''Panphagia'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

sauropodomorph Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lon ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

described in 2009. It lived around 231 million years ago, during the

Carnian The Carnian (less commonly, Karnian) is the lowermost stage (stratigraphy), stage of the Upper Triassic series (stratigraphy), Series (or earliest age (geology), age of the Late Triassic Epoch (reference date), Epoch). It lasted from 237 to 227 m ...

age of the

Late Triassic The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch ...

period in what is now northwestern

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the List of South American countries by area, second-largest ...

. Fossils of the genus were found in the La Peña Member of the

Ischigualasto Formation The Ischigualasto Formation is a Late Triassic fossiliferous formation and Lagerstätte in the Ischigualasto-Villa Unión Basin of the southwestern La Rioja Province and northeastern San Juan Province in northwestern Argentina. The formation ...

in the

Ischigualasto-Villa Unión Basin The Ischigualasto-Villa Unión Basin ( es, Cuenca de Ischigualasto-Villa Unión) is a small sedimentary basin located in the Argentine Northwest, Argentina. It is located in the southwestern part of La Rioja Province, Argentina, La Rioja Province a ...

. The name ''Panphagia'' comes from the

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

words ', meaning "all", and ', meaning "to eat", in reference to its inferred

omnivorous An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nut ...

diet. ''Panphagia'' is one of the earliest known dinosaurs, and is an important find which may mark the transition of diet in early sauropodomorph dinosaurs.

Discovery

Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

s of ''Panphagia'' were found in late 2006 by the Argentinean paleontologist Ricardo N. Martínez in rocks of the

of Valle Pintado,

Ischigualasto Provincial Park Ischigualasto Provincial Park ( es, Parque Provincial Ischigualasto), also called ''Valle de la Luna'' ("Valley of the Moon" or "Moon Valley"), due to its moon like appearance, is a provincial protected area in the north-east of San Juan Provin ...

, San Juan Province,

. The bones were found at approximately the same level as a 231.4 million year old ash layer, indicating it lived during the early

of the

. ''Panphagia'' is currently known from

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

PVSJ 874, the disarticulated remains of one partially grown individual of about long. Portions of the skull,

vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...

pectoral girdle The shoulder girdle or pectoral girdle is the set of bones in the appendicular skeleton which connects to the arm on each side. In humans it consists of the clavicle and scapula; in those species with three bones in the shoulder, it consists of t ...

pelvic girdle The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The p ...

, and hindlimb bones have been recovered. The russet-colored fossils were embedded in a greenish sandstone matrix and took several years to prepare and describe.

Classification

''Panphagia'' was described in 2009 by Ricardo N. Martínez and Oscar A. Alcober, both of the Museo de Ciencias Naturales, in San Juan, Argentina. They performed a

phylogenetic analysis In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...

and found it to be the most basal known sauropodomorph dinosaur: the fossils shared similar features to those of ''

Saturnalia Saturnalia is an ancient Roman festival and holiday in honour of the god Saturn, held on 17 December of the Julian calendar and later expanded with festivities through to 23 December. The holiday was celebrated with a sacrifice at the Temple ...

'', an early sauropodomorph, including similarities in the

ischium The ischium () form ...

astragalus ''Astragalus'' is a large genus of over 3,000 species of herbs and small shrubs, belonging to the legume family Fabaceae and the subfamily Faboideae. It is the largest genus of plants in terms of described species. The genus is native to tempe ...

, and the

scapula The scapula (plural scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on eithe ...

r blade. Yet the fossils also exhibited similar features to those of ''

Eoraptor ''Eoraptor'' () is a genus of small, lightly built, basal sauropodomorph. One of the earliest-known dinosaurs, it lived approximately 231 to 228 million years ago, during the Late Triassic in Western Gondwana, in the region that is now northwest ...

'', an early omnivorous sauropodomorph, including hollow bones, sublanceolate teeth, and overall proportions. Based on analysis and comparison of the ''Panphagia'' fossils and those of its closest kin, Martínez and Alcober concluded that the evolution of saurischian dinosaurs likely began with small, cursorial animals similar to ''Panphagia'', and that there is a "general similarity among all of these basal dinosaurs uggestingthat few structural changes stand" between ''Panphagia'', ''Eoraptor'', and two basal theropods which have yet to be described. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

of ''Panphagia'' is ''P. protos''; the specific name, meaning "the first" in Greek, is a reference to its basal position.

Diet

The teeth of ''Panphagia'' indicate a possible omnivorous diet, transitional in form between the mostly carnivorous theropods and the herbivorous sauropodomorphs. The teeth in the back of the jaw are shorter than those in the front, are leaf-shaped, and also have more marked serrations.

References

{{Portal bar, Dinosaurs Sauropodomorphs Dinosaur genera Carnian genera Late Triassic dinosaurs of South America Triassic Argentina Fossils of Argentina Ischigualasto Formation Fossil taxa described in 2009

Discovery

Classification

Diet

See also

References