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Paleontology in Oklahoma refers to
paleontological Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (geology), epoch (roughly 11,700 years before present). It includes t ...
research occurring within or conducted by people from the
U.S. state In the United States, a state is a constituent political entity, of which there are 50. Bound together in a political union, each state holds governmental jurisdiction over a separate and defined geographic territory where it shares its sover ...
of
Oklahoma Oklahoma (; Choctaw language, Choctaw: ; chr, ᎣᎧᎳᎰᎹ, ''Okalahoma'' ) is a U.S. state, state in the South Central United States, South Central region of the United States, bordered by Texas on the south and west, Kansas on the nor ...
. Oklahoma has a rich
fossil record A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...
spanning all three eras of the
Phanerozoic Eon The Phanerozoic Eon is the current geologic eon in the geologic time scale, and the one during which abundant animal and plant life has existed. It covers 538.8 million years to the present, and it began with the Cambrian Period, when anima ...
. Oklahoma is the best source of Pennsylvanian fossils in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
due to having an exceptionally complete geologic record of the epoch. From the
Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...
to the
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...
, all of Oklahoma was covered by a sea that would come to be home to creatures like
brachiopod Brachiopods (), phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, w ...
s, bryozoans,
graptolite Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and ...
s and
trilobite Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the At ...
s. During the
Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
, an expanse of coastal
deltaic A river delta is a landform shaped like a triangle, created by deposition of sediment that is carried by a river and enters slower-moving or stagnant water. This occurs where a river enters an ocean, sea, estuary, lake, reservoir, or (more rare ...
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s formed in areas of the state where early
tetrapod Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct theraps ...
s would leave behind
footprint Footprints are the impressions or images left behind by a person walking or running. Hoofprints and pawprints are those left by animals with hooves or paws rather than feet, while "shoeprints" is the specific term for prints made by shoes. They ...
s that would later fossilize. The sea withdrew altogether during the
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...
period. Oklahoma was home a variety of
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s as well as early
amphibian Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terres ...
s and
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s. Oklahoma stayed dry for most of the
Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceo ...
. During the
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
, carnivorous
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s left behind footprints that would later fossilize. During the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
, however, the state was mostly covered by the
Western Interior Seaway The Western Interior Seaway (also called the Cretaceous Seaway, the Niobraran Sea, the North American Inland Sea, and the Western Interior Sea) was a large inland sea that split the continent of North America into two landmasses. The ancient sea, ...
, which was home to huge
ammonite Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids (i.e., octopuses, squid and cuttlefish) ...
s and other marine invertebrates. During the
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
, Oklahoma became home to creatures like
bison Bison are large bovines in the genus ''Bison'' (Greek: "wild ox" (bison)) within the tribe Bovini. Two extant and numerous extinct species are recognised. Of the two surviving species, the American bison, ''B. bison'', found only in North Ame ...
,
camel A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. C ...
s,
creodont Creodonta ("meat teeth") is a former order of extinct carnivorous placental mammals that lived from the early Paleocene to the late Miocene epochs in North America, Europe, Asia and Africa. Originally thought to be a single group of animals ance ...
s, and
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million y ...
s. During the
Ice Age An ice age is a long period of reduction in the temperature of Earth's surface and atmosphere, resulting in the presence or expansion of continental and polar ice sheets and alpine glaciers. Earth's climate alternates between ice ages and gree ...
, the state was home to
mammoth A mammoth is any species of the extinct elephantid genus ''Mammuthus'', one of the many genera that make up the order of trunked mammals called proboscideans. The various species of mammoth were commonly equipped with long, curved tusks and, ...
s and
mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of th ...
s. Local Native Americans are known to have used fossils for
medicinal Medicine is the science and practice of caring for a patient, managing the diagnosis, prognosis, prevention, treatment, palliation of their injury or disease, and promoting their health. Medicine encompasses a variety of health care practic ...
purposes. The
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
dinosaur '' Saurophaganax maximus'' is the Oklahoma
state fossil Most American states have made a state fossil designation, in many cases during the 1980s. It is common to designate one species in which fossilization has occurred, rather than a single specimen, or a category of fossils not limited to a single ...
.


Prehistory


Paleozoic

No
Precambrian The Precambrian (or Pre-Cambrian, sometimes abbreviated pꞒ, or Cryptozoic) is the earliest part of Earth's history, set before the current Phanerozoic Eon. The Precambrian is so named because it preceded the Cambrian, the first period of the ...
fossils are known from Oklahoma, and the state's fossil record begins in the
Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...
. From the
Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...
to the
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...
, Oklahoma was covered by a sea. Cambrian life of Oklahoma included
brachiopod Brachiopods (), phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, w ...
s,
graptolite Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and ...
s,
sponge Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through t ...
s, and
trilobite Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the At ...
s. Oklahoma's
Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start ...
life included several species of brachiopods, bryozoans,
echinoderm An echinoderm () is any member of the phylum Echinodermata (). The adults are recognisable by their (usually five-point) radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the sea ...
s,
conodont Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which ...
s, and
ostracod Ostracods, or ostracodes, are a class of the Crustacea (class Ostracoda), sometimes known as seed shrimp. Some 70,000 species (only 13,000 of which are extant) have been identified, grouped into several orders. They are small crustaceans, typic ...
s. Abundant remains are fossilized at Rock Crossing in the
Criner Hills Criner is a surname. Notable people with the surname include: *Jim Criner (born 1940), American football player and coach *Juron Criner Juron Criner (born December 12, 1989) is a former American football wide receiver. Born in Las Vegas, Nevada ...
of southern Oklahoma. One common Oklahoman graptolite was ''
Climacograptus ''Climacograptus'' is an Ordovician to Silurian genus of graptolite Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding Filter feeders are a sub-group of ...
''. High quality specimens of the trilobite ''
Isotelus ''Isotelus'' is a genus of asaphid trilobites from the middle and upper Ordovician period, fairly common in the Northeastern United States, northwest Manitoba, southwestern Quebec and southeastern Ontario. One species, ''Isotelus rex,'' is curre ...
'' were preserved southwest of Ardmore. During the
Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozo ...
, Oklahoma was home to brachiopods, bryozoans, the trilobite ''
Calymene ''Calymene'' Brongniart, 1822, is a genus of trilobites in the order Phacopida, suborder Calymenina, that are found throughout North America, North Africa, and Europe in primarily Silurian outcrops. ''Calymene'' is closely related to ''Flexicaly ...
'', echinoderms, and
sponge Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through t ...
s, all of which are preserved south of Lawrence Creek. Oklahoma was home to an extremely diverse
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...
fauna in the Lawrence and White Mound areas. During the Mississippian, Oklahoma's local fauna included ''
Archimedes Archimedes of Syracuse (;; ) was a Greek mathematician, physicist, engineer, astronomer, and inventor from the ancient city of Syracuse in Sicily. Although few details of his life are known, he is regarded as one of the leading scientists ...
'', brachiopods,
conodont Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which ...
s, echinoderms, the
blastoid Blastoids (class Blastoidea) are an extinct type of stemmed echinoderm, often referred to as sea buds. They first appear, along with many other echinoderm classes, in the Ordovician period, and reached their greatest diversity in the Mississi ...
''
Pentremites ''Pentremites'' is an extinct genus of blastoid echinoderm belonging to the family Pentremitidae. Description These stalked echinoderms averaged a height of about but occasionally ranged up to about 3 times that size. They, like other blastoid ...
'', and trilobites. Contemporary brachiopod families included the
productid Productida is an extinct order of brachiopods in the extinct class Strophomenata. Members of Productida first appeared during the Silurian. They represented the most abundant group of brachiopods during the Permian period, accounting for 45-70% ...
s and
rhynchonellida The taxonomic order Rhynchonellida is one of the two main groups of living articulate brachiopods, the other being the order Terebratulida. They are recognized by their strongly ribbed wedge-shaped or nut-like shells, and the very short hing ...
. The best source of Mississippian fossils in Oklahoma is the state's northeastern region. During the Carboniferous, Oklahoma was a terrestrial environment characterized by vast river systems and accompanying deltas. These deltas were home to vast swamps responsible for leaving behind many coal deposits. During the Carboniferous, early
tetrapod Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct theraps ...
s left behind
footprints Footprints are the impressions or images left behind by a person walking or running. Hoofprints and pawprints are those left by animals with hoof, hooves or paws rather than foot, feet, while "shoeprints" is the specific term for prints made by ...
that would later fossilize. Oklahoma's diverse Pennsylvanian life included blastoids, brachiopods, bryozoans,
fusulinid The Fusulinida is an extinct order within the Foraminifera in which the tests are traditionally considered to have been composed of microgranular calcite. Like all forams, they were single-celled organisms. In advanced forms the test wall was di ...
s, and
pelecypod Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, biva ...
s. Vertebrates included various fishes, and the early tetrapods likely responsible for the trackways. Occasionally during this period, sea levels would rise and cover the state again. This sea gradually retreated from the state before the end of the Paleozoic era. Oklahoma preserves one of the richest fossil records of non-marine vertebrates from the Permian of North America with respect to both abundance of material and taxonomic diversity, with particularly notable records from early Permian sites such as
Richards Spur Richards Spur is a Permian fossil locality located at the Dolese Brothers Limestone Quarry north of Lawton, Oklahoma. The locality preserves clay and mudstone fissure fills of a karst system eroded out of Ordovician limestone and dolomite, with the ...
in Comanche County and South Grandfield in Tillman County. Most of these deposits are distributed across the western half of the state, including in
Logan Logan may refer to: Places * Mount Logan (disambiguation) Australia * Logan (Queensland electoral district), an electoral district in the Queensland Legislative Assembly * Logan, Victoria, small locality near St. Arnaud * Logan City, local gover ...
,
Noble A noble is a member of the nobility. Noble may also refer to: Places Antarctica * Noble Glacier, King George Island * Noble Nunatak, Marie Byrd Land * Noble Peak, Wiencke Island * Noble Rocks, Graham Land Australia * Noble Island, Great B ...
,
Grant Grant or Grants may refer to: Places *Grant County (disambiguation) Australia * Grant, Queensland, a locality in the Barcaldine Region, Queensland, Australia United Kingdom *Castle Grant United States * Grant, Alabama *Grant, Inyo County, C ...
,
Garfield ''Garfield'' is an American comic strip created by Jim Davis. Originally published locally as ''Jon'' in 1976, then in nationwide syndication from 1978 as ''Garfield'', it chronicles the life of the title character Garfield the cat, his human ...
, Kay,
Pawnee Pawnee initially refers to a Native American people and its language: * Pawnee people * Pawnee language Pawnee is also the name of several places in the United States: * Pawnee, Illinois * Pawnee, Kansas * Pawnee, Missouri * Pawnee City, Nebraska * ...
, and Payne Counties. In particular, there is extensive body fossil documentation of many groups of extinct vertebrates, including
lungfish Lungfish are freshwater vertebrates belonging to the order Dipnoi. Lungfish are best known for retaining ancestral characteristics within the Osteichthyes, including the ability to breathe air, and ancestral structures within Sarcopterygii, i ...
, various ' lepospondyls' like aïstopods,
nectridea Nectridea is the name of an extinct order of lepospondyl tetrapods from the Carboniferous and Permian periods, including animals such as ''Diplocaulus''. In appearance, they would have resembled modern newts or aquatic salamanders, although they ...
ns, and ' microsaurs,'
temnospondyl Temnospondyli (from Greek language, Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order (biology), order of small to giant tetrapods—often considered Labyrinthodontia, primitive amphi ...
amphibians,
parareptiles Parareptilia ("at the side of reptiles") is a subclass or clade of basal sauropsids ( reptiles), typically considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds). Parareptiles first arose near t ...
,
eureptiles Eureptilia ("true reptiles") is one of the two major subgroups of the clade Sauropsida, the other one being Parareptilia. Eureptilia includes Diapsida (the clade containing all modern reptiles and birds), as well as a number of primitive Permo-C ...
,
reptiliomorphs Reptiliomorpha (meaning reptile-shaped; in PhyloCode known as ''Pan-Amniota'') is a clade containing the amniotes and those tetrapods that share a more recent common ancestor with amniotes than with living amphibians (lissamphibians). It was defi ...
(stem amniotes),
synapsid Synapsids + (, 'arch') > () "having a fused arch"; synonymous with ''theropsids'' (Greek, "beast-face") are one of the two major groups of animals that evolved from basal amniotes, the other being the sauropsids, the group that includes reptil ...
s, and
diapsid Diapsids ("two arches") are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The group first appeared about three hundred million years ago ...
s. The giant Permian
foraminiferan Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly an ...
'' Pseudoschwagerina'' was preserved in the
Pawnee Pawnee initially refers to a Native American people and its language: * Pawnee people * Pawnee language Pawnee is also the name of several places in the United States: * Pawnee, Illinois * Pawnee, Kansas * Pawnee, Missouri * Pawnee City, Nebraska * ...
area. Many of these tetrapods likely produced a variety of trackways also known from the early Permian of Oklahoma. There is also an extensive record of invertebrates, such as
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s and
millipede Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a resu ...
s, as well as brachiopods and foraminifers. It remains controversial whether there are any middle Permian tetrapods known from Oklahoma, which would represent perhaps the only such record from this time period in all of North America and perhaps the entire globe; if tetrapod records from the
Chickasha Formation The Chickasha Formation is a geologic formation in Oklahoma. It preserves fossils dating back to the Permian period. See also * List of fossiliferous stratigraphic units in Oklahoma * Paleontology in Oklahoma Paleontology in Oklahoma refers ...
and the
Flowerpot Formation The Flowerpot Formation is a Formation (geology), geologic formation in Oklahoma. It preserves fossils dating back to the Permian Period (geology), period. See also * List of fossiliferous stratigraphic units in Oklahoma * Paleontology in Oklah ...
in Blaine,
Canadian Canadians (french: Canadiens) are people identified with the country of Canada. This connection may be residential, legal, historical or cultural. For most Canadians, many (or all) of these connections exist and are collectively the source of ...
, Grady, and
Kingfisher Kingfishers are a family, the Alcedinidae, of small to medium-sized, brightly colored birds in the order Coraciiformes. They have a cosmopolitan distribution, with most species found in the tropical regions of Africa, Asia, and Oceania, ...
Counties and their equivalents in Texas (the
San Angelo Formation The San Angelo Formation is a geologic formation in Texas. It preserves fossils dating back to the Permian period. It is one of the geologically youngest formations to preserve fossils of pelycosaurs. Stratigraphy and age The San Angelo Formati ...
) are not considered to be middle Permian in age, there would be a hiatus in the fossil record, which is termed 'Olson's Gap,', although records from other geographic regions may fill this gap regardless of whether it existed in North America.


Mesozoic

Oklahoma was a terrestrial environment for most of the ensuing
Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceo ...
era. The
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
Dockum Group The Dockum is a Late Triassic (approximately late Carnian through Rhaetian, or 223–200 Ma) geologic group found primarily on the Llano Estacado of western Texas and eastern New Mexico with minor exposures in southwestern Kansas, eastern Color ...
of western Oklahoma preserved remains of archosaurs and temnospondyls, although its fossil record is restricted to a narrow region of the panhandle and is far sparser than the equivalent records in Texas, New Mexico, and Arizona. During the Late Triassic, small carnivorous dinosaurs left behind tracks near Kenton now classified in the
ichnogenus An ichnotaxon (plural ichnotaxa) is "a taxon based on the fossilized work of an organism", i.e. the non-human equivalent of an artifact. ''Ichnotaxa'' comes from the Greek ίχνος, ''ichnos'' meaning ''track'' and ταξις, ''taxis'' meaning ...
''
Grallator ''Grallator'' GRA-luh-tor"is an ichnogenus (form taxon based on footprints) which covers a common type of small, three-toed print made by a variety of bipedal theropod dinosaurs. ''Grallator''-type footprints have been found in formations dating ...
''. The sediments preserving these tracks later became the
Sheep Pen Sandstone The Sheep Pen Sandstone is a late Triassic geologic formation exposed in northeastern New Mexico. Fossil theropod tracks have been reported from the formation. Description The formation consists of up to of thinly bedded light brown sandstone. ...
. Other local tracks have been referred to ''
Chirotherium ''Chirotherium'', also known as ''Cheirotherium'' (‘hand-beast’), is a Triassic trace fossil consisting of five-fingered (pentadactyle) footprints and whole tracks. These look, by coincidence, remarkably like the hands of apes and bears, with ...
'', but Martin G. Lockley and
Adrian Hunt Adrian is a form of the Latin given name Adrianus or Hadrianus. Its ultimate origin is most likely via the former river Adria from the Venetic and Illyrian word ''adur'', meaning "sea" or "water". The Adria was until the 8th century BC the main ...
have speculated that these might actually be '' Pseudotetrasauropus''. The
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
fossiliferous
Morrison Formation The Morrison Formation is a distinctive sequence of Late Jurassic, Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandsto ...
is exposed in the western part of the state and has produced extensive remains of sauropod dinosaurs. Most of Oklahoma was submerged under the
Western Interior Seaway The Western Interior Seaway (also called the Cretaceous Seaway, the Niobraran Sea, the North American Inland Sea, and the Western Interior Sea) was a large inland sea that split the continent of North America into two landmasses. The ancient sea, ...
during the Cretaceous.
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
life included "immense"
ammonite Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids (i.e., octopuses, squid and cuttlefish) ...
s,
echinoid Sea urchins () are spine (zoology), spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard s ...
s, and
pelecypod Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, biva ...
s. These fossils were preserved in
Love Love encompasses a range of strong and positive emotional and mental states, from the most sublime virtue or good habit, the deepest Interpersonal relationship, interpersonal affection, to the simplest pleasure. An example of this range of ...
and Marshall counties. The
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
rocks of Bryan,
Choctaw The Choctaw (in the Choctaw language, Chahta) are a Native American people originally based in the Southeastern Woodlands, in what is now Alabama and Mississippi. Their Choctaw language is a Western Muskogean language. Today, Choctaw people are ...
, and McCurtain counties bear abundant
oysters Oyster is the common name for a number of different families of salt-water bivalve molluscs that live in marine or brackish habitats. In some species, the valves are highly calcified, and many are somewhat irregular in shape. Many, but not al ...
like ''
Exogyra ''Exogyra'' is an extinct genus of fossil marine oysters in the family Gryphaeidae, the foam oysters or honeycomb oysters. These bivalves grew cemented by the more cupped left valve. The right valve is flatter, and the beak is curved to one si ...
'' and '' Ostraea''. However, there are also records of many terrestrial vertebrates, particularly from the
Antlers Antlers are extensions of an animal's skull found in members of the Cervidae (deer) family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally found only on male ...
and Cloverly Formations, including fish, amphibians, reptiles, crocodiles, dinosaurs, and mammals.


Cenozoic

As the
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico in ...
rose during the early
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
, rivers drained off them and into Oklahoma. Sediments deposited by these rivers would preserve
petrified wood Petrified wood, also known as petrified tree (from Ancient Greek meaning 'rock' or 'stone'; literally 'wood turned into stone'), is the name given to a special type of ''fossilized wood'', the fossilized remains of terrestrial vegetation. ''P ...
and mammal fossils. Sediments were generally being eroded away from Oklahoma during the later portion of the Cenozoic. The
High Plains High Plains refers to one of two distinct land regions: * High Plains (United States), land region of the western Great Plains *High Plains (Australia) The High Plains of south-eastern Australia are a sub-region, or more strictly a string of adja ...
of the western part of Oklahoma preserve evidence for the presence of
camel A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. C ...
s,
creodont Creodonta ("meat teeth") is a former order of extinct carnivorous placental mammals that lived from the early Paleocene to the late Miocene epochs in North America, Europe, Asia and Africa. Originally thought to be a single group of animals ance ...
s, and
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million y ...
s during the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58 During the ensuing
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
epoch, resident animals included
mammoth A mammoth is any species of the extinct elephantid genus ''Mammuthus'', one of the many genera that make up the order of trunked mammals called proboscideans. The various species of mammoth were commonly equipped with long, curved tusks and, ...
s and
mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of th ...
. Their fossils were preserved in several different regions of Oklahoma. Typical Oklahoman
proboscidean The Proboscidea (; , ) are a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. From ...
fossils are teeth and tusks, often preserved in gravel pits, but complete skeletons are also known. Other mammals found in Pleistocene Oklahoma included ''
Glyptotherium ''Glyptotherium'' (from Greek for 'grooved or carved beast') is a genus of glyptodont (an extinct group of large, herbivorous armadillos) that lived from the Early Pliocene, about 4.9 million years ago, to the Early Holocene, around 7,000 years ...
'', a large, heavily armored mammal related to the armadillo.


History


Indigenous interpretations

The
Comanche The Comanche or Nʉmʉnʉʉ ( com, Nʉmʉnʉʉ, "the people") are a Native American tribe from the Southern Plains of the present-day United States. Comanche people today belong to the federally recognized Comanche Nation, headquartered in La ...
people gathered fossils in Comanche County, near Indiahoma to be used as
medicine Medicine is the science and practice of caring for a patient, managing the diagnosis, prognosis, prevention, treatment, palliation of their injury or disease, and promoting their health. Medicine encompasses a variety of health care pract ...
for
sprain A sprain, also known as a torn ligament, is an acute soft tissue injury of the ligaments within a joint, often caused by a sudden movement abruptly forcing the joint to exceed its functional range of motion. Ligaments are tough, inelastic fibers ...
s and
bone fracture A bone fracture (abbreviated FRX or Fx, Fx, or #) is a medical condition in which there is a partial or complete break in the continuity of any bone in the body. In more severe cases, the bone may be broken into several fragments, known as a '' ...
s. The
Comanche The Comanche or Nʉmʉnʉʉ ( com, Nʉmʉnʉʉ, "the people") are a Native American tribe from the Southern Plains of the present-day United States. Comanche people today belong to the federally recognized Comanche Nation, headquartered in La ...
ground up the bone into a powder known as tsoapitsitsuhni, which translates to "ghost creature bone", and mixed it with water. This mixture could be made into a sort of plaster cast if the fossils used to make the powder contained sufficient
gypsum Gypsum is a soft sulfate mineral composed of calcium sulfate dihydrate, with the chemical formula . It is widely mined and is used as a fertilizer and as the main constituent in many forms of plaster, blackboard or sidewalk chalk, and drywall. ...
or
calcium sulphate Calcium sulfate (or calcium sulphate) is the inorganic compound with the formula CaSO4 and related hydrates. In the form of γ-anhydrite (the anhydrous form), it is used as a desiccant. One particular hydrate is better known as plaster of Pari ...
content. The local geology consist largely of
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...
-aged
red beds Red beds (or redbeds) are sedimentary rocks, typically consisting of sandstone, siltstone, and shale, that are predominantly red in color due to the presence of ferric oxides. Frequently, these red-colored sedimentary strata locally contain ...
, and Comanche County's eastern side contains
Richards Spur Richards Spur is a Permian fossil locality located at the Dolese Brothers Limestone Quarry north of Lawton, Oklahoma. The locality preserves clay and mudstone fissure fills of a karst system eroded out of Ordovician limestone and dolomite, with the ...
, the best source of Permian fossils in the entire state. Reptile and amphibian fossils like ''
Captorhinus ''Captorhinus'' (from el, καπτō , 'to gulp down' and el, ῥῑνός , 'nose') is an extinct genus of captorhinid reptiles that lived during the Permian period. Its remains are known from Oklahoma, Texas, Europe, India, the Pedra de Fogo ...
'' are found nearby in other counties. Such Permian remains are viable candidates for the fossils used medicinally by the Comanche, but local
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
and
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
remains like those of ''
Apatosaurus ''Apatosaurus'' (; meaning "deceptive lizard") is a genus of herbivorous sauropod dinosaur that lived in North America during the Late Jurassic period. Othniel Charles Marsh described and named the first-known species, ''A. ajax'', in 1877, an ...
'', ''
Saurophaganax ''Saurophaganax'' ("lord of lizard-eaters") is a genus of large allosaurid dinosaur from the Morrison Formation of Late Jurassic (latest Kimmeridgian age, about 151 million years ago) Oklahoma, United States.Turner, C.E. and Peterson, F., (1999) ...
'', ''
Sauroposeidon ''Sauroposeidon'' ( ; meaning "lizard earthquake god", after the Greek god Poseidon) is a genus of sauropod dinosaur known from several incomplete specimens including a bone bed and fossilized trackways that have been found in the U.S. states of ...
'' and ''
Tenontosaurus ''Tenontosaurus'' ( ; ) is a genus of medium- to large-sized ornithopod dinosaur. It was a relatively medium sized ornithopod, reaching in length and in body mass. It had an unusually long, broad tail, which like its back was stiffened with a n ...
'' are also candidates. More recent
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
fossils were also used by the Comanche for medicine like those of
bear Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the Nor ...
s, giant
bison Bison are large bovines in the genus ''Bison'' (Greek: "wild ox" (bison)) within the tribe Bovini. Two extant and numerous extinct species are recognised. Of the two surviving species, the American bison, ''B. bison'', found only in North Ame ...
,
camel A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. C ...
s,
glyptodont Glyptodonts are an extinct subfamily of large, heavily armoured armadillos. They arose in South America around 48 million years ago and spread to southern North America after the continents became connected several million years ago. The best-kn ...
s,
Columbian mammoth The Columbian mammoth (''Mammuthus columbi'') is an extinct species of mammoth that inhabited the Americas as far north as the Northern United States and as far south as Costa Rica during the Pleistocene epoch. It was one of the last in a line ...
s, and
mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of th ...
s. Comanches used bits of mammoth leg bone to draw out
boil A boil, also called a furuncle, is a deep folliculitis, which is an infection of the hair follicle. It is most commonly caused by infection by the bacterium ''Staphylococcus aureus'', resulting in a painful swollen area on the skin caused by an ...
s, infections, poisons and pain from wounds. This usage is fairly plausible as the porous nature of fossil bone causes a capillary effect that could be used to dry infected wounds and sores. Mammoth bone used for this purpose was known as medicinebone or madstone.


Scientific research

In 1931,
University of Oklahoma The University of Oklahoma (OU) is a Public university, public research university in Norman, Oklahoma. Founded in 1890, it had existed in Oklahoma Territory near Indian Territory for 17 years before the two Territories became the state of Oklahom ...
geologist
J. Willis Stovall John Willis Stovall (1891– July 24, 1953) was an American paleontologist at the University of Oklahoma. Along with his student Wann Langston, Jr., he named the theropod dinosaur ''Acrocanthosaurus ''Acrocanthosaurus'' ( ; ) is a genus of ...
received word that a road crew grading for the construction of
U.S. Route 64 U.S. Route 64 (US 64) is an east–west United States highway that runs for 2,326 miles (3,743 km) from Nags Head in eastern North Carolina to just southwest of the Four Corners in northeast Arizona. The western terminus is at U.S. Route 1 ...
uncovered a rich deposit of fossils east of Kenton. Stovall examined the site and was impressed by the fossils uncovered by the workers. He organized an expedition to the region. By 1935, Stovall assembled a team consisting of students and a handful of
Works Progress Administration The Works Progress Administration (WPA; renamed in 1939 as the Work Projects Administration) was an American New Deal agency that employed millions of jobseekers (mostly men who were not formally educated) to carry out public works projects, i ...
workers. He placed a local named
Crompton Tate Crompton may refer to Place names *Crompton (West Warwick), a community in West Warwick, Rhode Island, US *Crompton, Greater Manchester, in Shaw and Crompton, Greater Manchester, England formerly in Lancashire *Crompton Urban District, an obsole ...
in charge of the team. Stovall's team excavated the site for nearly three years, in the process digging through almost 100 metric tons of rock and sediment to extract the remains preserved there. The site was called Quarry 1, the first of seventeen quarries that the expedition would start in the region. The excavation uncovered the bones from many kinds of dinosaurs. Finds of previously documented species included both sizable and hatchling ''
Apatosaurus ''Apatosaurus'' (; meaning "deceptive lizard") is a genus of herbivorous sauropod dinosaur that lived in North America during the Late Jurassic period. Othniel Charles Marsh described and named the first-known species, ''A. ajax'', in 1877, an ...
'', hatchling ''
Camarasaurus ''Camarasaurus'' ( ) was a genus of quadrupedal, herbivorous dinosaurs and is the most common North American sauropod fossil. Its fossil remains have been found in the Morrison Formation, dating to the Late Jurassic epoch (Kimmeridgian to Titho ...
'', several ''
Camptosaurus ''Camptosaurus'' ( ) is a genus of plant-eating, beaked ornithischian dinosaurs of the Late Jurassic period of western North America and possibly also Europe. The name means 'flexible lizard' (Greek (') meaning 'bent' and (') meaning 'lizard') ...
'' of different age groups, and ''
Stegosaurus ''Stegosaurus'' (; ) is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been foun ...
'' fossils. The new
theropoda Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ca ...
species that would come to be known as Saurophaganax was also discovered there. By December 1939, excavation had commenced on the Stovall team's fifth quarry. The most significant remains uncovered there are referable to the large sauropod ''
Diplodocus ''Diplodocus'' (, , or ) was a genus of diplodocid sauropod dinosaurs, whose fossils were first discovered in 1877 by S. W. Williston. The generic name, coined by Othniel Charles Marsh in 1878, is a neo-Latin term derived from Greek διπ ...
''. Prior to the cessation of digging at Quarry 5 in the middle of 1941, this quarry had attained impressive dimensions. Its walls were nine meters (30 feet) high and the breadth of the excavation wide. Other notable quarries excavated by the Stovall team include the eighth, which produced fossils of
ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (), that started out as small, bipedal running grazers and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous world ...
and
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaurs as well as other reptiles like a new species of
crocodilia Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...
n, ''
Cteniogenys ''Cteniogenys'' is a genus of choristodere, a morphologically diverse group of aquatic reptiles. It is part of the monotypic family Cteniogenidae. The type and only named species, ''C. antiquus'', was named in 1928 by Charles W. Gilmore. The hol ...
'', and
turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked tu ...
s. Lungfish were also preserved there. Funding for Stovall's field work ended with the advent of World War II in 1942, interrupting excavations at Quarries 9 and 10. In 1964, Charles Mook named the new crocodilian species uncovered by the Stovall team '' Goniopholis stovalli'' in his honor. The new theropod from Quarry 1 was named ''
Saurophagus The great kiskadee (''Pitangus sulphuratus''), called ''bem-te-vi'' in Brazil and ''benteveo'' in Argentina, is a passerine bird in the tyrant flycatcher family Tyrannidae. It is the only member of the genus ''Pitangus''. It breeds in open woodl ...
''. In 1995, Dan Chure published a new name for ''Saurophagus'' since that name had already been used for another kind of animal; he renamed it '' Saurophaganax maximus''. More recently, in 2004, Matt Bonnan and Matt Wedel noticed the presence of at least one ''
Brachiosaurus ''Brachiosaurus'' () is a genus of sauropod dinosaur that lived in North America during the Late Jurassic, about 154to 150million years ago. It was first described by American paleontologist Elmer S. Riggs in 1903 from fossils found in th ...
'' bone among the fossils excavated by the Stovall Crew at Quarry 1.


Natural history museums

*
Sam Noble Oklahoma Museum of Natural History Sam, SAM or variants may refer to: Places * Sam, Benin * Sam, Boulkiemdé, Burkina Faso * Sam, Bourzanga, Burkina Faso * Sam, Kongoussi, Burkina Faso * Sam, Iran * Sam, Teton County, Idaho, United States, a populated place People and fictio ...
,
Norman Norman or Normans may refer to: Ethnic and cultural identity * The Normans, a people partly descended from Norse Vikings who settled in the territory of Normandy in France in the 10th and 11th centuries ** People or things connected with the Norm ...
*
The Museum of Osteology ''The'' () is a grammatical Article (grammar), article in English language, English, denoting persons or things already mentioned, under discussion, implied or otherwise presumed familiar to listeners, readers, or speakers. It is the definite ...
,
Oklahoma City Oklahoma City (), officially the City of Oklahoma City, and often shortened to OKC, is the capital and largest city of the U.S. state of Oklahoma. The county seat of Oklahoma County, it ranks 20th among United States cities in population, a ...


See also

*
Bromide Formation The Bromide Formation is a geological formation in Oklahoma, USA. It is well known for its diverse echinoderm and trilobite fossil fauna. Location The Bromide Formation crops out in the Arbuckle and Wichita Mountains and in the Criner Hills ...
*
Paleontology in Arkansas Paleontology in Arkansas refers to paleontological research occurring within or conducted by people from the United States, U.S. state of Arkansas. The fossil Fossil record, record of Arkansas spans from the Ordovician to the Eocene. Nearly all of ...
*
Paleontology in Kansas Paleontology in Kansas refers to paleontological research occurring within or conducted by people from the U.S. state of Kansas. Kansas has been the source of some of the most spectacular fossil discoveries in US history. The fossil record of Kansa ...
*
Paleontology in Missouri Paleontology in Missouri refers to paleontological research occurring within or conducted by people from the U.S. state of Missouri. The geologic column of Missouri spans all of geologic history from the Precambrian to present with the exception ...
*
Paleontology in New Mexico Paleontology in New Mexico refers to paleontological research occurring within or conducted by people from the U.S. state of New Mexico. The fossil record of New Mexico is exceptionally complete and spans almost the entire stratigraphic column. ...
*
Paleontology in Texas Paleontology in Texas refers to paleontological research occurring within or conducted by people from the U.S. state of Texas. Author Marian Murray has remarked that "Texas is as big for fossils as it is for everything else." Some of the most impor ...


Footnotes


References

* Everhart, M. J. 2005. ''Oceans of Kansas - A Natural History of the Western Interior Sea''. Indiana University Press, 320 pp. * Foster, J. (2007). ''Jurassic West: The Dinosaurs of the Morrison Formation and Their World.'' Indiana University Press. 389pp. . * Lockley, Martin and Hunt, Adrian. '' Dinosaur Tracks of Western North America''. Columbia University Press. 1999. * Mayor, Adrienne. ''Fossil Legends of the First Americans''. Princeton University Press. 2005. . * * Springer, Dale, Judy Scotchmoor. July 14, 2010.
Oklahoma, US

The Paleontology Portal
Accessed September 21, 2012.


External links


Geologic units in Oklahoma

Sam Noble Museum: Common Fossils of Oklahoma

Paleoportal: Oklahoma
{{Paleontology in the United States
Oklahoma Oklahoma (; Choctaw language, Choctaw: ; chr, ᎣᎧᎳᎰᎹ, ''Okalahoma'' ) is a U.S. state, state in the South Central United States, South Central region of the United States, bordered by Texas on the south and west, Kansas on the nor ...
Natural history of Oklahoma Science and technology in Oklahoma