Owls In Art
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Owls are birds from the
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
Strigiformes (), which includes over 200 species of mostly solitary and
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers adapted for silent flight. Exceptions include the diurnal northern hawk-owl and the gregarious burrowing owl. Owls hunt mostly small
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s, insects, and other birds, although a few species specialize in hunting fish. They are found in all regions of the Earth except the polar ice caps and some remote islands. Owls are divided into two families: the true (or typical) owl family, Strigidae, and the barn-owl family, Tytonidae. A group of owls is called a "parliament".


Anatomy

Owls possess large, forward-facing eyes and ear-holes, a
hawk Hawks are bird of prey, birds of prey of the family Accipitridae. They are widely distributed and are found on all continents except Antarctica. * The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks and others. Th ...
-like beak, a flat face, and usually a conspicuous circle of feathers, a facial disc, around each eye. The feathers making up this disc can be adjusted to sharply focus sounds from varying distances onto the owls' asymmetrically placed ear cavities. Most birds of prey have eyes on the sides of their heads, but the stereoscopic nature of the owl's forward-facing eyes permits the greater sense of
depth perception Depth perception is the ability to perceive distance to objects in the world using the visual system and visual perception. It is a major factor in perceiving the world in three dimensions. Depth perception happens primarily due to stereopsis an ...
necessary for low-light hunting. Although owls have binocular vision, their large eyes are fixed in their sockets—as are those of most other birds—so they must turn their entire heads to change views. As owls are farsighted, they are unable to clearly see anything within a few centimetres of their eyes. Caught prey can be felt by owls with the use of filoplumes—hairlike feathers on the beak and feet that act as "feelers". Their far vision, particularly in low light, is exceptionally good. Owls can rotate their heads and necks as much as 270°. Owls have 14 neck vertebrae compared to seven in humans, which makes their necks more flexible. They also have adaptations to their circulatory systems, permitting rotation without cutting off blood to the brain: the foramina in their vertebrae through which the vertebral arteries pass are about 10 times the diameter of the artery, instead of about the same size as the artery as in humans; the vertebral arteries enter the cervical vertebrae higher than in other birds, giving the vessels some slack, and the carotid arteries unite in a very large
anastomosis An anastomosis (, plural anastomoses) is a connection or opening between two things (especially cavities or passages) that are normally diverging or branching, such as between blood vessels, leaf#Veins, leaf veins, or streams. Such a connection m ...
or junction, the largest of any bird's, preventing blood supply from being cut off while they rotate their necks. Other anastomoses between the carotid and vertebral arteries support this effect. The smallest owl—weighing as little as and measuring some —is the elf owl (''Micrathene whitneyi'').Konig, Claus; Welck, Friedhelm and Jan-Hendrik Becking (1999) ''Owls: A Guide to the Owls of the World'', Yale University Press, . Around the same diminutive length, although slightly heavier, are the lesser known long-whiskered owlet (''Xenoglaux loweryi'') and Tamaulipas pygmy owl (''Glaucidium sanchezi''). The largest owls are two similarly sized eagle owls; the Eurasian eagle-owl (''Bubo bubo'') and Blakiston's fish owl (''Bubo blakistoni''). The largest females of these species are long, have a wing span, and weigh . Different species of owls produce different sounds; this distribution of calls aids owls in finding mates or announcing their presence to potential competitors, and also aids
ornithologist Ornithology is a branch of zoology that concerns the "methodological study and consequent knowledge of birds with all that relates to them." Several aspects of ornithology differ from related disciplines, due partly to the high visibility and th ...
s and birders in locating these birds and distinguishing species. As noted above, their facial discs help owls to funnel the sound of prey to their ears. In many species, these discs are placed asymmetrically, for better directional location. Owl
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
is generally
cryptic Cryptic may refer to: In science: * Cryptic species complex, a group of species that are very difficult to distinguish from one another * Crypsis, the ability of animals to blend in to avoid observation * Cryptic era, earliest period of the Earth ...
, although several species have facial and head markings, including face masks, ear tufts, and brightly colored irises. These markings are generally more common in species inhabiting open habitats, and are thought to be used in signaling with other owls in low-light conditions.


Sexual dimorphism

Sexual dimorphism is a physical difference between males and females of a species. Female owls are typically larger than the males. The degree of size dimorphism varies across multiple populations and species, and is measured through various traits, such as wing span and body mass. One theory suggests that selection has led males to be smaller because it allows them to be efficient foragers. The ability to obtain more food is advantageous during breeding season. In some species, female owls stay at their nest with their eggs while it is the responsibility of the male to bring back food to the nest. If food is scarce, the male first feeds himself before feeding the female. Small birds, which are agile, are an important source of food for owls. Male burrowing owls have been observed to have longer wing chords than females, despite being smaller than females. Furthermore, owls have been observed to be roughly the same size as their prey. This has also been observed in other predatory birds, which suggests that owls with smaller bodies and long wing chords have been selected for because of the increased agility and speed that allows them to catch their prey. Another popular theory suggests that females have not been selected to be smaller like male owls because of their sexual roles. In many species, female owls may not leave the nest. Therefore, females may have a larger mass to allow them to go for a longer period of time without starving. For example, one hypothesized sexual role is that larger females are more capable of dismembering prey and feeding it to their young, hence female owls are larger than their male counterparts. A different theory suggests that the size difference between male and females is due to sexual selection: since large females can choose their mate and may violently reject a male's sexual advances, smaller male owls that have the ability to escape unreceptive females are more likely to have been selected. If the character is stable, there can be different optimums for both sexes. Selection operates on both sexes at the same time; therefore it is necessary to explain not only why one of the sexes is relatively bigger, but also why the other sex is smaller. If owls are still evolving toward smaller bodies and longer wing chords, according to V. Geodakyan's Evolutionary Theory of Sex, males should be more advanced on these characters. Males are viewed as an evolutionary vanguard of a population, and sexual dimorphism on the character, as an evolutionary “distance” between the sexes. “Phylogenetic rule of sexual dimorphism” states that if there exists a sexual dimorphism on any character, then the evolution of this trait goes from the female form toward the male one.


Hunting adaptations

All owls are carnivorous birds of prey and live on diets of insects, small rodents and lagomorphs. Some owls are also specifically adapted to hunt fish. They are very adept in hunting in their respective environments. Since owls can be found in nearly all parts of the world and across a multitude of ecosystems, their hunting skills and characteristics vary slightly from species to species, though most characteristics are shared among all species.


Flight and feathers

Most owls share an innate ability to fly almost silently and also more slowly in comparison to other birds of prey. Most owls live a mainly nocturnal lifestyle and being able to fly without making any noise gives them a strong advantage over prey alert to the slightest sound in the night. A silent, slow flight is not as necessary for diurnal and crepuscular owls given that prey can usually see an owl approaching. Owls’ feathers are generally larger than the average birds’ feathers, have fewer radiates, longer pennulum, and achieve smooth edges with different rachis structures. Serrated edges along the owl's remiges bring the flapping of the wing down to a nearly silent mechanism. The serrations are more likely reducing aerodynamic disturbances, rather than simply reducing noise. The surface of the flight feathers is covered with a velvety structure that absorbs the sound of the wing moving. These unique structures reduce noise frequencies above 2 kHz, making the sound level emitted drop below the typical hearing spectrum of the owl's usual prey and also within the owl's own best hearing range. This optimizes the owl's ability to silently fly to capture prey without the prey hearing the owl first as it flies, and to hear any noise the prey makes. It also allows the owl to monitor the sound output from its flight pattern. The feather adaption that allows silent flight means that barn owl feathers are not waterproof. To retain the softness and silent flight, the barn owl cannot use the
preen oil The uropygial gland, informally known as the preen gland or the oil gland, is a bilobed sebaceous gland possessed by the majority of birds used to distribute the gland's oil through the plumage by means of preening. It is located dorsally at the ...
or powder dust that other species use for waterproofing. In wet weather, they cannot hunt and this may be disastrous during the breeding season. Barn owls are frequently found drowned in livestock drinking troughs, since they land to drink and bathe, but are unable to climb out. Owls can struggle to keep warm, because of their lack of waterproofing, so large numbers of downy feathers help them to retain body heat.


Vision

Eyesight is a particular characteristic of the owl that aids in nocturnal prey capture. Owls are part of a small group of birds that live nocturnally, but do not use echolocation to guide them in flight in low-light situations. Owls are known for their disproportionally large eyes in comparison to their skulls. An apparent consequence of the evolution of an absolutely large eye in a relatively small skull is that the eye of the owl has become tubular in shape. This shape is found in other so-called nocturnal eyes, such as the eyes of strepsirrhine primates and bathypelagic fishes. Since the eyes are fixed into these sclerotic tubes, they are unable to move the eyes in any direction. Instead of moving their eyes, owls swivel their heads to view their surroundings. Owls' heads are capable of swiveling through an angle of roughly 270°, easily enabling them to see behind them without relocating the torso. This ability keeps bodily movement at a minimum, thus reduces the amount of sound the owl makes as it waits for its prey. Owls are regarded as having the most frontally placed eyes among all avian groups, which gives them some of the largest binocular fields of vision. Owls are farsighted and cannot focus on objects within a few centimetres of their eyes. These mechanisms are only able to function due to the large-sized retinal image. Thus, the primary nocturnal function in the vision of the owl is due to its large posterior nodal distance; retinal image brightness is only maximized to the owl within secondary neural functions. These attributes of the owl cause its nocturnal eyesight to be far superior to that of its average prey.


Hearing

Owls exhibit specialized hearing functions and ear shapes that also aid in hunting. They are noted for asymmetrical ear placements on the skull in some genera. Owls can have either internal or external ears, both of which are asymmetrical. Asymmetry has not been reported to extend to the middle or internal ear of the owl. Asymmetrical ear placement on the skull allows the owl to pinpoint the location of its prey. This is especially true for strictly nocturnal species such as the barn owls ''Tyto'' or Tengmalm's owl. With ears set at different places on its skull, an owl is able to determine the direction from which the sound is coming by the minute difference in time that it takes for the sound waves to penetrate the left and right ear
The Hearing of the Barn Owl
The owl turns its head until the sound reaches both ears at the same time, at which point it is directly facing the source of the sound. This time difference between ears is about 30 microseconds. Behind the ear openings are modified, dense feathers, densely packed to form a facial ruff, which creates an anterior-facing, concave wall that cups the sound into the ear structure. This facial ruff is poorly defined in some species, and prominent, nearly encircling the face, in other species. The facial disk also acts to direct sound into the ears, and a downward-facing, sharply triangular beak minimizes sound reflection away from the face. The shape of the facial disk is adjustable at will to focus sounds more effectively. The prominences above a great horned owl's head are commonly mistaken as its ears. This is not the case; they are merely feather tufts. The ears are on the sides of the head in the usual location (in two different locations as described above).


Talons

While the auditory and visual capabilities of the owl allow it to locate and pursue its prey, the talons and beak of the owl do the final work. The owl kills its prey using these talons to crush the skull and knead the body. The crushing power of an owl's talons varies according to prey size and type, and by the size of the owl. The burrowing owl (''Athene cunicularia''), a small, partly insectivorous owl, has a release force of only 5 N. The larger
barn owl The barn owl (''Tyto alba'') is the most widely distributed species of owl in the world and one of the most widespread of all species of birds, being found almost everywhere except for the polar and desert regions, Asia north of the Himalaya ...
(''Tyto alba'') needs a force of 30 N to release its prey, and one of the largest owls, the great horned owl (''Bubo virginianus'') needs a force over 130 N to release prey in its talons. An owl's talons, like those of most birds of prey, can seem massive in comparison to the body size outside of flight. The Tasmanian masked owl has some of the proportionally longest talons of any bird of prey; they appear enormous in comparison to the body when fully extended to grasp prey. An owl's claws are sharp and curved. The family Tytonidae has inner and central toes of about equal length, while the family Strigidae has an inner toe that is distinctly shorter than the central one. These different morphologies allow efficiency in capturing prey specific to the different environments they inhabit.


Beak

The beak of the owl is short, curved, and downward-facing, and typically hooked at the tip for gripping and tearing its prey. Once prey is captured, the scissor motion of the top and lower bill is used to tear the tissue and kill. The sharp lower edge of the upper bill works in coordination with the sharp upper edge of the lower bill to deliver this motion. The downward-facing beak allows the owl's field of vision to be clear, as well as directing sound into the ears without deflecting sound waves away from the face.


Camouflage

The coloration of the owl's plumage plays a key role in its ability to sit still and blend into the environment, making it nearly invisible to prey. Owls tend to mimic the coloration and sometimes the texture patterns of their surroundings, the barn owl being an exception. The snowy owl (''Bubo scandiacus'') appears nearly bleach-white in color with a few flecks of black, mimicking their snowy surroundings perfectly, while the speckled brown plumage of the tawny owl (''Strix aluco'') allows it to lie in wait among the deciduous woodland it prefers for its habitat. Likewise, the
mottled wood-owl The mottled wood owl (''Strix ocellata'') is a species of large owl found in India. They are found in gardens and thin deciduous forests adjacent to dry thorn forests or farmland. They are easily detected by their distinctive tremulous eerie call ...
(''Strix ocellata'') displays shades of brown, tan and black, making the owl nearly invisible in the surrounding trees, especially from behind. Usually, the only tell-tale sign of a perched owl is its vocalizations or its vividly colored eyes.


Behavior

Most owls are
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
, actively hunting their prey in darkness. Several types of owls are
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of daylig ...
—active during the twilight hours of dawn and dusk; one example is the pygmy owl (''Glaucidium''). A few owls are active during the day, also; examples are the burrowing owl (''Speotyto cunicularia'') and the short-eared owl (''Asio flammeus''). Much of the owls' hunting strategy depends on stealth and surprise. Owls have at least two adaptations that aid them in achieving stealth. First, the dull coloration of their feathers can render them almost invisible under certain conditions. Secondly, serrated edges on the leading edge of owls' remiges muffle an owl's wing beats, allowing an owl's flight to be practically silent. Some fish-eating owls, for which silence has no evolutionary advantage, lack this adaptation. An owl's sharp beak and powerful talons allow it to kill its prey before swallowing it whole (if it is not too big). Scientists studying the diets of owls are helped by their habit of regurgitating the indigestible parts of their prey (such as bones, scales, and fur) in the form of pellets. These "owl pellets" are plentiful and easy to interpret, and are often sold by companies to schools for dissection by students as a lesson in biology and ecology.


Breeding and reproduction

Owl eggs typically have a white color and an almost spherical shape, and range in number from a few to a dozen, depending on species and the particular season; for most, three or four is the more common number. In at least one species, female owls do not mate with the same male for a lifetime. Female burrowing owls commonly travel and find other mates, while the male stays in his territory and mates with other females.


Evolution and systematics

Recent phylogenetic studies place owls within the
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
Telluraves, most closely related to the Accipitrimorphae and the Coraciimorphae,H Kuhl, C Frankl-Vilches, A Bakker, G Mayr, G Nikolaus, S T Boerno, S Klages, B Timmermann, M Gahr (2020
An unbiased molecular approach using 3’UTRs resolves the avian family-level tree of life
''Molecular Biology and Evolution''. https://doi.org/10.1093/molbev/msaa191
although the exact placement within Telluraves is disputed.Prum, R.O. ''et al.'' (2015

Nature 526, 569–573.
See below cladogram: Cladogram of Telluraves relationships based on Braun & Kimball (2021)Braun, E.L. & Kimball, R.T. (2021) Data types and the phylogeny of Neoaves. ''Birds'', 2(1), 1-22; https://doi.org/10.3390/birds2010001 Some 220 to 225 extant species of owls are known, subdivided into two families: 1. true owls or typical owls family ( Strigidae) and 2. barn-owls family ( Tytonidae). Some entirely extinct families have also been erected based on fossil remains; these differ much from modern owls in being less specialized or specialized in a very different way (such as the terrestrial
Sophiornithidae The Sophiornithidae (literally "wisdom birds") are an extinct family of chicken-sized predatory birds that lived from the Paleocene to the Eocene periods of the Cenozoic, and were found primarily in Europe, and are thought to be primitive owls. ...
). The Paleocene genera '' Berruornis'' and '' Ogygoptynx'' show that owls were already present as a distinct lineage some 60–57 million years ago (Mya), hence, possibly also some 5 million years earlier, at the extinction of the non-avian dinosaurs. This makes them one of the oldest known groups of non- Galloanserae landbirds. The supposed " Cretaceous owls" ''
Bradycneme ''Bradycneme'' (meaning "ponderous leg") is a genus of theropod dinosaur from the Maastrichtian-age Upper Cretaceous Sânpetru Formation of the Hațeg Basin, Transylvania, Romania. The genus contains a single species, ''Bradycneme draculae'', k ...
'' and ''
Heptasteornis ''Heptasteornis'' is the name given to a potentially dubious genus of alvarezsaurid dinosaur from the Late Cretaceous. The type (and only known) species is ''Heptasteornis andrewsi'', described as a presumed gigantic prehistoric owl in 1975. It w ...
'' are apparently non- avialan
maniraptors Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to ''Ornithomimus velox''. It contains the major subgroups Avialae, Deinonychosauria, Oviraptorosa ...
. During the
Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...
, the Strigiformes radiated into ecological niches now mostly filled by other groups of birds. The owls as known today, though, evolved their characteristic morphology and
adaptation In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the po ...
s during that time, too. By the early
Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. ...
, the other lineages had been displaced by other bird orders, leaving only barn owls and typical owls. The latter at that time was usually a fairly generic type of (probably earless) owl similar to today's North American spotted owl or the European tawny owl; the diversity in size and ecology found in typical owls today developed only subsequently. Around the Paleogene-Neogene boundary (some 25 Mya), barn owls were the dominant group of owls in southern Europe and adjacent Asia at least; the distribution of fossil and present-day owl lineages indicates that their decline is contemporary with the evolution of the different major lineages of true owls, which for the most part seems to have taken place in Eurasia. In the Americas, rather, an expansion of immigrant lineages of ancestral typical owls occurred. The supposed fossil
heron The herons are long-legged, long-necked, freshwater and coastal birds in the family Ardeidae, with 72 recognised species, some of which are referred to as egrets or bitterns rather than herons. Members of the genera ''Botaurus'' and ''Ixobrychus ...
s ''"Ardea" perplexa'' (Middle Miocene of Sansan, France) and ''"Ardea" lignitum'' (Late Pliocene of Germany) were more probably owls; the latter was apparently close to the modern genus '' Bubo''. Judging from this, the Late Miocene remains from France described as ''"Ardea" aureliensis'' should also be restudied. The Messelasturidae, some of which were initially believed to be
basal Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''. Science * Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure * Basal (medicine), a minimal level that is nec ...
Strigiformes, are now generally accepted to be diurnal birds of prey showing some convergent evolution toward owls. The taxa often united under '' Strigogyps'' were formerly placed in part with the owls, specifically the Sophiornithidae; they appear to be Ameghinornithidae instead. For fossil species and paleosubspecies of extant taxa, see the genus and species articles. For a full list of extant and recently extinct owls, see the article List of owl species. Unresolved and basal forms (all fossil) * '' Berruornis'' (Late Paleocene of France) basal? Sophornithidae? * Strigiformes gen. et sp. indet. (Late Paleocene of Zhylga, Kazakhstan) * '' Primoptynx'' (Early Eocene of Wyoming, U.S.) * ''
Palaeoglaux ''Palaeoglaux'' is a genus of fossil owls from the Eocene epoch. The two known species are ''P. perrierensis'' from the Upper Eocene of Quercy, France, and ''P. artophoron'' from the Middle Eocene Messel shales, Germany.Mourer-Chauviré, Cécile ...
'' (Middle-Late Eocene of West-Central Europe) own family Palaeoglaucidae or Strigidae? * ''Palaeobyas'' (Late Eocene/Early Oligocene of Quercy, France) Tytonidae? Sophiornithidae? * ''Palaeotyto'' (Late Eocene/Early Oligocene of Quercy, France) Tytonidae? Sophiornithidae? * Strigiformes ''gen. et spp. indet.'' (Early Oligocene of Wyoming, U.S.)


Ogygoptyngidae

* '' Ogygoptynx'' (Middle/Late Paleocene of Colorado, U.S.)


Protostrigidae

* '' Eostrix'' (Early Eocene of United States, Europe, and Mongolia). ''E. gulottai'' is the smallest known fossil (or living) owl. * '' Minerva'' (Middle – Late Eocene of western U.S.) formerly ''Protostrix'', includes ''"Aquila" ferox'', ''"Aquila" lydekkeri'', and ''"Bubo" leptosteus'' * ''Oligostrix'' (mid-Oligocene of Saxony, Germany)


Sophiornithidae

* ''Sophiornis''


Tytonidae

* Genus '' Tyto'' – the barn owls, grass owls, and masked owls, stand up to tall; some 15 extant species and possibly one recently extinct * Genus '' Phodilus'' – the bay owls, two to three extant species and possibly one recently extinct Fossil genera * ''Nocturnavis'' (Late Eocene/Early Oligocene) includes ''"Bubo" incertus'' * ''Selenornis'' (Late Eocene/Early Oligocene) – includes ''"Asio" henrici'' * ''Necrobyas'' (Late Eocene/Early Oligocene – Late Miocene) includes ''"Bubo" arvernensis'' and ''Paratyto'' * ''Prosybris'' (Early Oligocene? – Early Miocene) Placement unresolved * Tytonidae ''gen. et sp. indet.'' "TMT 164" (Middle Miocene) – ''Prosybris''?


Strigidae

* Genus '' Aegolius'' – the saw-whet owls, four species * Genus '' Asio'' – the eared owls, eight species * Genus '' Athene'' – two to four species (depending on whether the genera ''Speotyto'' and ''Heteroglaux'' are included or not) * Genus '' Bubo'' – the horned owls, eagle-owls and fish-owls; paraphyletic with the genera ''Nyctea'', ''Ketupa'', and ''Scotopelia'', some 25 species * Genus '' Glaucidium'' – the pygmy owls, about 30–35 species * Genus '' Gymnasio'' – the Puerto Rican owl * Genus '' Gymnoglaux'' – the bare-legged owl or Cuban screech-owl * Genus ''
Lophostrix The crested owl (''Lophostrix cristata'') is a species of owl in the family Strigidae. It is the only species (monotypic) in the genus ''Lophostrix''. It is a resident bird and occurs in Central America and northern South America. It is a medium- ...
'' – the crested owl * Genus '' Jubula'' – the maned owl * Genus ''
Megascops Screech owls are typical owls (Strigidae) belonging to the genus ''Megascops'' with 23 living species. For most of the 20th century, this genus was merged with the Old World scops owls in ''Otus'', but nowadays it is again considered separat ...
'' – the screech owls, some 20 species * Genus '' Micrathene'' – the elf owl * Genus '' Ninox'' – the Australasian hawk-owls or boobooks, some 20 species * Genus '' Otus'' – the scops owls; probably
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
, about 45 species * Genus '' Pseudoscops'' – the Jamaican owl * Genus ''
Psiloscops The flammulated owl (''Psiloscops flammeolus'') is a small migratory North American owl in the family Strigidae. It is the only species placed in the genus ''Psiloscops''. Taxonomy The flammulated owl was formally described in 1852 by the Germ ...
'' – the flammulated owl * Genus '' Ptilopsis'' – the white-faced owls, two species * Genus ''
Pulsatrix ''Pulsatrix'' is a genus of owl in the family Strigidae. They are called spectacled owls because of their prominent facial pattern. The genus contains the following species: * Spectacled owl, '' Pulsatrix perspicillata'' * Tawny-browed owl, '' ...
'' – the spectacled owls, three species * Genus ''
Strix Strix may refer to: * Strix (mythology), a legendary creature of ancient Roman mythology * ''Strix'' (bird), a genus of large "earless" wood-owls * Strix Ltd, manufacturer of kettle controls, thermostats and water boiling elements for domestic ap ...
'' – the earless owls, about 15 species, including four previously assigned to ''Ciccaba'' * Genus ''
Surnia ''Surnia'' is a genus of owl that contains a single living species, the northern hawk-owl ''(Surnia ulula)''. Two fossil species are known as well; ''Surnia capeki'' and ''Surnia robusta'', both from the Plio-Pleistocene of Europe Europe ...
'' – the northern hawk-owl * Genus '' Taenioptynx'' - the collared owlet * Genus '' Uroglaux'' – the Papuan hawk-owl * Genus '' Xenoglaux'' – the long-whiskered owlet Extinct genera * Genus '' Grallistrix'' – the stilt-owls, four species;
prehistoric Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of ...
* Genus '' Ornimegalonyx'' – the Caribbean giant owls, one to two species;
prehistoric Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of ...
Fossil genera * ''Mioglaux'' (Late Oligocene? – Early Miocene of West-Central Europe) – includes ''"Bubo" poirreiri'' * ''Intutula'' (Early/Middle – ?Late Miocene of Central Europe) – includes ''"Strix/Ninox" brevis'' * ''Alasio'' (Middle Miocene of Vieux-Collonges, France) – includes ''"Strix" collongensis'' * ''
Oraristrix The La Brea owl (''Oraristix brea'') is an extinct owl reported from the upper Pleistocene asphalt deposits of the La Brea Tar Pits in Los Angeles, California. It was first described in 1933 by Hildegarde Howard as ''Strix brea'', but this extinc ...
'' – the Brea owl (Late Pleistocene) Placement unresolved * ''"Otus/Strix" wintershofensis'': fossil (Early/Middle Miocene of Wintershof West, Germany) – may be close to extant genus ''Ninox'' * ''"Strix" edwardsi'' – fossil (Middle/Late? Miocene) * ''"Asio" pygmaeus'' – fossil (Early Pliocene of Odessa, Ukraine) * Strigidae gen. et sp. indet. UMMP V31030 (Late Pliocene) – ''Strix/Bubo''? * the Ibizan owl, Strigidae ''gen. et sp. indet.'' –
prehistoric Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of ...


Symbolism and mythology


African cultures

Among the Kikuyu of Kenya, it was believed that owls were harbingers of death. If one saw an owl or heard its hoot, someone was going to die. In general, owls are viewed as harbingers of bad luck, ill health, or death. The belief is widespread even today.


Asia

In Mongolia, the owl is regarded as a benign omen. In one story,
Genghis Khan ''Chinggis Khaan'' ͡ʃʰiŋɡɪs xaːŋbr />Mongol script: ''Chinggis Qa(gh)an/ Chinggis Khagan'' , birth_name = Temüjin , successor = Tolui (as regent)Ögedei Khan , spouse = , issue = , house = Borjigin , ...
was hiding from enemies in a small coppice when an owl roosted in the tree above him, which caused his pursuers to think no man could be hidden there. In modern Japan, owls are regarded as lucky and are carried in the form of a talisman or charm.
Hootum Pyanchar Naksha ''Hootum Pyanchar Naksha'' (literally "Sketches by a Watching Owl") is a book by Kaliprasanna Singha (1841–1870), first published in 1861. The book has been influential in Bengali literature. It reflects the tension that arose within several are ...
by Kaliprasanna Singha (1841–1870), first published in 1861, is a book of social commentaries influential in
Bengali literature Bengali literature ( bn, বাংলা সাহিত্য, Bangla Sahityô) denotes the body of writings in the Bengali language and which covers Old Bengali, Middle- Bengali and Modern Bengali with the changes through the passage of time ...
. The name literally means "Sketches by a Watching Owl".


Sumerian and ancient Semitic cultures

In Sumerian, Akkadian, and Babylonian culture, the owl was associated with Lilith. This association also occurs in the Bible (in some translations) in Isaiah 34:14.


Ancient European and modern Western culture

The
modern West Kevin Costner & Modern West is an American country rock band founded and fronted by actor Kevin Costner. He began a worldwide tour with the band in October 2007, which included shows in Istanbul and Rome. It also performed at NASCAR Sprint Cup Ser ...
generally associates owls with wisdom and vigilance. This link goes back at least as far as Ancient Greece, where Athens, noted for art and scholarship, and Athena, Athens' patron goddess and the goddess of wisdom, had the owl as a symbol.Deacy, Susan, and Villing, Alexandra (2001). Athena in the Classical World. Koninklijke Brill NV, Leiden, The Netherlands, . Marija Gimbutas traces veneration of the owl as a goddess, among other birds, to the culture of Old Europe, long pre-dating Indo-European cultures.
T. F. Thiselton-Dyer The Reverend Thomas Firminger Thiselton-Dyer, MA, Oxon (25 July 1848 – 14 July 1923) was a son of William George Thiselton-Dyer, physician and of Catherine Jane, née Firminger. He was educated at King's College School and at Pembroke College, O ...
, in his 1883 ''Folk-lore of Shakespeare'', says that "from the earliest period it has been considered a bird of ill-omen," and Pliny tells us how, on one occasion, even Rome itself underwent a lustration, because one of them strayed into the Capitol. He represents it also as a funereal bird, a monster of the night, the very abomination of humankind. Virgil describes its death howl from the top of the temple by night, a circumstance introduced as a precursor of Dido's death. Ovid, too, constantly speaks of this bird's presence as an evil omen; and indeed the same notions respecting it may be found among the writings of most of the ancient poets." A list of "omens drear" in
John Keats John Keats (31 October 1795 – 23 February 1821) was an English poet of the second generation of Romantic poets, with Lord Byron and Percy Bysshe Shelley. His poems had been in publication for less than four years when he died of tuberculo ...
' ''
Hyperion Hyperion may refer to: Greek mythology * Hyperion (Titan), one of the twelve Titans * ''Hyperion'', a byname of the Sun, Helios * Hyperion of Troy or Yperion, son of King Priam Science * Hyperion (moon), a moon of the planet Saturn * ''Hyp ...
'' includes the "gloom-bird's hated screech." Pliny the Elder reports that owls' eggs were commonly used as a hangover cure. One of the etymologies offered for the name of the German folk hero Till Eulenspiegel is that it means "Mirror for Owls". File:Aryballos owl Louvre CA1737.jpg, An owl-shaped protocorinthian aryballos, c. 640 BCE, from Greece File:Itálica Owl.jpg, A Roman owl mosaic from Italica, Spain File:Hispano-Moresque - Plate - Walters 481099.jpg, A Manises plate, c. 1535. A fantastical owl wearing a crown, a characteristic Manises design during the first half of the 16th century File:Albrecht Dürer - The Little Owl - WGA7367.jpg, ''The Little Owl'', 1506, by
Albrecht Dürer Albrecht Dürer (; ; hu, Ajtósi Adalbert; 21 May 1471 – 6 April 1528),Müller, Peter O. (1993) ''Substantiv-Derivation in Den Schriften Albrecht Dürers'', Walter de Gruyter. . sometimes spelled in English as Durer (without an umlaut) or Due ...
File:Natungram Dolls, West Bengal.jpg, Wooden Owls of Natungram, West Bengal, India. The wooden owl is an integral part of an ancient and indigenous tradition and art form in Bengal along with its auspicious association with Goddess of wealth, Laxmi.


Hinduism

In Hinduism, an owl is the '' vahana'' (mount) of the goddess Lakshmi, especially in the eastern region of India. Owl is considered a symbol of wealth, prosperity, wisdom, good luck, and Fortune. This is the reason why Owl is seen with Godden Lakshmi, who is also the goddess the fortune, wealth, and prosperity. The Goddess Lakshmi is known to have a White Barn Owl as her ''vahana''. At the same time, owls are also associated with evil times in Hinduism. At times,
Chamunda Chamunda (Sanskrit: चामुण्डा, ISO-15919: Cāmuṇḍā), also known as Chamundeshwari, Chamundi or Charchika, is a fearsome form of Chandi, the Hindu Divine Mother Shakti and is one of the seven Matrikas (mother goddesses).W ...
(fearsome form of Chandi) is depicted seated on an owl, her vahana (mount or vehicle). Hindus believed that owls are messengers of death.


Native American cultures

People often allude to the reputation of owls as bearers of supernatural danger when they tell misbehaving children, "the owls will get you", and in most Native American folklore, owls are a symbol of death. According to the
Apache The Apache () are a group of culturally related Native American tribes in the Southwestern United States, which include the Chiricahua, Jicarilla, Lipan, Mescalero, Mimbreño, Ndendahe (Bedonkohe or Mogollon and Nednhi or Carrizaleño an ...
and Seminole tribes, hearing owls hooting is considered the subject of numerous " bogeyman" stories told to warn children to remain indoors at night or not to cry too much, otherwise the owl may carry them away. In some tribal legends, owls are associated with spirits of the dead, and the bony circles around an owl's eyes are said to comprise the fingernails of apparitional humans. Sometimes owls are said to carry messages from beyond the grave or deliver supernatural warnings to people who have broken tribal taboos. The Aztecs and the Maya, along with other natives of Mesoamerica, considered the owl a symbol of death and destruction. In fact, the Aztec god of death, Mictlantecuhtli, was often depicted with owls. There is an old saying in Mexico that is still in use: ''Cuando el tecolote canta, el indio muere'' ("When the owl cries/sings, the Indian dies"). The '' Popol Vuh'', a Mayan religious text, describes owls as messengers of Xibalba (the Mayan "Place of Fright"). The belief that owls are messengers and harbingers of the dark powers is also found among the Hočągara (Winnebago) of Wisconsin. When in earlier days the Hočągara committed the sin of killing enemies while they were within the sanctuary of the chief's lodge, an owl appeared and spoke to them in the voice of a human, saying, "From now on, the Hočągara will have no luck." This marked the beginning of the decline of their tribe. An owl appeared to
Glory of the Morning Glory of the Morning (died c. 1832) was the first woman ever described in the written history of Wisconsin, and the only known female chief of the Hocąk (Winnebago) nation. At least one source has rendered her name as ''Hopokoekau'', which is a co ...
, the only female chief of the Hočąk nation, and uttered her name. Soon after, she died. According to the culture of the
Hopi The Hopi are a Native American ethnic group who primarily live on the Hopi Reservation in northeastern Arizona, United States. As of the 2010 census, there are 19,338 Hopi in the country. The Hopi Tribe is a sovereign nation within the Unite ...
, a Uto-Aztec tribe, taboos surround owls, which are associated with sorcery and other evils. The Ojibwe tribes, as well as their Aboriginal Canadian counterparts, used an owl as a symbol for both evil and death. In addition, they used owls as a symbol of very high status of spiritual leaders of their spirituality. The Pawnee tribes viewed owls as the symbol of protection from any danger within their realms. The Puebloan peoples associated owls with Skeleton Man, the god of death and the spirit of fertility. The Yakama tribes use an owl as a totem, to guide where and how forests and natural resources are useful with management.


Rodent control

Encouraging natural predators to control rodent population is a natural form of pest control, along with excluding food sources for rodents. Placing a nest box for owls on a property can help control rodent populations (one family of hungry barn owls can consume more than 3,000 rodents in a nesting season) while maintaining the naturally balanced food chain.


Attacks on humans

Although humans and owls frequently live together in harmony, there have been incidents when owls have attacked humans. For example, in January 2013, a man from Inverness, Scotland suffered heavy bleeding and went into shock after being attacked by an owl, which was likely a
eagle-owl The American (North and South America) horned owls and the Old World eagle-owls make up the genus ''Bubo'', at least as traditionally described. The genus name ''Bubo'' is Latin for the Eurasian eagle-owl. This genus contains 19 species that ar ...
. The photographer
Eric Hosking Eric John Hosking Order of the British Empire, OBE (2 October 1909 – 22 February 1991) was an English photographer noted for his Nature photography, bird photography. Life and career Eric Hosking OBE, Hon FRPS, was a pioneering wildlife photo ...
lost his left eye after attempting to photograph a tawny owl, which inspired the title of his 1970 autobiography, ''An Eye for a Bird''.


Conservation issues

Almost all owls are listed in Appendix II of the international CITES treaty (the Convention on Illegal Trade in Endangered Species of Wild Fauna and Flora) with four species listed in Appendix I. Although owls have long been hunted, a 2008 news story from Malaysia indicates that the magnitude of owl poaching may be on the rise. In November 2008, TRAFFIC reported the seizure of 900 plucked and "oven-ready" owls in Peninsular Malaysia. Said Chris Shepherd, Senior Programme Officer for TRAFFIC's Southeast Asia office, "This is the first time we know of where 'ready-prepared' owls have been seized in Malaysia, and it may mark the start of a new trend in wild meat from the region. We will be monitoring developments closely." TRAFFIC commended the Department of Wildlife and National Parks in Malaysia for the raid that exposed the huge haul of owls. Included in the seizure were dead and plucked barn owls, spotted wood owls, crested serpent eagles, barred eagles, and brown wood owls, as well as 7,000 live lizards.


References


Further reading

* Calaprice, Alice & Heinrich, Bernd (1990): ''Owl in the House: A Naturalist's Diary''. Joy Street Books, Boston. . * Duncan, James (2013). ''The Complete Book of North American Owls.'' Thunder Bay Press, San Diego. . * Duncan, James (2003). ''Owls of the World''. Key Porter Books, Toronto. . * Heinrich, Bernd (1987): ''One Man's Owl''. Princeton, N.J.: Princeton University Press. . . * Johnsgard, Paul A. (2002): ''North American Owls: Biology and Natural History'', 2nd ed. Smithsonian Institution Press, Washington, D.C. . * Maslow, Jonathan Evan (1983): ''The Owl Papers'', 1st Vintage Books ed. Vintage Books, New York. . * Sibley, Charles Gald & Monroe, Burt L. Jr. (1990): ''Distribution and taxonomy of the birds of the world: A Study in Molecular Evolution''. Yale University Press, New Haven, CT.


External links


The Owl Pages

Owl Brain Atlas



World Owl Trust

Athenian Owl coins
Eurasia:
World of Owls
– Northern Ireland's only owl, bird of prey and exotic animal centre
Current Blakiston's Fish Owl Research in Russia
North America:

Oceania:

info. re Australian owls and frogmouths {{Authority control Extant Paleocene first appearances