Neovenatoridae is a proposed clade of
carcharodontosaurian
Allosauroidea is a superfamily or clade of theropod dinosaurs which contains four families — the Metriacanthosauridae, Allosauridae, Carcharodontosauridae, and Neovenatoridae. Allosauroids, alongside the family Megalosauroidea, were among th ...
dinosaurs uniting some primitive members of the group such as ''
Neovenator
''Neovenator'' (nee-o-ven-a-tor meaning "new hunter") is a genus of carcharodontosaurian theropod dinosaur. It is known from several skeletons found in the Early Cretaceous (Barremian~130-125 million years ago) Wessex Formation on the south coa ...
'' with the
Megaraptora
Megaraptora is a clade of carnivorous Tetanurae, tetanuran theropod dinosaurs with controversial relations to other theropods. Its Derived (phylogenetics), derived members, the Megaraptoridae are noted for their elongated hand claws and proporti ...
, a group of theropods with controversial affinities. Other studies recover megaraptorans as basal
coelurosaurs
Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs.
Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyran ...
unrelated to carcharodontosaurs. Other theropods with uncertain affinities such as ''
Gualicho
''Gualicho'' (named in reference to the gualichu) is a genus of theropod dinosaur. The type species is ''Gualicho shinyae''. It lived in what is now northern Patagonia, on what was then a South American island continent split off from the superco ...
'', ''
Chilantaisaurus
''Chilantaisaurus'' (" lizard") is a genus of large theropod dinosaur, possibly a neovenatorid or a primitive coelurosaur, from the Late Cretaceous Ulansuhai Formation of China (Turonian age, about 92 million years ago). The type species, ''C. ...
'' and ''
Deltadromeus
''Deltadromeus'' (meaning "delta runner") is a genus of theropod dinosaur from Northern Africa. It had long, unusually slender hind limbs for its size, suggesting that it was a swift runner. The skull is not known. One fossil specimen of a sin ...
'' are also sometimes included.
Classification
Phylogenetic studies conducted by Benson, Carrano and Brusatte (2010) and Carrano, Benson and Sampson (2012) recovered the group
Megaraptora
Megaraptora is a clade of carnivorous Tetanurae, tetanuran theropod dinosaurs with controversial relations to other theropods. Its Derived (phylogenetics), derived members, the Megaraptoridae are noted for their elongated hand claws and proporti ...
as members of the Neovenatoridae. This would make neovenatorids the latest-surviving allosauroids; at least one megaraptoran, ''
Orkoraptor
''Orkoraptor'' is a genus of medium-sized megaraptoran theropod dinosaur from the late Cretaceous Period of Argentina. It is known from incomplete fossil remains including parts of the skull, teeth, tail vertebrae, and a partial tibia. The speci ...
'', lived near the end of the
Mesozoic
The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretace ...
era, dating to the early
Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval ...
stage of the latest
Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
period, about 70 million years ago.
On the other hand, Novas ''et al.'' (2012), while confirming that ''
Neovenator
''Neovenator'' (nee-o-ven-a-tor meaning "new hunter") is a genus of carcharodontosaurian theropod dinosaur. It is known from several skeletons found in the Early Cretaceous (Barremian~130-125 million years ago) Wessex Formation on the south coa ...
'' was closely related to carcharodontosaurids, simultaneously found ''
Megaraptor
''Megaraptor'' () is a genus of large theropod dinosaur that lived in the ages of the Late Cretaceous. Its fossils have been discovered in the Patagonian Portezuelo Formation of Argentina, South America. Initially thought to have been a giant dr ...
'' and related genera to be
coelurosaurs
Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs.
Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyran ...
closely related to
tyrannosaurids.
However, Novas ''et al.'' subsequently found that megaraptorans lacked most of the key features in the hands of derived coelurosaurs including ''
Guanlong
''Guanlong'' (冠龍) is a genus of extinct proceratosaurid tyrannosauroid from the Late Jurassic of China. The taxon was first described in 2006 by Xu Xing ''et al.'', who found it to represent a new taxon related to ''Tyrannosaurus''. The na ...
'' and ''
Deinonychus
''Deinonychus'' ( ; ) is a genus of dromaeosaurid theropod dinosaur with one described species, ''Deinonychus antirrhopus''. This species, which could grow up to long, lived during the early Cretaceous Period, about 115–108 million y ...
''. Instead, their hands retain a number of primitive characteristics seen in basal tetanurans such as ''
Allosaurus
''Allosaurus'' () is a genus of large carnosaurian theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic epoch ( Kimmeridgian to late Tithonian). The name "''Allosaurus''" means "different lizard" alludin ...
''. Nevertheless, there are still a number of other traits that support megaraptorans as members of the Coelurosauria.
The cladogram below follows a 2016 analysis by Sebastián Apesteguía, Nathan D. Smith, Rubén Juarez Valieri, and Peter J. Makovicky based on the dataset of Carrano ''et al.'' (2012).
References
{{Taxonbar, from=Q133002
Carcharodontosauria
Prehistoric dinosaur families