Paraclinus Fasciatus
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''Paraclinus fasciatus'', the banded blenny, is a species of labrisomid blenny native to the
Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
including the
Gulf of Mexico The Gulf of Mexico ( es, Golfo de México) is an oceanic basin, ocean basin and a marginal sea of the Atlantic Ocean, largely surrounded by the North American continent. It is bounded on the northeast, north and northwest by the Gulf Coast of ...
and the
Caribbean Sea The Caribbean Sea ( es, Mar Caribe; french: Mer des Caraïbes; ht, Lanmè Karayib; jam, Kiaribiyan Sii; nl, Caraïbische Zee; pap, Laman Karibe) is a sea of the Atlantic Ocean in the tropics of the Western Hemisphere. It is bounded by Mexico ...
from southern
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
to
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
. The banded blenny is a marine organism, meaning it lives in an aquatic environment. This species lives in the vicinity of
coral reefs A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of Colony (biology), colonies of coral polyp (zoology), polyps held together by calcium carbonate. Most coral reefs are built from stony corals, wh ...
preferring
sea grass Seagrasses are the only flowering plants which grow in marine environments. There are about 60 species of fully marine seagrasses which belong to four families (Posidoniaceae, Zosteraceae, Hydrocharitaceae and Cymodoceaceae), all in the orde ...
beds in shallow waters down to . Although they are typically located in shallow waters, their colors camouflage them exceptionally well, leading to them rarely being seen or recognized. They have also been found within floating algae. They are part of the family perciform teleost fishes within the sub order Blenniodei. The suborder blenniodei currently comprises 6 families, 151 genera, and 883 species. The perciform fish lineage is distinguished by its small body size, as well as their coastal and benthic habitats. The banded blenny has a typical lifespan of one year.


Taxonomic characteristics

''Paraclinus fasciatus'' is a species within the class
Osteichthyes Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartilage ...
, or
bony fishes Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartilage ...
. This class represents the largest class of
vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...
s, with more than 31,000 species. One of the distinguishing characteristics of this class is that the organisms have a skeleton at least partially made of bone. Another characteristic being, this class typically has cycloid or ctenoid scales in comparison to pointed
placoid A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scale (zoology), scales, which can also provide effective Underwater camouflage, camouflage through the us ...
scales. These scales are distinguished by their bony makeup and the protective layer of mucous covering them.Castro, Peter (Ph.D.), Huber, Michael (Ph.D.), Rotjan, Randi (Professor) (2019) Marine Biology: Biology 260, Boston University, USA: McGraw-Hill Education (Mcgraw-create)] The Operculum (fish), operculum is another defining characteristic of this class, it is a gill cover which, as its name implies, protects the gills with a flap. Bony fishes typically have fin rays rather than a fleshy makeup that you see in cartilaginous fishes. Often, you will also see a
swim bladder The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled Organ (anatomy), organ that contributes to the ability of many bony fish (but not cartilaginous fish) to control their buoyancy, and thus to stay at their curren ...
within bony fishes which acts as a floatation tool, allowing the fish to adjust its depth in the water.


Morphology

Banded blennies are small and slender. Their standard length is 15 mm. The color of their body ranges from different shades of brown and has thick black bar-like stripes. They have three well defined dorsal-fin spines.Lin, H., & Hastings, P. (2011). "{Evolution of a Neotropical marine fish lineage (Subfamily Chaenopsinae, Suborder Blennioidei) based on phylogenetic analysis of combined molecular and morphological data". ''Molecular Phylogenetics and Evolution'', 60(2), 236-248. Link text] They have fin-hook characteristics which are significant of the adaptations these blennies have made to their bottom-dwelling lifestyle. Adaptations of this pectoral fin has drawn great attention to the blennies from researchers. The lower part of the
pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
has a fibrous cord as well as deeply incised fin-web between neighboring rays. Banded blennies are also referred to as "
combtooth blennies Combtooth blennies are blenniiformids; percomorph marine fish of the family Blenniidae, part of the order Blenniiformes. They are the largest family of blennies with around 401 known species in 58 genera. Combtooth blennies are found in tropica ...
" because they have ctenoid scales. These are overlapping fish scales. Ctenoid scales are similar to
cycloid scale A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as w ...
s which are large and thin with a circular shape. The main difference here is that ctenoid scales have spines or comb-like spikes at their free edges.


Reproduction

Blenniids are typically sexually dimorphic. The males are larger than the females and exhibit secondary sex characteristics (SSC). The SSC are especially seen during breeding seasons, which is typically early summer resulting in a large increase in population in late Summer/early Fall. The male SSC include, an enlarged dorsal fin, glands on the anal spine, and color marks on the head or body. Banded blennys are
oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...
, meaning they release eggs as means of reproduction. There are breeding patterns observed among Blenniids, this includes males defending the breeding territory in which females come to spawn in a small hole or cavity. The territory is typically a sheltered area. The males provide the exclusive parental care and guard the eggs until they have hatched.


Habitat

The banded blennies tend to be found in
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
/
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
locations. Within these locations, they are found in shallow waters, in coral, rock, and sandy bottoms. These blennies also seem to favor sea grass beds.Gibson, R 2010 "The Biology of Blennies" ‐ Edited by R. A. Patzner, E. J. Gonçalves, P. A. Hastings and B. G. Kapoor. ''Journal of Fish Biology'', 76(7), 1871–1872 They are classified as
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
, coastal fishes, although some have been found at depths of 30m. There have also been studies conducted that correlate the population density of banded blennies to the amount of unattached red algae, also known as drift algae. The study conducted by Stoner shows that the more drift algae there is, the more blennies there also are. This floating
red algae Red algae, or Rhodophyta (, ; ), are one of the oldest groups of eukaryotic algae. The Rhodophyta also comprises one of the largest phyla of algae, containing over 7,000 currently recognized species with taxonomic revisions ongoing. The majority ...
is able to provide shelter, as well as food for the fish. These fish are also known to be exceptionally tolerant of physical and chemical conditions within the shallow waters along coastal
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
.


Feeding

Banded blennies are carnivores with highly specialized diets. Meaning, they feed on very specific species. They have teeth in their jaws to help capture prey as do most bony-fishes. Blennies tend to solely prey on amphipods when they are juveniles or below 15 mm in size. If they are above 15 mm in size their diet expands to not only amphipods, but also
shrimp Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are refer ...
,
crab Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all the ...
, and isopods. Although this species has a highly specialized diet, it also seems to vary with seasonal changes, leading to differences in
prey Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
species abundance. A study conducted in Apalachee Bay, Florida, found that the most important prey species were ''Cymadusa compta'' and ''Hippolyte zostericola''.


References

fasciatus Fish described in 1876 Fish of the Atlantic Ocean {{Labrisomidae-stub