae and small invertebrates. Some authors (e.g., Bütschli) used the term as a synonym for Ciliophora. In modern formal classifications, the term i ...
or newly hatched
brine shrimp ''Artemia'' is a genus of aquatic crustaceans also known as brine shrimp. It is the only genus in the family Artemiidae. The first historical record of the existence of ''Artemia'' dates back to the first half of the 10th century AD from Urmia La ...
. An
albino Albinism is the congenital absence of melanin in an animal or plant resulting in white hair, feathers, scales and skin and pink or blue eyes. Individuals with the condition are referred to as albino. Varied use and interpretation of the term ...
form of ''Macropodus opercularis'' is available. Many aquarists consider this form to be less aggressive than the wild type, but also less hardy, having more trouble with low temperatures.


References


External links

{{Taxonbar, from=Q304256 Macropodus Fish of East Asia Freshwater fish of Taiwan Fauna of Hong Kong
paradise fish The paradise fish, paradise-fish, paradisefish, or paradise gourami (''Macropodus opercularis'') is a species of gourami found in most types of fresh water in East Asia, ranging from the Korean Peninsula to northern Vietnam. This species can ...
paradise fish The paradise fish, paradise-fish, paradisefish, or paradise gourami (''Macropodus opercularis'') is a species of gourami found in most types of fresh water in East Asia, ranging from the Korean Peninsula to northern Vietnam. This species can ...