Natural History Of California
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The ecology of California can be understood by dividing the state into a number of
ecoregion An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas of l ...
s, which contain distinct
ecological Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps wi ...
communities A community is a Level of analysis, social unit (a group of living things) with commonality such as place (geography), place, Norm (social), norms, religion, values, Convention (norm), customs, or Identity (social science), identity. Communiti ...
of plants and animals in a contiguous region. The ecoregions of California can be grouped into four major groups: desert ecoregions (such as the
Mojave Desert The Mojave Desert ( ; mov, Hayikwiir Mat'aar; es, Desierto de Mojave) is a desert in the rain shadow of the Sierra Nevada mountains in the Southwestern United States. It is named for the indigenous Mojave people. It is located primarily in ...
), Mediterranean ecoregions (such as the Central Valley), forested mountains (such as the
Sierra Nevada The Sierra Nevada () is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primarily ...
), and coastal forests. Different authorities define the boundaries of ecoregions somewhat differently: this article follows the definitions of the
World Wide Fund for Nature The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the Wor ...
(WWF) and the
United States Environmental Protection Agency The Environmental Protection Agency (EPA) is an independent executive agency of the United States federal government tasked with environmental protection matters. President Richard Nixon proposed the establishment of EPA on July 9, 1970; it be ...
(EPA)


Deserts

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
's high mountains block most moisture from reaching the eastern parts of the state, which are home to California's desert and xeric shrub ecoregions. The low desert of southeastern California is part of the
Sonoran desert The Sonoran Desert ( es, Desierto de Sonora) is a desert in North America and ecoregion that covers the northwestern Mexican states of Sonora, Baja California, and Baja California Sur, as well as part of the southwestern United States (in Arizona ...
ecoregion, which extends into
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
and parts of northern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
. California has two high deserts: the
Mojave desert The Mojave Desert ( ; mov, Hayikwiir Mat'aar; es, Desierto de Mojave) is a desert in the rain shadow of the Sierra Nevada mountains in the Southwestern United States. It is named for the indigenous Mojave people. It is located primarily in ...
and the
Great Basin Desert The Great Basin Desert is part of the Great Basin between the Sierra Nevada and the Wasatch Range. The desert is a geographical region that largely overlaps the Great Basin shrub steppe defined by the World Wildlife Fund, and the Central Basin a ...
. The Mojave desert ecoregion is marked by the presence of
Joshua tree ''Yucca brevifolia'' is a plant species belonging to the genus ''Yucca''. It is tree-like in habit, which is reflected in its common names: Joshua tree, yucca palm, tree yucca, and palm tree yucca. This monocotyledonous tree is native to the ar ...
s. The dry cold Great Basin desert of California consists of the
Owens Valley Owens Valley (Numic Numic is a branch of the Uto-Aztecan language family. It includes seven languages spoken by Native American peoples traditionally living in the Great Basin, Colorado River basin, Snake River basin, and southern Great Pl ...
, and is classified into
Great Basin shrub steppe The Great Basin Desert is part of the Great Basin between the Sierra Nevada and the Wasatch Range. The desert is a geographical region that largely overlaps the Great Basin shrub steppe defined by the World Wildlife Fund, and the Central Basin a ...
by the WWF, and into the
Central Basin and Range ecoregion The Great Basin Desert is part of the Great Basin between the Sierra Nevada and the Wasatch Range. The desert is a geographical region that largely overlaps the Great Basin shrub steppe defined by the World Wildlife Fund, and the Central Basin a ...
by the
EPA The Environmental Protection Agency (EPA) is an independent executive agency of the United States federal government tasked with environmental protection matters. President Richard Nixon proposed the establishment of EPA on July 9, 1970; it be ...
. The deserts in California receive between of rain per year. Plants in these deserts are brush and scrub, adapted to the low rainfall. Common plant species include
creosote bush ''Larrea tridentata'', called creosote bush and greasewood as a plant, chaparral as a medicinal herb, and ''gobernadora'' (Spanish language, Spanish for "governess") in Mexico, due to its ability to secure more water by inhibiting the growth of n ...
,
blackbrush ''Coleogyne ramosissima'' or blackbrush, is a low lying, dark grayish-green, aromatic,Turner, Raymond M. 1982. Great Basin desertscrub. In: Brown, David E., ed. Biotic communities of the American Southwest--United States and Mexico. Desert Plan ...
,
greasewood Greasewood is a common name shared by several plants: * ''Adenostoma fasciculatum'' is a plant with white flowers that is native to Oregon, Nevada, California, and northern Baja California. This shrub is one of the most widespread plants of the ...
,
saltbush Saltbush is a vernacular plant name that most often refers to ''Atriplex'', a genus of about 250 plants distributed worldwide from subtropical to subarctic regions. ''Atriplex'' species are native to Australia, North and South America, and Eurasia. ...
,
big sagebrush ''Artemisia tridentata'', commonly called big sagebrush,MacKay, Pam (2013), ''Mojave Desert Wildflowers'', 2nd ed., , p. 264. Great Basin sagebrush or (locally) simply sagebrush, is an aromatic shrub from the family Asteraceae, which grows in ari ...
,
low sagebrush ''Artemisia arbuscula'' is a North American species of sagebrush (plant), sagebrush known by the common names little sagebrush, low sagebrush, or black sagebrush. It is native to the western United States from Washington (U.S. state), Washington, ...
, and
shadscale ''Atriplex confertifolia'', the shadscale or spiny saltbush, is a species of evergreen shrub in the family Amaranthaceae, which is native to the western United States and northern Mexico Mexico (Spanish: México), officially the United Me ...
. Higher elevations have more precipitation, which allows drought-resistant trees to grow, such as
western juniper ''Juniperus occidentalis'', known as the western juniper, is a shrub or tree native to the Western United States, growing in mountains at altitudes of and rarely down to . It is listed as Least Concern on the IUCN Red List because it is a widesp ...
and
pinyon pine The pinyon or piñon pine group grows in southwestern North America, especially in New Mexico, Arizona, and Utah. The trees yield edible nuts, which are a staple food of Native Americans, and widely eaten as a snack and as an ingredient in New ...
.


Mediterranean ecoregions

The coast of California from
Monterey Bay Monterey Bay is a bay of the Pacific Ocean located on the coast of the U.S. state of California, south of the San Francisco Bay Area and its major city at the south of the bay, San Jose. San Francisco itself is further north along the coast, by a ...
south to the Mexican border, and inland from
San Francisco Bay Area The San Francisco Bay Area, often referred to as simply the Bay Area, is a populous region surrounding the San Francisco, San Pablo, and Suisun Bay estuaries in Northern California. The Bay Area is defined by the Association of Bay Area Go ...
to the Sierra Nevada foothills contain California's
Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the e ...
ecoregions. This region is divided by the WWF into three
California chaparral and woodlands The California chaparral and woodlands is a terrestrial ecoregion of southwestern Oregon, northern, central, and southern California (United States) and northwestern Baja California (Mexico), located on the west coast of North America. It is a ...
ecoregions, plus the Central Valley grasslands. The EPA divides the region between the Central Valley (ecoregion 7), the Southern California chaparral (ecoregion 6), the Southern California mountains (ecoregion 8), and the Southern California coast (ecoregion 85). The WWF distinguishes between different chaparral ecoregions based on species
endemism Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
. In the south, the
California coastal sage and chaparral The California coastal sage and chaparral ( es, Salvia y chaparral costero de California) is a Mediterranean forests, woodlands, and scrub ecoregion located in southwestern California (United States) and northwestern Baja California (Mexico). It i ...
extends across the Mexican border into northwestern
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
and Los Angeles. The coastal sage ecoregion is notable for having the highest number of native bees in the United States, although much of the ecoregion is now urbanized. The
California montane chaparral and woodlands The California montane chaparral and woodlands is an ecoregion defined by the World Wildlife Fund, spanning of mountains in the Transverse Ranges, Peninsular Ranges, and Coast Ranges of southern and central California. The ecoregion is part of ...
include the
Transverse Ranges The Transverse Ranges are a group of mountain ranges of southern California, in the Pacific Coast Ranges physiographic region in North America. The Transverse Ranges begin at the southern end of the California Coast Ranges and lie within Santa B ...
north of Los Angeles as well as the
Santa Lucia Range The Santa Lucia Mountains (sæntə luˈsiːə) or Santa Lucia Range is a rugged mountain range in coastal central California, running from Carmel southeast for to the Cuyama River in San Luis Obispo County. The range is never more than from t ...
on the Central Coast. The montane chaparral consists of a mosaic of sage scrub, chaparral, and montane species, depending on altitude. The
California interior chaparral and woodlands The California interior chaparral and woodlands ecoregion covers in an elliptical ring around the California Central Valley. It occurs on hills and mountains ranging from to . It is part of the Mediterranean forests, woodlands, and scrub biom ...
form a ring around the Central Valley, covering the hills around the Bay Area as well as the foothills of the Sierra Nevada. The interior woodland ecoregion contains several endemic species, due to unique soil types such as serpentine. These chaparral ecoregions contain numerous plant communities, including oak savanna, oak woodland, conifer woodlands, chamise chaparral,
coastal sage scrub Coastal sage scrub, also known as coastal scrub, CSS, or soft chaparral, is a low scrubland plant community of the California coastal sage and chaparral subecoregion, found in coastal California and northwestern coastal Baja California. It is w ...
, and coastal grassland. These plant communities often occur as a mosaic, caused by fire. California's Central Valley was once a large
temperate grassland Temperate grasslands, savannas, and shrublands is a terrestrial biome defined by the World Wide Fund for Nature. The predominant vegetation in this biome consists of grass and/or shrubs. The climate is temperate and ranges from semi-arid to semi ...
containing native
bunchgrass Tussock grasses or bunch grasses are a group of grass species in the family Poaceae. They usually grow as singular plants in clumps, tufts, hummocks, or bunches, rather than forming a sod or lawn, in meadows, grasslands, and prairies. As perennial ...
es and
vernal pools Vernal pools, also called vernal ponds or ephemeral pools, are seasonal pools of water that provide habitat for distinctive plants and animals. They are considered to be a distinctive type of wetland usually devoid of fish, and thus allow the safe ...
.
Grizzly bear The grizzly bear (''Ursus arctos horribilis''), also known as the North American brown bear or simply grizzly, is a population or subspecies of the brown bear inhabiting North America. In addition to the mainland grizzly (''Ursus arctos horri ...
,
gray wolf The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly ...
,
tule elk The tule elk (''Cervus canadensis nannodes'') is a subspecies of elk found only in California, ranging from the grasslands and marshlands of the Central Valley to the grassy hills on the coast. The subspecies name derives from the tule (), a s ...
, and
pronghorn antelope The pronghorn (, ) (''Antilocapra americana'') is a species of artiodactyl (even-toed, hoofed) mammal indigenous to interior western and central North America. Though not an antelope, it is known colloquially in North America as the American ante ...
used to inhabit the grasslands. The native grasslands and pools have now been largely replaced by livestock ranches and farms. The
Carrizo Plain The Carrizo Plain ( Obispeño: ''tšɨłkukunɨtš'', "Place of the rabbits") is a large enclosed grassland plain, approximately long and up to across, in southeastern San Luis Obispo County, California, about northwest of Los Angeles. It con ...
, where the native grass is preserved, is referred to as the "Serengeti of California".


Forested mountains

The cooler and wetter mountains of northern California are covered by forest ecoregions. Both the WWF and the EPA divide the mountains into three ecoregions: the
Sierra Nevada The Sierra Nevada () is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primarily ...
, the
Klamath Mountains The Klamath Mountains are a rugged and lightly populated mountain range in northwestern California and southwestern Oregon in the western United States. As a mountain system within both the greater Pacific Coast Ranges and the California Coast R ...
, and the Eastern Cascades Slopes and Foothills (occurring on the
Modoc Plateau __NOTOC__ The Modoc Plateau lies in the northeast corner of California as well as parts of Oregon and Nevada. Nearly of the Modoc National Forest are on the plateau between the Medicine Lake Highlands in the west and the Warner Mountains in the ...
). The Sierra Nevada are home to half of the
vascular plant Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They al ...
species of California, with 400 species that are
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to the region. Like many mountain ranges, the plant communities of the Sierra group into
biotic zone The life zone concept was developed by C. Hart Merriam in 1889 as a means of describing areas with similar plant and animal communities. Merriam observed that the changes in these communities with an increase in latitude at a constant elevation a ...
s by altitude, because of the increasingly harsh climate as elevation increases. These biotic zones include montane forest dominated by
conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
s such as
Jeffrey pine ''Pinus jeffreyi'', also known as Jeffrey pine, Jeffrey's pine, yellow pine and black pine, is a North American pine tree. It is mainly found in California, but also in the westernmost part of Nevada, southwestern Oregon, and northern Baja Califo ...
and
Lodgepole pine ''Pinus contorta'', with the common names lodgepole pine and shore pine, and also known as twisted pine, and contorta pine, is a common tree in western North America. It is common near the ocean shore and in dry montane forests to the subalpine, ...
,
subalpine forest Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
dominated by
whitebark pine ''Pinus albicaulis'', known by the common names whitebark pine, white bark pine, white pine, pitch pine, scrub pine, and creeping pine, is a conifer tree native to the mountains of the western United States and Canada, specifically subalpine ...
, up to
alpine tundra Alpine tundra is a type of natural region or biome that does not contain trees because it is at high elevation, with an associated alpine climate, harsh climate. As the latitude of a location approaches the poles, the threshold elevation for alp ...
which cannot support trees. The Sierra are also notable for
giant sequoia ''Sequoiadendron giganteum'' (giant sequoia; also known as giant redwood, Sierra redwood, Sierran redwood, California big tree, Wellingtonia or simply big treea nickname also used by John Muir) is the sole living species in the genus ''Sequoiade ...
trees: the most massive on earth. The Klamath and Siskiyou Mountains are a notable
biodiversity hotspot A biodiversity hotspot is a biogeographic region with significant levels of biodiversity that is threatened by human habitation. Norman Myers wrote about the concept in two articles in ''The Environmentalist'' in 1988 and 1990, after which the co ...
, containing one of the four most biodiverse temperate forests in the world. The diversity is caused by the ecoregion being adjacent to a number of other ecoregions, diverse soil, and having refugia caused by isolation in the last
ice age An ice age is a long period of reduction in the temperature of Earth's surface and atmosphere, resulting in the presence or expansion of continental and polar ice sheets and alpine glaciers. Earth's climate alternates between ice ages and gree ...
. Some endemic species in the Klamath mountains are limited to only one mountain or valley. The Eastern Cascades slopes of the Modoc Plateau are characterized by a mosaic of open
ponderosa pine ''Pinus ponderosa'', commonly known as the ponderosa pine, bull pine, blackjack pine, western yellow-pine, or filipinus pine is a very large pine tree species of variable habitat native to mountainous regions of western North America. It is the ...
forest, grasslands, and shrublands. Although high, these slopes and mountains are in the rain shadow of the
Cascade Range The Cascade Range or Cascades is a major mountain range of western North America, extending from southern British Columbia through Washington and Oregon to Northern California. It includes both non-volcanic mountains, such as the North Cascades, ...
, and hence are drier and more open.


Coastal forests

The coast of California north of San Francisco contains the
Northern California coastal forests The Northern California coastal forests are a temperate coniferous forests ecoregion of coastal Northern California and southwestern Oregon. Setting The ecoregion covers , extending from just north of the California-Oregon border south, to south ...
(as defined by the WWF) and the southern section of the Coast Range ecoregion (as defined by the EPA). This ecoregion is dominated by
redwood forest ''Sequoia sempervirens'' ()''Sunset Western Garden Book,'' 1995:606–607 is the sole living species of the genus '' Sequoia'' in the cypress family Cupressaceae (formerly treated in Taxodiaceae). Common names include coast redwood, coastal ...
, containing the tallest and some of the oldest trees in the world. The redwood forests thrive in a thin belt up to wide next to the coast, where the trees are kept moist by winter rains and summer fog. The redwood forests are also notable for having the highest forest
productivity Productivity is the efficiency of production of goods or services expressed by some measure. Measurements of productivity are often expressed as a ratio of an aggregate output to a single input or an aggregate input used in a production proces ...
in the world.


See also

* * * List of California birds *
Fauna of California The fauna of the State of California may be the most diverse in the United States of America. Of the Lower 48 conterminous states, California has the greatest diversity in climate, terrain and geology in general. The state's six life zones are t ...
*
California Floristic Province The California Floristic Province (CFP) is a floristic province with a Mediterranean-type climate located on the Pacific Coast of North America with a distinctive flora similar to other regions with a winter rainfall and summer drought climate l ...
*
Geography of California California is a U.S. state on the western coast of North America. Covering an area of , California is among the most geographically diverse states. The Sierra Nevada, the fertile farmlands of the Central Valley, and the arid Mojave Desert of t ...
*
California native plants California native plants are plants that existed in California prior to the arrival of European explorers and colonists in the late 18th century. California includes parts of at least three phytochoria. The largest is the California Floristic ...
* Protected areas of California


References

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