''Niassodon'' is an extinct

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

of endothiodontid

dicynodont Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typic ...

therapsid Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more ...

known from the

Late Permian Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, ...

of Niassa Province, northern

Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a country located in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi ...

. It contains a single species, ''Niassodon mfumukasi''.

Discovery

''Niassodon'' was first described and named by Rui Castanhinha, Ricardo Araújo, Luís C. Júnior, Kenneth D. Angielczyk, Gabriel G. Martins, Rui M. S. Martins, Claudine Chaouiya, Felix Beckmann and Fabian Wilde in

2013 File:2013 Events Collage V2.png, From left, clockwise: Edward Snowden becomes internationally famous for leaking classified NSA wiretapping information; Typhoon Haiyan kills over 6,000 in the Philippines and Southeast Asia; The Dhaka garment fa ...

and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...

is ''Niassodon mfumukasi''. The generic name combines the word ''Niassa'', meaning "

lake A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much large ...

" in

Chiyao Yao is a Bantu language in Africa with approximately two million speakers in Malawi, and half a million each in Tanzania and Mozambique. There are also some speakers in Zambia. In Malawi, the main dialect is Mangochi, mostly spoken around Lak ...

and the name of the northwestern province in Mozambique where ''Niassodon'' was found, with ''odontos'', meaning "tooth" in

Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...

. The specific name, ''Mfumukasi'' means "queen" in

Nyanja Chewa (also known as Nyanja, ) is a Bantu language spoken in much of Southern, Southeast and East Africa, namely the countries of Malawi , where it is an official language, and Mozambique and Zambia. The noun class prefix ''chi-'' is used for l ...

, in honor of the members of the local Nyanja matriarchal society and all Mozambican women. ''Niassodon'' is known solely from the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...

ML1620, a partial skeleton which has returned to the Museu Nacional de Geologia,

Maputo Maputo (), formerly named Lourenço Marques until 1976, is the capital, and largest city of Mozambique. Located near the southern end of the country, it is within of the borders with Eswatini and South Africa. The city has a population of 1,0 ...

, Mozambique. The holotype consists of a complete

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...

, a

mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...

, series of 19

dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal c ...

sacral Sacral may refer to: *Sacred Sacred describes something that is dedicated or set apart for the service or worship of a deity; is considered worthy of spiritual respect or devotion; or inspires awe or reverence among believers. The property ...

and caudal vertebrae, ribs, both ilia and a partial

femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates wit ...

, from a single individual. It was discovered by Ricardo Araújo and collected during a 2009 expedition in the Metangula Graben, under the supervision of Projecto PaleoMoz. ML1620 came from the K5 Formation' '''' located near Tulo, a small village situated along the Metangula-Cóbue road. The fossil bed is composed of a grey mudstone with abundant septaria-like calcareous

concretion A concretion is a hard, compact mass of matter formed by the precipitation of mineral cement within the spaces between particles, and is found in sedimentary rock or soil. Concretions are often ovoid or spherical in shape, although irregular ...

s, dating to the

References

Dicynodonts Lopingian synapsids of Africa Lopingian genus first appearances Lopingian genus extinctions Fossils of Mozambique Fossil taxa described in 2013 Articles containing video clips Anomodont genera {{anomodont-stub