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The Palaeonisciformes (Palaeoniscida) are an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
of early ray-finned fishes (
Actinopterygii Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...
). Palaeonisciformes ''
sensu lato ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular co ...
'' first appeared in the
fossil record A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...
in the Late
Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozo ...
and last appeared in the Late
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
. The name is derived from the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
words παλαιός (''palaiós'', ancient) and ὀνίσκος (''oniskos'', 'cod-fish' or woodlouse), probably pertaining to the organization of the fishes' scales, similar to the exoskeletal plating of woodlice. In an early interpretation of the group, the Palaeonisciformes are divided in two suborders: Palaeoniscoidei (includes ''
Palaeoniscum ''Palaeoniscum'' (from el, παλαιός , 'ancient' and el, ὀνίσκος 'cod-fish' or 'woodlouse') is an extinct genus of ray-finned fish from the Permian period (Guadalupian-Lopingian) of Europe (England, Germany, Czech Republic?, Turk ...
'' and fossil
taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
with a broadly similar appearance) and Platysomoidei (includes ''
Platysomus ''Platysomus'' (from el, πλατύς , 'broad' and el, σῶμα 'body') is an extinct genus of ray-finned fish that lived in the Carboniferous and Permian periods. Fossils have been found worldwide. ''Platysomus'' was about long, and ...
'' and other deep-bodied early actinopterygians). These groupings are considered
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
today. In the
cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...
sense, Palaeonisciformes ''
sensu stricto ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular co ...
'' should only refer to the
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...
''Palaeoniscum'', the name giving taxon, and all other taxa that fall on the same
lineage Lineage may refer to: Science * Lineage (anthropology), a group that can demonstrate its common descent from an apical ancestor or a direct line of descent from an ancestor * Lineage (evolution), a temporal sequence of individuals, populati ...
as ''Palaeoniscum''. Additionally, the term Palaeopterygii ("ancient fins") is sometimes used to group fossil and
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...
actinopterygians that are neither members of the
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
Neopterygii Neopterygii (from Greek νέος ''neos'' 'new' and πτέρυξ ''pteryx'' 'fin') is a subclass of ray-finned fish (Actinopterygii). Neopterygii includes the Holostei and the Teleostei, of which the latter comprise the vast majority of extant f ...
nor the also monophyletic
Cladistia Cladistia is a clade of bony fishes whose only living members are the bichirs. Their major synapomorphies are a heterocercal tail in which the dorsal fin has independent rays, and a posteriorly elongated parasphenoid. Cladistia are the earlie ...
. Like the Palaeonisciformes ''sensu lato'', the Palaeopterygii are also a paraphyletic assemblage.


Historic background

The
systematics Biological systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: cladograms, phylogenetic tre ...
of fossil and extant fishes has puzzled
ichthyologists Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of October ...
since the time of
Louis Agassiz Jean Louis Rodolphe Agassiz ( ; ) FRS (For) FRSE (May 28, 1807 – December 14, 1873) was a Swiss-born American biologist and geologist who is recognized as a scholar of Earth's natural history. Spending his early life in Switzerland, he rec ...
, who first grouped all
Palaeozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and '' ...
ray-finned fishes together with
Chondrostei Chondrostei is a group of non-neopterygian ray-finned fish, while the term originally referred to a paraphyletic group of all non-neopterygian ray-finned fish, it was redefined by Patterson in 1982 to be a clade comprising the Acipenseriformes (w ...
(
sturgeons Sturgeon is the common name for the 27 species of fish belonging to the family Acipenseridae. The earliest sturgeon fossils date to the Late Cretaceous, and are descended from other, earlier acipenseriform fish, which date back to the Early ...
,
paddlefish Paddlefish (family Polyodontidae) are a family of ray-finned fish belonging to order Acipenseriformes, and one of two living groups of the order alongside sturgeons (Acipenseridae). They are distinguished from other fish by their titular elongla ...
es),
gar Gars are members of the family Lepisosteidae, which are the only surviving members of the Ginglymodi, an ancient holosteian group of ray-finned fish, which first appeared during the Triassic, over 240 million years ago. Gars comprise seven livin ...
s,
lungfishes Lungfish are freshwater vertebrates belonging to the order Dipnoi. Lungfish are best known for retaining ancestral characteristics within the Osteichthyes, including the ability to breathe air, and ancestral structures within Sarcopterygii, i ...
, and
acanthodians Acanthodii or acanthodians is an extinct class of gnathostomes (jawed fishes), typically considered a paraphyletic group. They are currently considered to represent a grade of various fish lineages leading up to the extant Chondrichthyes, which ...
in his Ganoidei. Carl Hermann Müller later proposed to divide actinopterygians into three groups: Chondrostei,
Holostei Holostei is a group of ray-finned bony fish. It is divided into two major clades, the Halecomorphi, represented by a single living species, the bowfin (''Amia calva''), as well as the Ginglymodi, the sole living representatives being the gars (Le ...
, and
Teleostei Teleostei (; Ancient Greek, Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts ), is, by far, the largest class (biology), infraclass in the class Actinopterygii, the ray-finned fishes, containing 96% of a ...
. Later,
Edward Drinker Cope Edward Drinker Cope (July 28, 1840 – April 12, 1897) was an American zoologist, paleontologist, comparative anatomist, herpetologist, and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested ...
included these three groups within
Actinopteri Actinopteri is the sister group of Cladistia in the class Actinopterygii (ray-finned fish). Dating back to the Permian period, the Actinopteri comprise the Chondrostei (sturgeons and paddlefish) and the Neopterygii (bowfin, gars, and teleosts). ...
. The same classification is also used today, though the definitions of these groups have changed significantly over the years. The
sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...
to Actinopteri are the
Cladistia Cladistia is a clade of bony fishes whose only living members are the bichirs. Their major synapomorphies are a heterocercal tail in which the dorsal fin has independent rays, and a posteriorly elongated parasphenoid. Cladistia are the earlie ...
, which include ''
Polypterus ''Polypterus'' is a genus of freshwater fish in the bichir family (Polypteridae) of order Polypteriformes. The type species is the Nile bichir (''P. bichir''). Fish in this genus live in various areas in Africa. ''Polypterus'' is the only know ...
'' (bichirs), ''
Erpetoichthys The reedfish, ropefish (more commonly used in the United States), or snakefish, ''Erpetoichthys calabaricus'', is a species of fish in the bichir family and order. It is the only member of the genus ''Erpetoichthys''. It is native to fresh and ...
'' and their fossil relatives. All together are grouped as Actinopterygii. A few additional classification schemes were proposed over the years.
Lev Berg Lev Semyonovich Berg, also known as Leo S. Berg (russian: Лев Семёнович Берг; 14 March 1876 – 24 December 1950) was a leading Russian geographer, biologist and ichthyologist who served as President of the Soviet Geographical So ...
erected the superorder Palaeonisci, in which he included early actinopterygians that belonged to neither Chondrostei nor Polypteri (Cladistia). Mostly following Berg, Jean-Pierre Lehman grouped the Actinopterygii into 26 orders, among others the Palaeonisciformes with the two suborders Palaeoniscoidei and Platysomoidei. Numerous
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
of early actinopterygians have been referred to either Palaeonisciformes or to one of its suborders based on superficial resemblance with either ''Palaeoniscum'' (Palaeoniscoidei) or ''Platysomus'' (Platysomoidei), especially during the early and middle parts of the 20th century. Palaeonisciformes, Palaeoniscoidei, and Platysomoidei have therefore become wastebasket taxa. They are not natural groups, but instead
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
assemblages of the early members of several ray-finned fish lineages. Palaeoniscoidei have traditionally encompassed most
Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...
actinopterygians, except those that exhibit atypical body forms (such as the deep-bodied Platysomoidei, or those assigned securely to any of the living groups of ray-finned fishes. The same can also be said about the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Palaeoniscidae Palaeoniscidae is an extinct family of ray-finned fishes (Actinopterygii) ascribed to the order Palaeonisciformes. The family includes the genus '' Palaeoniscum'' and potentially other Palaeozoic and Mesozoic early actinopterygian genera. The ...
''sensu lato'', to which several genera not closely related to ''Palaeoniscum'' have been referred in the past. The grouping of "palaeonisciforms" was based largely on shared
plesiomorphic In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, and ...
features, such as the forward position of the eye, the large gape or the presence of rhombic scales. However, such
symplesiomorphies In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, an ...
are not informative with regard to
phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...
, but rather an indication of common ancestry. In modern biology,
taxonomists In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given ...
group taxa based on shared
apomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have ...
(
synapomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have ...
) in order to detect
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
groups (natural groups). They use computer software (e.g., PAUP) to determine the most likely
evolutionary relationships Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation t ...
between taxa, thereby putting previous
hypotheses A hypothesis (plural hypotheses) is a proposed explanation for a phenomenon. For a hypothesis to be a scientific hypothesis, the scientific method requires that one can test it. Scientists generally base scientific hypotheses on previous obser ...
of such relationships to the test. As a consequence, many genera have been subsequently removed from Palaeonisciformes and referred to distinct orders (e.g.,
Saurichthyiformes Saurichthyiformes is an extinct order of ray-finned fish which existed in Asia, Africa, Australia, Europe and North America, during the late Permian to early Middle Jurassic. Saurichthyiiformes comprise two families, Saurichthyidae and Yela ...
). The term Palaeonisciformes has mostly disappeared from the modern literature or is nowadays only used to refer to the "primitive"
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
of a taxon (e.g., "palaeonisciform skull shape" or "palaeoniscoid body shape"). In order to make the Paleonisciformes, Palaeoniscoidei or Palaeoniscidae monophyletic, these terms should only be used in a strict sense, i.e., when referring to the clade of actinopterygians that includes ''Palaeoniscum'' and the taxa closely related to it. A monophyletic clade including several taxa classically referred to the Palaeonisciformes (e.g., '' Aesopichthys'', ''
Birgeria ''Birgeria'' is a genus of carnivorous marine ray-finned fish from the Triassic period. ''Birgeria'' had a global distribution. Fossils were found in Madagascar, Spitsbergen, Germany, Switzerland, Italy, Slovenia, China, Russia, Canada and N ...
'', '' Boreosomus'', '' Canobius'', ''
Pteronisculus ''Pteronisculus'' is an extinct genus of prehistoric ray-finned fish that lived during the Early Triassic and Middle Triassic epochs of the Triassic period. It was first described under the name "''Glaucolepis''" by Erik Stensiö in 1921 and w ...
'', ''
Rhadinichthys ''Rhadinichthys'' is an extinct genus of prehistoric bony fish. It is known from several species that lived in the Late Devonian epoch (geology), epoch, the Carboniferous period (geology), period and the Cisuralian epoch (early Permian) in what ...
'') was recovered in the
cladistic analysis Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived chara ...
by Lund et al. This clade, coined Palaeoniscimorpha, is also used in subsequent publications. Recent cladistic analyses also recovered
clades A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, t ...
containing several genera that have historically been grouped within Palaeonisciformes, while excluding others. Due to the delicate nature of fossils of ray-finned fishes and the incomplete knowledge of several taxa (especially with regard to the internal cranial anatomy), there is still no consensus about the evolutionary relationships of several early actinopterygians previously grouped within Palaeonisciformes.


Classification

The following list includes
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
that have been referred to Palaeonisciformes (or
Palaeoniscidae Palaeoniscidae is an extinct family of ray-finned fishes (Actinopterygii) ascribed to the order Palaeonisciformes. The family includes the genus '' Palaeoniscum'' and potentially other Palaeozoic and Mesozoic early actinopterygian genera. The ...
, respectively), usually because of superficial resemblance with ''
Palaeoniscum ''Palaeoniscum'' (from el, παλαιός , 'ancient' and el, ὀνίσκος 'cod-fish' or 'woodlouse') is an extinct genus of ray-finned fish from the Permian period (Guadalupian-Lopingian) of Europe (England, Germany, Czech Republic?, Turk ...
freieslebeni''. Many of these species are poorly known and have never been included in any
cladistic analysis Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived chara ...
. Their inclusion in Palaeonisciformes (or Palaeoniscidae) is in most cases doubtful and requires confirmation by cladistic studies. Which taxa should be included in Palaeonisciformes ''sensu stricto'' (or Palaeoniscidae ''sensu stricto'') and which ones moved to other
orders Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
or
families Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideall ...
, respecitively, is a matter of ongoing research. * Order †Palaeonisciformes Hay, 1902 ''sensu stricto'' alaeoniscida Moy-Thomas & Miles, 1971ref name="mikko">
** Family †Palaeoniscidae Vogt, 1852 *** Genus ?†'' Agecephalichthys'' Wade, 1935 **** Species †''Agecephalichthys granulatus'' Wade, 1935 *** Genus ?†'' Atherstonia'' Woodward, 18989 'Broometta''_Chabakov,_1927.html" ;"title="Broometta.html" ;"title="'Broometta">'Broometta'' Chabakov, 1927">Broometta.html" ;"title="'Broometta">'Broometta'' Chabakov, 1927**** Species †''Atherstonia scutata'' Woodward, 1889 [''Atherstonia cairncrossi'' Broom, 1913; ''
Amblypterus ''Amblypterus'' (from el, ἀμβλύς , 'blunt' and el, πτερόν 'wing' or 'fin') is an extinct genus of ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise ...
capensis'' Broom, 1913; ''Broometta cairncrossi'' Chabakov, 1927] **** Species †''Atherstonia minor'' Woodward, 1893 *** Genus ?†''Cryphaeiolepis'' Traquair, 1881 **** Species †''Cryphaeiolepis scutata'' Traquair, 1881 *** Genus ?†'' Cteniolepidotrichia'' Poplin & Su, 1992 **** Species †''Cteniolepidotrichia turfanensis'' Poplin & Su, 1992 *** Genus †'' Dicellopyge'' Brough, 1931 **** Species †''Dicellopyge macrodentata'' Brough, 1931 **** Species †''Dicellopyge lissocephalus'' Brough, 1931 *** Genus ?†'' Duwaichthys'' Liu ''et al.'', 1990 **** Species †''Duwaichthys mirabilis'' Liu ''et al.'', 1990 *** Genus ?†'' Ferganiscus'' Sytchevskaya & Yakolev, 1999 **** Species †''Ferganiscus osteolepis'' Sytchevskaya & Yakolev, 1999 *** Genus †''
Gyrolepis ''Gyrolepis'' is an extinct genus of prehistoric ray-finned fish from the Middle-Late Triassic epochs in what is now Europe. It is known both from complete specimens and isolated skeletal elements, such as scales or teeth. See also * Prehist ...
'' Agassiz, 1833 non Kade, 1858 **** Species †''G. albertii'' Agassiz, 1833 **** Species †''G. gigantea'' Agassiz, 1833 **** Species †''G. maxima'' Agassiz, 1833 **** Species †''G. quenstedti'' Dames, 1888 **** Species †''G. tenuistriata'' Agassiz, 1833 *** Genus †''
Gyrolepidoides ''Gyrolepidoides'' is an extinct genus of prehistoric bony fish that lived during the Late Triassic epoch in what is now Mendoza, Argentina. See also * Prehistoric fish * List of prehistoric bony fish A ''list'' is any set of items in a ro ...
'' Cabrera, 1944 **** Species †''G. creyanus'' Schaeffer, 1955 **** Species †''G. cuyanus'' Cabrera, 1944 **** Species †''G. multistriatus'' Rusconi, 1948 *** Genus ?†'' Palaeoniscinotus'' Rohon, 1890 **** Species †''P. czekanowskii'' Rohon, 1890 *** Genus †''
Palaeoniscum ''Palaeoniscum'' (from el, παλαιός , 'ancient' and el, ὀνίσκος 'cod-fish' or 'woodlouse') is an extinct genus of ray-finned fish from the Permian period (Guadalupian-Lopingian) of Europe (England, Germany, Czech Republic?, Turk ...
'' de Blainville, 1818 'Palaeoniscus''_Agassiz,_1833_non_Von_Meyer,_1858;_''Palaeoniscas.html" ;"title="Palaeoniscus.html" ;"title="'Palaeoniscus">'Palaeoniscus'' Agassiz, 1833 non Von Meyer, 1858; ''Palaeoniscas">Palaeoniscus.html" ;"title="'Palaeoniscus">'Palaeoniscus'' Agassiz, 1833 non Von Meyer, 1858; ''Palaeoniscas'' Rzchak, 1881; ''Eupalaeoniscus'' Rzchak, 1881; ''Palaeomyzon'' Weigelt, 1930; ''Geomichthys'' Sauvage, 1888] **** Species †''P. angustum'' (Rzehak, 1881) [''Palaeoniscas angustus'' Rzehak, 1881] **** Species †''P. antipodeum'' (Egerton, 1864) 'Palaeoniscus antipodeus'' Egerton, 1864**** Species †''P. antiquum'' Williams, 1886 **** Species †''P. arenaceum'' Berger, 1832 **** Species †''P. capense'' (Bloom, 1913) 'Palaeoniscus capensis'' Bloom, 1913**** Species †''P. comtum'' (Agassiz, 1833) 'Palaeoniscus comtus'' Agassiz, 1833**** Species †''P. daedalium'' Yankevich & Minich, 1998 **** Species †''P. devonicum'' Clarke, 1885 **** Species †''P. elegans'' (Sedgwick, 1829) 'Palaeoniscus elegans'' Sedgwick, 1829**** Species †''P. freieslebeni'' de Blainville, 1818 'Eupalaeoniscus freieslebeni'' (de Brainville, 1818); ''Palaeoniscus freieslebeni'' (de Brainville, 1818)**** Species †''P. hassiae'' (Jaekel, 1898) 'Galeocerdo contortus hassiae'' Jaekel, 1898; ''Palaeomyzon hassiae'' (Jaekel, 1898)**** Species †''P. kasanense'' Geinitz & Vetter, 1880 **** Species †''P. katholitzkianum'' (Rzehak, 1881) 'Palaeoniscas katholitzkianus'' Rzehak, 1881**** Species †''P. landrioti'' (le Sauvage, 1890) 'Palaeoniscus landrioti'' le Sauvage, 1890**** Species †''P. longissimum'' (Agassiz, 1833) 'Palaeoniscus longissimus'' Agassiz, 1833**** Species †''P. macrophthalmum'' (McCoy, 1855) 'Palaeoniscus macrophthalmus'' McCoy, 1855**** Species †''P. magnum'' (Woodward, 1937) 'Palaeoniscus magnus'' Woodward, 1937**** Species †''P. moravicum'' (Rzehak, 1881) 'Palaeoniscas moravicus'' Rzehak, 1881**** Species †''P. promtu'' (Rzehak, 1881) 'Palaeoniscas promtus'' Rzehak, 1881**** Species †''P. reticulatum'' Williams, 1886 **** Species †''P. scutigerum'' Newberry, 1868 **** Species †''P. vratislavensis'' (Agassiz, 1833) 'Palaeoniscus vratislavensis'' Agassiz, 1833*** Genus †''
Palaeothrissum ''Palaeothrissum'' is an extinct genus of prehistoric bony fish. See also * Prehistoric fish The evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates develop ...
'' de Blainville, 1818 **** Species †''P. elegans'' Sedgwick, 1829 **** Species †''P. macrocephalum'' de Blainville, 1818 **** Species †''P. magnum'' de Blainville, 1818 *** Genus ?†'' Shuniscus'' Su, 1983 **** Species †''Shuniscus longianalis'' Su, 1983 *** Genus ?†'' Suchonichthys'' Minich, 2001 **** Species †''Suchonichthys molini'' Minich, 2001 *** Genus ?†'' Trachelacanthus'' Fischer De Waldheim, 1850 **** Species †''Trachelacanthus stschurovskii'' Fischer De Waldheim, 1850 *** Genus ?†'' Triassodus'' Su, 1984 **** Species †''Triassodus yanchangensis'' Su, 1984 *** Genus ?†'' Turfania'' Liu & Martínez, 1973 **** Species †''T. taoshuyuanensis'' Liu & Martínez, 1973 **** Species †''T. varta'' Wang, 1979 *** Genus ?†'' Turgoniscus'' Jakovlev, 1968 **** Species †''Turgoniscus reissi'' Jakovlev, 1968 *** Genus ?†'' Weixiniscus'' Su & Dezao, 1994 **** Species †''Weixiniscus microlepis'' Su & Dezao, 1994 *** Genus ?†'' Xingshikous'' Liu, 1988 **** Species †''Xingshikous xishanensis'' Liu, 1988 *** Genus ?†'' Yaomoshania'' Poplin ''et al.'', 1991 **** Species †''Yaomoshania minutosquama'' Poplin ''et al.'', 1991


Other families attributed to Palaeonisciformes

This list includes
families Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideall ...
that at one time or another were placed in the order Palaeonisciformes. The species included in these families are often poorly known, and a close relationship with the family Palaeoniscidae is therefore doubtful unless confirmed by cladistic analyses. These families are therefore better treated as Actinopterygii ''
incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...
'' for the time being. The
evolutionary relationships Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation t ...
of early actinopterygians is a matter of ongoing studies. * † Acropholidae Kazantseva-Selezneva, 1977 * † Atherstoniidae Gardiner, 1969 * † Brazilichthyidae Cox & Hutchinson, 1991 * † Centrolepididae Gardier, 1960 * †
Coccolepididae Coccolepididae is an extinct family of ray-finned fish, known from the Early Jurassic to Early Cretaceous, most of which were originally referred to the type genus ''Coccolepis.'' They had a widespread distribution, being found in North and South ...
Berg, 1940 corrig. * † Commentryidae Gardiner, 1963 * † Cryphiolepididae MoyThomas, 1939 corrig. * † Dwykiidae Gardiner, 1969 * † Holuridae Moy-Thomas, 1939 * † Igornichthyidae Heyler, 1977 * † Irajapintoseidae Beltan, 1978 * † Monesedeiphidae Beltan, 1989 * † Moythomasiidae Kazantseva, 1971 * † Rhabdolepididae Gardiner, 1963 * † Stegotrachelidae Gardiner, 1963 * † Thrissonotidae Berg, 1955 * † Tienshaniscidae Lu & Chen, 2010 * † Turseodontidae Bock, 1959 corrig. * † Uighuroniscidae Jin, 1996 * † Urosthenidae Woodward, 1931


Timeline of genera

''
Andreolepis hedei ''Andreolepis'' is an extinct genus of prehistoric fish, which lived around 420 million years ago. It was described by Walter Gross in 1968 based on scales found in the Hemse Formation in Gotland, Sweden. It is placed in the monogeneric f ...
'', previously grouped within Palaeonisciformes, has proven so far to be the earliest-known actinopterygiian, living around 420 million years ago (
Late Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozo ...
in
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
,
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
,
Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, a ...
, and
Latvia Latvia ( or ; lv, Latvija ; ltg, Latveja; liv, Leţmō), officially the Republic of Latvia ( lv, Latvijas Republika, links=no, ltg, Latvejas Republika, links=no, liv, Leţmō Vabāmō, links=no), is a country in the Baltic region of ...
. Actinopterygians underwent an extensive diversification during the
Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
, after the end-
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...
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Early Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early ...
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Early Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early ...
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Late Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, ...
from: -199.6 till: -175.6 color:earlyjurassic text: E. from: -175.6 till: -161.2 color:middlejurassic text: M. from: -161.2 till: -145.5 color:latejurassic text: L. from: -145.5 till: -99.6 color:earlycretaceous text:
Early Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early ...
from: -99.6 till: -66 color:latecretaceous text:
Late Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, ...
bar:eratop from: -542 till: -488.3 color:cambrian text:
Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...
from: -488.3 till: -443.7 color:ordovician text:
Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start ...
from: -443.7 till: -416 color:silurian text:
Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozo ...
from: -416 till: -359.2 color:devonian text:
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...
from: -359.2 till: -299 color:carboniferous text:
Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
from: -299 till: -251 color:permian text:
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...
from: -251 till: -199.6 color:triassic text:
Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...
from: -199.6 till: -145.5 color:jurassic text:
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
from: -145.5 till: -66 color:cretaceous text:
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
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Andreolepis ''Andreolepis'' is an extinct genus of prehistoric fish, which lived around 420 million years ago. It was described by Walter Gross in 1968 based on scales found in the Hemse Formation in Gotland, Sweden. It is placed in the monogeneric family ...
color:llandovery bar:NAM2 from: -422.9 till: -418.7 text: Naxilepis color:llandovery bar:NAM3 from: -422.9 till: -416 text: Lophosteus color:llandovery bar:NAM4 from: -422.9 till: -397.5 text:
Ligulalepis ''Ligulalepis'' is an extinct genus of prehistoric bony fish. ''Ligulalepis'' was first described from isolated scales found in the Taemas-Wee jasper limestones of New South Wales (Early Devonian age) by Dr Hans-Peter Schultze (1968) and further ...
color:earlydevonian bar:NAM5 from: -416 till: -397.5 text: Dialiipina color:earlydevonian bar:NAM6 from: -416 till: -359.2 text: Osorioichthys color:middledevonian bar:NAM7 from: -397.5 till: -391.8 text: Stegotrachelus color:middledevonian bar:NAM8 from: -397.5 till: -385.3 text: Orvikuina color:middledevonian bar:NAM9 from: -397.5 till: -359.2 text:
Moythomasia ''Moythomasia'' (named for James Alan Moy-Thomas) is an extinct genus of early ray-finned fish from the Devonian period of Europe and Australia. ''Moythomasia'' was a small freshwater fish, long. It had relatively large eyes, presumably to fin ...
color:latedevonian bar:NAM10 from: -385.3 till: -374.5 text: Mimia color:latedevonian bar:NAM11 from: -364.3 till: -359.2 text: Tegeolepis color:mississippian bar:NAM12 from: -359.2 till: -345.3 text: Whiteichthys color:mississippian bar:NAM13 from: -359.2 till: -345.3 text: Watsonichthyes color:mississippian bar:NAM14 from: -359.2 till: -345.3 text: Wardichthys color:mississippian bar:NAM15 from: -359.2 till: -345.3 text: Styracopterus color:mississippian bar:NAM16 from: -359.2 till: -345.3 text: Protamblyptera color:mississippian bar:NAM17 from: -359.2 till: -345.3 text:
Platysomus ''Platysomus'' (from el, πλατύς , 'broad' and el, σῶμα 'body') is an extinct genus of ray-finned fish that lived in the Carboniferous and Permian periods. Fossils have been found worldwide. ''Platysomus'' was about long, and ...
color:mississippian bar:NAM18 from: -359.2 till: -345.3 text: Paramesolepis color:mississippian bar:NAM19 from: -359.2 till: -345.3 text: Paragonatodus color:mississippian bar:NAM20 from: -359.2 till: -345.3 text:
Paradrydenius ''Paradrydenius'' is an extinct genus of prehistoric bony fish that lived during the Tournaisian stage of the Mississippian age, Mississippian epoch. See also * Prehistoric fish * List of prehistoric bony fish References

Palaeoniscifor ...
color:mississippian bar:NAM21 from: -359.2 till: -345.3 text: Nematoptychius color:mississippian bar:NAM22 from: -359.2 till: -345.3 text: Mesolepis color:mississippian bar:NAM23 from: -359.2 till: -345.3 text: Gonatodus color:mississippian bar:NAM24 from: -359.2 till: -345.3 text:
Elonichthys ''Elonichthys'' is an extinct genus of prehistoric bony fish. The genus is represented by several species from Carboniferous and Permian of Europe, Greenland, South Africa, and North America. Species See also * Prehistoric fish The evo ...
color:mississippian bar:NAM25 from: -359.2 till: -345.3 text: Drydenius color:mississippian bar:NAM26 from: -359.2 till: -345.3 text:
Cosmoptychius ''Cosmoptychius'' is an extinct genus of prehistoric bony fish that lived during the Tournaisian stage of the Mississippian epoch. See also * Prehistoric fish * List of prehistoric bony fish A ''list'' is any set of items in a row. List or ...
color:mississippian bar:NAM27 from: -359.2 till: -345.3 text:
Cornuboniscus ''Cornuboniscus'' is an extinct genus of prehistoric bony fish that lived during the Tournaisian age of the Mississippian epoch ( Carboniferous) in what is now Cornwall, United Kingdom. Etymology The genus ''Cornubonisus'' was named after th ...
color:mississippian bar:NAM28 from: -359.2 till: -345.3 text: Cheirodopsis color:mississippian bar:NAM29 from: -359.2 till: -345.3 text:
Brachypareion ''Brachypareion'' is an extinct genus of prehistoric bony fish that lived during the Tournaisian stage of the Mississippian epoch. See also * Prehistoric fish * List of prehistoric bony fish A ''list'' is any set of items in a row. List or ...
color:mississippian bar:NAM30 from: -359.2 till: -345.3 text:
Acrolepis ''Acrolepis'' (Ancient Greek for "tip scale") is an extinct genus of prehistoric bony fish that lived from the Tournaisian stage of the Mississippian (early Carboniferous) to the late Permian epoch. Some species from the Early Triassic of Tas ...
color:mississippian bar:NAM31 from: -359.2 till: -318.1 text: Sundayichthys color:mississippian bar:NAM32 from: -359.2 till: -318.1 text: Strepeoschema color:mississippian bar:NAM33 from: -359.2 till: -318.1 text: Phanerosteon color:mississippian bar:NAM34 from: -359.2 till: -318.1 text:
Kentuckia ''Kentuckia'' is an extinct genus of prehistoric bony fish. Taxonomy * Family Kentuckiidae Gardiner 1993 ** Genus ''Kentuckia'' Rayner 1951 *** ''K. deani'' (Eastman 1907) 'Rhadinichthys deani'' Eastman 1907*** ''K. hlavini'' Dunkle 1964 See ...
color:mississippian bar:NAM35 from: -359.2 till: -318.1 text: Canobius color:mississippian bar:NAM36 from: -359.2 till: -318.1 text: Bendenius color:mississippian bar:NAM37 from: -359.2 till: -318.1 text: Aetheretmon color:mississippian bar:NAM38 from: -359.2 till: -299 text:
Rhadinichthys ''Rhadinichthys'' is an extinct genus of prehistoric bony fish. It is known from several species that lived in the Late Devonian epoch (geology), epoch, the Carboniferous period (geology), period and the Cisuralian epoch (early Permian) in what ...
color:mississippian bar:NAM39 from: -359.2 till: -299 text:
Cycloptychius ''Cycloptychius'' is an extinct genus of prehistoric bony fish that existed in the Late Devonian epoch and the Carboniferous period in what is now France, Russia and the United Kingdom. See also * Prehistoric fish * List of prehistoric bony f ...
color:mississippian bar:NAM40 from: -345.3 till: -328.3 text: Willomorichthys color:mississippian bar:NAM41 from: -345.3 till: -328.3 text: Dwykia color:mississippian bar:NAM42 from: -345.3 till: -318.1 text: Borichthys color:mississippian bar:NAM43 from: -318.1 till: -323.63 text: Wendyichths color:mississippian bar:NAM44 from: -318.1 till: -323.63 text: Cyranorhis color:mississippian bar:NAM45 from: -318.1 till: -318.1 text:
Paramblypterus ''Paramblypterus'' is an extinct genus of bony fish. This taxon would often fall under predation from Paleozoic tetrapods such as ''Sclerocephalus'' See also * Prehistoric fish * List of prehistoric bony fish A ''list'' is any set of item ...
color:mississippian bar:NAM46 from: -318.1 till: -270.6 text: Coccocephalichthys color:pennsylvanian bar:NAM47 from: -308.23 till: -306.5 text: Nozamichthys color:pennsylvanian bar:NAM48 from: -308.23 till: -306.5 text: Illiniichthys color:pennsylvanian bar:NAM49 from: -308.23 till: -306.5 text:
Haplolepis ''Haplolepis'' is an extinct genus of prehistoric ray-finned fish that lived during the late Moscovian stage of the Pennsylvanian epoch. See also * Prehistoric fish * List of prehistoric bony fish A ''list'' is any set of items in a row. Li ...
color:earlypermian bar:NAM50 from: -299 till: -294.6 text: Charleuxia color:earlypermian bar:NAM51 from: -299 till: -270.6 text: Uydenia color:earlypermian bar:NAM52 from: -299 till: -270.6 text:
Palaeothrissum ''Palaeothrissum'' is an extinct genus of prehistoric bony fish. See also * Prehistoric fish The evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates develop ...
color:earlypermian bar:NAM53 from: -299 till: -270.6 text: Eigilia color:earlypermian bar:NAM54 from: -280 till: -270.6 text: Westollia color:earlypermian bar:NAM55 from: -280 till: -270.6 text: Igornella color:earlypermian bar:NAM56 from: -280 till: -270.6 text:
Decazella ''Decazella'' is an extinct genus of prehistoric bony fish that lived during the Gzhelian ( Stephanian) age ( Carboniferous period) in what is now Occitania, France (near Decazeville). See also * Prehistoric fish The evolution of fish b ...
color:earlypermian bar:NAM57 from: -280 till: -270.6 text:
Burbonella ''Bourbonnella'' is an extinct genus of prehistoric freshwater bony fish that lived during the late Mississippian (geology), Mississippian (Carboniferous) and Asselian (Cisuralian/early Permian epoch) in what is now Burgundy (Autun, France), Rh ...
color:earlypermian bar:NAM58 from: -280 till: -270.6 text: Aedulla color:middlepermian bar:NAM59 from: -270.6 till: -260.4 text:
Palaeoniscum ''Palaeoniscum'' (from el, παλαιός , 'ancient' and el, ὀνίσκος 'cod-fish' or 'woodlouse') is an extinct genus of ray-finned fish from the Permian period (Guadalupian-Lopingian) of Europe (England, Germany, Czech Republic?, Turk ...
color:middlepermian bar:NAM60 from: -270.6 till: -260.4 text:
Boreolepis ''Boreolepis'' is an extinct genus of prehistoric bony fish that lived during the Wuchiapingian age (Lopingian/late Permian epoch) in what is now Clavering Island, Greenland, and Vologda Oblast, Russia. See also * Prehistoric fish * List of ...
color:latepermian bar:NAM61 from: -260 till: -251 text: Trachelacanthus color:latepermian bar:NAM62 from: -260 till: -251 text: Tienshaniscus color:latepermian bar:NAM63 from: -260 till: -251 text: Sinoniscus color:latepermian bar:NAM64 from: -260 till: -251 text: Rhabdolepis color:latepermian bar:NAM65 from: -260 till: -251 text: Paralogoniscus color:latepermian bar:NAM66 from: -260 till: -251 text: Neuburgella color:latepermian bar:NAM67 from: -260 till: -251 text: Korutichthys color:latepermian bar:NAM68 from: -260 till: -251 text: Inichthys color:latepermian bar:NAM69 from: -260 till: -251 text: Gardinerichthys color:latepermian bar:NAM70 from: -260 till: -251 text:
Eurynotoides ''Eurynotoides'' is an extinct genus of prehistoric bony fish. See also * Prehistoric fish * List of prehistoric bony fish A ''list'' is any set of items in a row. List or lists may also refer to: People * List (surname) Organizations ...
color:latepermian bar:NAM71 from: -260 till: -251 text: Chichia color:latepermian bar:NAM72 from: -260 till: -251 text:
Amblypterus ''Amblypterus'' (from el, ἀμβλύς , 'blunt' and el, πτερόν 'wing' or 'fin') is an extinct genus of ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise ...
color:latepermian bar:NAM73 from: -260 till: -251 text:
Amblypterina ''Amblypterina'' is an extinct genus of prehistoric ray-finned fish. See also * Prehistoric fish * List of prehistoric bony fish References

Prehistoric ray-finned fish genera Palaeonisciformes {{Palaeonisciformes-stub ...
color:latepermian bar:NAM74 from: -260 till: -228 text: Urosthenes color:earlytriassic bar:NAM75 from: -251 till: -245 text:
Stichopterus ''Stichopterus'' is an extinct genus of chondrostean ray-finned fish that lived during the Early Cretaceous epoch in Asia. It has been found in Russia (Murtoi Formation) and Mongolia. The type species, ''Stichopterus woodwardi'', was named an ...
color:earlytriassic bar:NAM76 from: -251 till: -245 text: Sakamenichthys color:earlytriassic bar:NAM77 from: -251 till: -245 text:
Pteronisculus ''Pteronisculus'' is an extinct genus of prehistoric ray-finned fish that lived during the Early Triassic and Middle Triassic epochs of the Triassic period. It was first described under the name "''Glaucolepis''" by Erik Stensiö in 1921 and w ...
color:earlytriassic bar:NAM78 from: -251 till: -245 text:
Helichthys ''Helichthys'' is an extinct genus of prehistoric bony fish that lived during the Early Triassic epoch (geology), epoch in what is now South Africa. Although several species of ''Helichthys'' have been erected (''H. elegans'', ''H. stegopyge'', ...
color:earlytriassic bar:NAM79 from: -251 till: -245 text: Evenkia color:earlytriassic bar:NAM80 from: -251 till: -245 text: Dictyopype color:earlytriassic bar:NAM81 from: -251 till: -245 text: Daedalichthys color:earlytriassic bar:NAM82 from: -251 till: -245 text: Broovalia color:earlytriassic bar:NAM83 from: -251 till: -245 text: Atopocephala color:earlytriassic bar:NAM84 from: -251 till: -228 text: Boreosomus color:earlytriassic bar:NAM85 from: -251 till: -175.6 text:
Birgeria ''Birgeria'' is a genus of carnivorous marine ray-finned fish from the Triassic period. ''Birgeria'' had a global distribution. Fossils were found in Madagascar, Spitsbergen, Germany, Switzerland, Italy, Slovenia, China, Russia, Canada and N ...
color:earlytriassic bar:NAM86 from: -249.7 till: -199.6 text:
Gyrolepis ''Gyrolepis'' is an extinct genus of prehistoric ray-finned fish from the Middle-Late Triassic epochs in what is now Europe. It is known both from complete specimens and isolated skeletal elements, such as scales or teeth. See also * Prehist ...
color:middletriassic bar:NAM87 from: -245 till: -237 text: Dorsolepis color:middletriassic bar:NAM88 from: -245 till: -237 text: Caruichthys color:middletriassic bar:NAM89 from: -245 till: -228 text:
Gyrolepidoides ''Gyrolepidoides'' is an extinct genus of prehistoric bony fish that lived during the Late Triassic epoch in what is now Mendoza, Argentina. See also * Prehistoric fish * List of prehistoric bony fish A ''list'' is any set of items in a ro ...
color:middletriassic bar:NAM90 from: -245 till: -228 text: Aegicephalichthys color:latetriassic bar:NAM91 from: -203.6 till: -199.6 text: Scanilepis color:latetriassic bar:NAM92 from: -203.6 till: -199.6 text:
Fukangichthys ''Fukangichthys'' is an extinct genus of prehistoric bony fish that lived during the Rhaetian stage of the Late Triassic epoch. See also * Prehistoric fish * List of prehistoric bony fish A ''list'' is any set of items in a row. List or lists ...
color:earlyjurassic bar:NAM93 from: -199.6 till: -196.5 text: Browneichthys color:earlyjurassic bar:NAM94 from: -199.6 till: -194.2 text:
Cosmolepis ''Cosmolepis'' is an extinct genus of prehistoric bony fish. See also * Prehistoric fish * List of prehistoric bony fish A ''list'' is any set of items in a row. List or lists may also refer to: People * List (surname) Organizations ...
color:earlyjurassic bar:NAM95 from: -196.5 till: -189.6 text: Plesiococcolepis color:earlyjurassic bar:NAM96 from: -196.5 till: -189.6 text:
Centrolepis ''Centrolepis'' is a genus of small herbaceous plants in the family Restionaceae known as thorn grass scales, with about 25 species native to Australia, New Zealand, New Guinea, and south-east Asia as far north as Hainan Dao. APG III system class ...
color:earlyjurassic bar:NAM97 from: -196.5 till: -175.6 text:
Chondrosteus ''Chondrosteus'' is a genus of extinct actinopterygian (ray-finned fish) belonging to the family Chondrosteidae. It lived during the Sinemurian (early Early Jurassic) in what is now England. ''Chondrosteus'' is remotely related to sturgeons and ...
color:earlyjurassic bar:NAM98 from: -196.5 till: -140.2 text:
Coccolepis ''Coccolepis'' is an extinct genus of prehistoric ray-finned fish in the family Coccolepididae. Originally including most species within the family, it is now restricted to two species from the Late Jurassic Solnhofen Limestone of Germany. The ...
color:earlyjurassic bar:NAM99 from: -183 till: -180.53 text:
Strongylosteus ''Strongylosteus'' is an extinct genus of prehistoric ray-finned fish that lived during the early Toarcian age of the Early Jurassic epoch.Hennig, E. (1925). ''Chondrosteus Hindenburgi'' Pomp.---Ein «Stör» des württembergischen Ölschiefers ...
color:earlyjurassic bar:NAM100 from: -183 till: -164.7 text:
Gyrosteus ''Gyrosteus'' is an extinct genus of very large ray-finned fish belonging to the family Chondrosteidae. It comprises the type species, ''Gyrosteus mirabilis'', which lived during the early Toarcian (Late Early Jurassic) in what is now northern E ...
color:middlejurassic bar:NAM101 from: -167.7 till: -161.2 text:
Hulettia ''Hulettia'' is an extinct genus of ray-finned fish found in the Morrison Formation in the western United States, measuring approximately three to four inches in length. This fish genus contains one species, ''H. americana''. Its body was covere ...
color:latejurassic bar:NAM102 from: -150.8 till: -149.03 text: Songanella color:earlycretaceous bar:NAM103 from: -145.5 till: -130 text:
Psilichthys ''Psilichthys'' is an extinct genus of prehistoric bony fish from Eumeralla Formation, the Lower Cretaceous epoch of what is now Victoria, Australia, known from single species ''P. selwyni''. This is the first Mesozoic fossil vertebrate named ...
color:latecretaceous bar:NAM104 from: -83.5 till: -70.6 text:
Asarotus ''Asarotus'' is an extinct genus of prehistoric bony fish that lived during the Campanian. References

Palaeonisciformes Late Cretaceous fish Late Cretaceous fish of North America Prehistoric ray-finned fish genera {{Cretaceous-fish-s ...
PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 bar:period from: -542 till: -513 color:earlycambrian text:
Early Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early ...
from: -513 till: -501 color:middlecambrian text: M. from: -501 till: -488.3 color:latecambrian text: L. from: -488.3 till: -471.8 color:earlyordovician text: E. from: -471.8 till: -460.9 color:middleordovician text: M. from: -460.9 till: -443.7 color:lateordovician text: L. from: -443.7 till: -428.2 color:llandovery text: L. from: -428.2 till: -422.9 color:wenlock text: W. from: -422.9 till: -418.7 color:ludlow text: L. from: -418.7 till: -416 color:pridoli text: P from: -416 till: -397.5 color:earlydevonian text: E. from: -397.5 till: -385.3 color:middledevonian text: M. from: -385.3 till: -359.2 color:latedevonian text: L. from: -359.2 till: -318.1 color:mississippian text: Miss. from: -318.1 till: -299 color:pennsylvanian text: Penn. from: -299 till: -270.6 color:earlypermian text:
Early Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early ...
from: -270.6 till: -260.4 color:middlepermian text: M. from: -260.4 till: -251 color:latepermian text: L. from: -251 till: -245 color:earlytriassic text: E. from: -245 till: -228 color:middletriassic text: Mid. from: -228 till: -199.6 color:latetriassic text:
Late Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, ...
from: -199.6 till: -175.6 color:earlyjurassic text: E. from: -175.6 till: -161.2 color:middlejurassic text: M. from: -161.2 till: -145.5 color:latejurassic text: L. from: -145.5 till: -99.6 color:earlycretaceous text:
Early Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early ...
from: -99.6 till: -66 color:latecretaceous text:
Late Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, ...
bar:era from: -542 till: -488.3 color:cambrian text:
Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...
from: -488.3 till: -443.7 color:ordovician text:
Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start ...
from: -443.7 till: -416 color:silurian text:
Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozo ...
from: -416 till: -359.2 color:devonian text:
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...
from: -359.2 till: -299 color:carboniferous text:
Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
from: -299 till: -251 color:permian text:
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...
from: -251 till: -199.6 color:triassic text:
Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...
from: -199.6 till: -145.5 color:jurassic text:
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
from: -145.5 till: -66 color:cretaceous text:
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...


References


External links

* * *
Palaeonisciformes
at
University of Bristol , mottoeng = earningpromotes one's innate power (from Horace, ''Ode 4.4'') , established = 1595 – Merchant Venturers School1876 – University College, Bristol1909 – received royal charter , type ...
* {{Taxonbar, from=Q144538 Prehistoric ray-finned fish orders Paraphyletic groups