Naashoibitosaurus
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''Naashoibitosaurus'' (from
Navajo The Navajo (; British English: Navaho; nv, Diné or ') are a Native American people of the Southwestern United States. With more than 399,494 enrolled tribal members , the Navajo Nation is the largest federally recognized tribe in the United ...
—"creek lizard") is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
hadrosaur Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which includ ...
id
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
that lived about 73 million years ago, in the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
, and was found in the Kirtland Formation of the
San Juan San Juan, Spanish for Saint John, may refer to: Places Argentina * San Juan Province, Argentina * San Juan, Argentina, the capital of that province * San Juan, Salta, a village in Iruya, Salta Province * San Juan (Buenos Aires Underground), ...
Basin in
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
. Only a partial skeleton has been found to date. It was first described as a specimen of ''Kritosaurus'' by
Jack Horner Jack Horner may refer to: *''Little Jack Horner'', a nursery rhyme People * Jack Horner (baseball) (1863–1910), American professional baseball player *Jack Horner (journalist) (1912–2005), Gordon John Horner, Minnesota sportscaster * Jack B. H ...
, and has been intertwined with ''
Kritosaurus ''Kritosaurus'' is an incompletely known genus of hadrosaurid (duck-billed) dinosaur. It lived about 74.5-66 million years ago, in the Late Cretaceous of North America. The name means "separated lizard" (referring to the arrangement of the cheek ...
'' since its description.


Description

''Naashoibitosaurus'', based as it is on a single partial skeleton ( NMMNH P-16106), is not well known in terms of anatomy. Its skull, the most thoroughly described portion, has a low
nasal Nasal is an adjective referring to the nose, part of human or animal anatomy. It may also be shorthand for the following uses in combination: * With reference to the human nose: ** Nasal administration, a method of pharmaceutical drug delivery ** ...
crest that peaks in front of the eyes, but does not strongly arch as in ''
Gryposaurus ''Gryposaurus'' (meaning "hooked-nosed (Ancient Greek, Greek ''grypos'') lizard"; sometimes incorrectly translated as "griffin (Latin ''gryphus'') lizard") was a genus of hadrosaur, duckbilled dinosaur that lived about 80 to 75 million years ...
''.


Classification

''Naashoibitosaurus'' is a
saurolophine Saurolophinae is a subfamily (biology), subfamily of hadrosaurid dinosaurs. It has since the mid-20th century generally been called the Hadrosaurinae, a group of largely non-crested hadrosaurs related to the crested sub-family Lambeosaurinae. How ...
hadrosaurid, a "flat-headed or solid-crested duckbill". It is closest to the solid-crested forms like ''
Saurolophus ''Saurolophus'' (; meaning "lizard crest") is a genus of large hadrosaurid dinosaur from the Late Cretaceous period of Asia and North America, that lived in what is now the Horseshoe Canyon and Nemegt formations about 70 million to 68 million ...
''. If it is the same as ''Kritosaurus'', ''Kritosaurus'' would be used because it is the older name, but its position vis-a-vis other duckbills would be the same.


History

Befitting a genus with a confusing
taxonomic Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
history, the name of this genus is based on an error. David Gillette and David Thomas collected the type and only known specimen from what was thought to be the Naashoibito Member of the Kirtland Formation, the youngest member of the Kirtland; this was commemorated in the name. Instead, it came from the older, late Campanian-age De-na-zin Member. Horner described the skull in 1992 as that of an
immature Mature is the adjectival form of maturity, as immature is the adjectival form of immaturity, which have several meanings. Mature or immature may also refer to: *Mature, a character from ''The King of Fighters'' series *"Mature 17+", a rating in ...
''Kritosaurus'', using it as evidence that ''Gryposaurus'' was different from ''Kritosaurus''. At the same time, Hunt and Lucas were describing the
postcrania Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated sk ...
as belonging to '' Edmontosaurus saskatchewanensis''. When the disconnect became apparent, Hunt and Lucas gave the specimen its own generic name, because the skull did not agree with an edmontosaur, and they considered ''Kritosaurus''
indeterminate Indeterminate may refer to: In mathematics * Indeterminate (variable), a symbol that is treated as a variable * Indeterminate system, a system of simultaneous equations that has more than one solution * Indeterminate equation, an equation that ha ...
and thus not usable. Their assessment was challenged in 2000 by Thomas Williamson, who found the morphological differences between it and ''Kritosaurus'' to be due to the skulls in question being from individuals of different ages. Unlike Hunt and Lucas, he considered the type skull of ''Kritosaurus'' to be diagnostic and thus could compare it to ''Naashoibitosaurus''. He returned to Horner's argument, noting as Horner had that the nasal crest in other hadrosaurines like ''
Prosaurolophus ''Prosaurolophus'' (; meaning "before ''Saurolophus''", in comparison to the later dinosaur with a similar head crest) is a genus of hadrosaurid (or duck-billed) dinosaur from the Late Cretaceous of North America. It is known from the remains of ...
'' moves back on the skull during growth. He also pointed out that the specimen's
provenance Provenance (from the French ''provenir'', 'to come from/forth') is the chronology of the ownership, custody or location of a historical object. The term was originally mostly used in relation to works of art but is now used in similar senses i ...
was incorrect; instead of being from younger rocks than ''Kritosaurus'', it is from rocks of about the same age. The next major review, in the second edition of ''The Dinosauria'', again separated the two. The absence of the beak and lower jaw in the partial skull of ''Naashoibitosaurus'' makes comparison even more difficult. A 2014 study agreed with previous authors that ''Naashoibitosaurus'' is similar to ''Kritosaurus'', but found it to be a distinct species.


Paleobiology

As a hadrosaurid, ''Naashoibitosaurus'' would have been a large bipedal/ quadrupedal
herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
, eating
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...
s with a sophisticated skull that permitted a grinding motion analogous to chewing. Its
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
were continually replacing and packed into
dental batteries Dinosaur teeth have been studied since 1822 when Mary Ann Mantell (1795-1869) and her husband Dr Gideon Algernon Mantell (1790-1852) discovered an ''Iguanodon'' tooth in Sussex in England. Unlike mammal teeth, individual dinosaur teeth are gener ...
that contained hundreds of teeth, only a relative handful of which were in use at any time. Plant material would have been cropped by its broad beak, and held in the jaws by a cheek-like organ. Feeding would have been from the ground up to ~4 meters (13 ft) above. If it was a separate genus, how it would have partitioned resources with the similar ''Kritosaurus'' is unknown.


See also

*
Timeline of hadrosaur research A timeline is a display of a list of events in chronological order. It is typically a graphic design showing a long bar labelled with dates paralleling it, and usually contemporaneous events. Timelines can use any suitable scale representin ...


References


External links


Status of fossil speciesPDF file on hadrosaurs
{{Taxonbar, from=Q744334 Late Cretaceous dinosaurs of North America Saurolophines Taxa named by Spencer G. Lucas Fossil taxa described in 1993 Paleontology in New Mexico Campanian genus first appearances Campanian genus extinctions Ornithischian genera