Praeaulacidae is an extinct family of
Mesozoic
The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceo ...
parasitic wasps
Parasitoid wasps are a large group of hymenopteran superfamilies, with all but the wood wasps (Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on or in the bodies of other arthropods, sooner or later causin ...
in the suborder
Evanioidea
The Evanioidea are a small hymenopteran superfamily that includes three families, two of which (Aulacidae and Gasteruptiidae) are much more closely related to one another than they are to the remaining family, Evaniidae. The rich fossil record, h ...
. It among the earliest known families of the group and is characterised by more complete wing venation in comparison to other members of the suborder.
It has been found that Othniodellithidae is nested within Praeaulacidae via
cladistic
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...
analysis.
Genera
Taxonomy according to Jouault et al, 2020
* †''
Albiogonalys'' Nel et al. 2003
Charentese amber, France, Late Cretaceous (
Cenomanian
The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
)
*†subfamily Cretocleistogastrinae Rasnitsyn 1990
**†''
Cretocleistogaster'' Rasnitsyn 1975
Zaza Formation
The Zaza Formation is a geological formation located in Buryatia (Russia). It dates to the Lower Cretaceous period. It is Aptian in age and consists of Sandstone, sandstones, Siltstone, siltstones, Marl, marls and bituminous Shale, shales, deposi ...
,
Turga Formation, Russia, Early Cretaceous (
Aptian
The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
)
**†''
Miniwestratia'' Rasnitsyn 1990
Dzun-Bain Formation, Mongolia, Aptian
**†''
Nanowestratia'' Rasnitsyn 1990, Zaza Formation, Russia, Aptian
**†''
Sinowestratia'' Zhang and Zheng 2000
Yixian Formation
The Yixian Formation (; formerly transcribed as Yihsien Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the late Barremian and early Aptian stages of the Early Cretaceous. It is known for its ex ...
, China, Aptian
**†''
Westratia'' Jell and Duncan 1986
Tsagaantsav Formation
The Tsagaantsav Formation, Tsagantsab Formation or Tsagan-Tsab Formation (Russian: Tsagaantsav Svita) is an Early Cretaceous (Hauterivian to Barremian) geologic formation in Mongolia.[Hauterivian
The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.9 ± 2 Ma and 129.4 ± 1.5 Ma (million years ago). The Hauterivian is preceded by the Va ...]
/
Barremian
The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
)
Koonwarra fossil bed, Australia, Aptian Dzun-Bain Formation, Mongolia, Aptian Zaza Formation,
Gidarinskaya Formation, Russia, Aptian
*†subfamily Nevaniinae Zhang and Rasnitsyn 2007
** †''
Eonevania'' Rasnitsyn and Zhang 2010
Daohugou
The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China.
The formation consists of coarse conglomerates ...
, China, Middle Jurassic (
Callovian
In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 166.1 ± 4.0 Ma (million years ago) and 163.5 ± 4.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
)
** †''
Nevania'' Zhang and Rasnitsyn 2007 Daohugou, China, Callovian,
Karabastau Formation
The Karabastau Formation ( kk, Qarabastaý svıtasy) is a geological formation and lagerstätte in the Karatau Mountains of southern Kazakhstan whose strata date to the Middle to Late Jurassic. It is an important locality for insect fossils that ...
, Kazakhstan, Late Jurassic
** †''
Rasnitsevania'' Jouault et al. 2020
Burmese amber
Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The ...
, Myanmar,
Cenomanian
The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
* †subfamily Praeaulacinae Rasnitsyn 1972
** †''
Archaulacus'' Li et al. 2014 Daohugou, China, Callovian
** †''
Aulacogastrinus'' Rasnitsyn 1983 Daohugou, China, Callovian, Karabastau Formation, Kazakhstan, Callovian/Oxfordian
** †''
Eosaulacus'' Zhang and Rasnitsyn 2008 Daohugou, China, Callovian
** †''
Evanigaster'' Rasnitsyn 1972 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
** †''
Evaniops'' Rasnitsyn 1972 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
** †''
Gulgonga'' Oberprieler et al. 2012
Talbragar Fossil Bed, Australia, Late Jurassic (
Tithonian
In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
)
** †''
Habraulacus'' Li et al. 2015 Burmese amber, Myanmar, Cenomanian
** †''
Praeaulacinus'' Rasnitsyn 1972 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
** †''
Praeaulacites'' Rasnitsyn 1972 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
** †''
Praeaulacon'' Rasnitsyn 1972 Daohugou, China, Callovian, Karabastau Formation, Kazakhstan, Callovian/Oxfordian
** †''
Praeaulacops'' Rasnitsyn 1972 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
**†''
Praeaulacus'' Rasnitsyn 1972 Daohugou, China, Callovian, Karabastau Formation, Kazakhstan, Callovian/Oxfordian,
Shar Teg, Mongolia, Tithonian
** ?†''
Sinaulacogastrinus'' Zhang and Rasnitsyn 2008 Daohugou, China, Callovian
** ?†''
Sinevania'' Rasnitsyn and Zhang 2010 Daohugou, China, Callovian
*†Family Othniodellithidae Engel and Huang, 2016 (found to be nested within Praeaulacidae by cladistic analysis
)
**†''
Keradellitha'' Jouault, Maréchal, Wang & Perrichot, 2021 Burmese amber, Myanmar, Cenomanian
**†''
Othniodellitha'' Engel and Huang, 2016 Burmese amber, Myanmar, Cenomanian
**†''
Xenodellitha'' Engel, 2017, Burmese amber, Myanmar, Cenomanian
References
{{Taxonbar, from=Q21073876
Evanioidea
Prehistoric insect families