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''Moringa'' is the sole
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
in the plant
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Moringaceae. It contains 13 species from
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
and
subtropical The subtropical zones or subtropics are geographical and climate zones to the north and south of the tropics. Geographically part of the temperate zones of both hemispheres, they cover the middle latitudes from to approximately 35° north a ...
regions of Africa and Asia that range in size from tiny
herbs In general use, herbs are a widely distributed and widespread group of plants, excluding vegetables and other plants consumed for macronutrients, with savory or aromatic properties that are used for flavoring and garnishing food, for medicina ...
to massive trees. ''Moringa'' species grow quickly in many types of environments. The most widely cultivated species is ''
Moringa oleifera ''Moringa oleifera'' is a fast-growing, drought-resistant tree of the family Moringaceae, native to the Indian subcontinent. Common names include moringa, drumstick tree (from the long, slender, triangular seed-pods), horseradish tree (from t ...
'', native to the foothill of the
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 10 ...
in northwestern India, a
multipurpose tree Multipurpose trees or multifunctional trees are trees that are deliberately grown and managed for more than one output. They may supply food in the form of fruit, nut (fruit), nuts, or leaves that can be used as a vegetable; while at the same tim ...
cultivated throughout the tropics and marketed as a
dietary supplement A dietary supplement is a manufactured product intended to supplement one's diet by taking a pill, capsule, tablet, powder, or liquid. A supplement can provide nutrients either extracted from food sources or that are synthetic in order ...
,
health food A healthy diet is a diet that maintains or improves overall health. A healthy diet provides the body with essential nutrition: fluid, macronutrients such as protein, micronutrients such as vitamins, and adequate fibre and food energy. A health ...
or source for herbalism practices. The fruit pods of ''Moringa oleifera'' ("drumsticks") are increasingly consumed as food in many parts of the world, but particularly in South Asia. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
are commonly used to make tea.
Oils An oil is any nonpolar chemical substance that is composed primarily of hydrocarbons and is hydrophobic (does not mix with water) & lipophilic (mixes with other oils). Oils are usually flammable and surface active. Most oils are unsaturate ...
are made from the seeds, while powders can be made from the leaves and
roots A root is the part of a plant, generally underground, that anchors the plant body, and absorbs and stores water and nutrients. Root or roots may also refer to: Art, entertainment, and media * ''The Root'' (magazine), an online magazine focusing ...
. '' M. stenopetala'', an African species, is also widely grown, but to a much lesser extent than ''M. oleifera''. The genus name is derived from ''murungai'', the
Tamil Tamil may refer to: * Tamils, an ethnic group native to India and some other parts of Asia ** Sri Lankan Tamils, Tamil people native to Sri Lanka also called ilankai tamils **Tamil Malaysians, Tamil people native to Malaysia * Tamil language, na ...
word for drumstick, and in India the plant is commonly referred to as the drumstick tree.


Description

''Moringa'' is considered one of the most widely diverse genera for its size ranging from small shrubs ('' M. pygmaea'') to large
pachycaul Pachycauls are plants with a disproportionately thick trunk for their height, and few branches. This can be the product of exceptional primary growth (as with palms and cycads) or disproportioate secondary growth as with Adansonia. The word is d ...
trees ('' M. ovalifolia'').


Growth habit

''Moringa'' contains a wide range of growth
habits A habit (or wont as a humorous and formal term) is a routine of behavior that is repeated regularly and tends to occur subconsciously.
that may be subdivided into the following categories: * Bottle (pachycaul) trees: '' M. drouhardii, M. hildebrandtii,'' '' M. ovalifolia, M. stenopetala'' * Slender trees: '' M. concanensis, M. oleifera'', '' M. peregrina'' * Sarcorhizal trees: '' M. arborea, M. ruspoliana'' *
Tuber Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growing ...
ous shrubs: '' M. borziana, M. longituba, M. pygmaea, M. rivae''


Leaves

Leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
are typically pinnately compound with entire margins.


Flowers

Flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s may be either bilaterally or radially symmetric. Bottle trees typically produce small, radially symmetric flowers, while other members of the genus produce radially symmetric flowers. Most flowers range in color from white to cream to brown with the notable exception of '' M. longituba'' which produces bright red flowers.


Fruit

Fruits are typically elongate, slender, 3-valved " pods" resembling an
indehiscent Dehiscence is the splitting of a mature plant structure along a built-in line of weakness to release its contents. This is common among fruits, anthers and sporangia. Sometimes this involves the complete detachment of a part; structures that op ...
silique A silique or siliqua (plural ''siliques'' or ''siliquae'') is a type of fruit (seed capsule) having two fused carpels with the length being more than three times the width. When the length is less than three times the width of the dried fruit i ...
(in contrast with a true dehiscent silique). Fruits of ''M. oleifera'' ( drumstick), are a major agricultural product of
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
, eaten as a
vegetable Vegetables are parts of plants that are consumed by humans or other animals as food. The original meaning is still commonly used and is applied to plants collectively to refer to all edible plant matter, including the edible flower, flowers, ...
and used for
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...
.


Phytochemistry

''Moringa'' contain a number of
sulfurous Sulfur (or sulphur in British English) is a chemical element with the symbol S and atomic number 16. It is abundant, multivalent and nonmetallic. Under normal conditions, sulfur atoms form cyclic octatomic molecules with a chemical formula ...
biochemical compounds called "mustard-oil glycosides" or
glucosinolate Glucosinolates are natural components of many pungent plants such as mustard, cabbage, and horseradish. The pungency of those plants is due to mustard oils produced from glucosinolates when the plant material is chewed, cut, or otherwise damage ...
s commonly found in
cruciferous Cruciferous vegetables are vegetables of the family Brassicaceae (also called Cruciferae) with many genera, species, and cultivars being raised for food production such as cauliflower, cabbage, kale, garden cress, bok choy, broccoli, Brussels spr ...
vegetables of
Brassicaceae Brassicaceae () or (the older) Cruciferae () is a medium-sized and economically important family of flowering plants commonly known as the mustards, the crucifers, or the cabbage family. Most are herbaceous plants, while some are shrubs. The leav ...
. Benzyl glucosinolate along with family-specific glucomoringin and glucosoonjnain have been detected from various ''Moringa'' species and are thought to be the cause of the bitter taste in some ''Moringa'' leaves.


Taxonomy


Higher-level classification

The
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...
family, Moringaceae, containing genus ''Moringa'' has been placed in the
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of ...
Brassicales The Brassicales (or Cruciales) are an order of flowering plants, belonging to the eurosids II group of dicotyledons under the APG II system. One character common to many members of the order is the production of glucosinolate (mustard oil) compo ...
according to most modern taxonomic systems, including the
APG IV system The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was published ...
. Molecular data has suggested a close relationship between Moringaceae and
Caricaceae The Caricaceae are a family of flowering plants in the order Brassicales, found primarily in tropical regions of Central and South America and Africa. They are usually short-lived evergreen pachycaul shrubs or small to medium-sized trees growing ...
with many identifying a "Caricaceae-Moringaceae" clade within Brassicales. Prior to the availability of molecular data, morphological classification of Moringaceae placed the family in either Brassicales or
Sapindales Sapindales is an order of flowering plants. Well-known members of Sapindales include citrus; maples, horse-chestnuts, lychees and rambutans; mangos and cashews; frankincense and myrrh; mahogany and neem. The APG III system of 2009 includes it i ...
due to the unusual morphological diversity of the family.


Classification within the genus

''Moringa'' contains three widely recognized clades—Donaldsonia, Moringa, and Dysmoringa. Donaldsonia, once thought to be a
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between ...
of ''Moringa'', is a non-monophyletic clade identifiable by radially symmetric flowers and containing the bottle trees '' M. drouhardii, M. hildebrandtii,'' '' M. ovalifolia,'' and '' M. stenopetala.'' The Moringa clade contains all other members of genus ''Moringa'' (except ''M. longituba'') characterized by irregular floral symmetry, perigynous flowers, and short receptacles. The Dysmoringa clade contains the species ''M. longituba'' which diverges from common Moringa clade characteristics due to its long receptacle and red flowers. The exact phylogenetic relationship between members of ''Moringa'' continues to evolve with growing molecular data, though the Donaldsonia clade is consistently identified as the basalmost clade within the family.


List of species

* '' Moringa arborea'' Verdc. (indigenous to
Kenya ) , national_anthem = " Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
) * '' Moringa borziana'' Mattei (indigenous to
Somalia Somalia, , Osmanya script: 𐒈𐒝𐒑𐒛𐒐𐒘𐒕𐒖; ar, الصومال, aṣ-Ṣūmāl officially the Federal Republic of SomaliaThe ''Federal Republic of Somalia'' is the country's name per Article 1 of thProvisional Constitut ...
) * ' Nimmo (indigenous to northern
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
) * '' Moringa drouhardii'' Jum. – bottle tree (indigenous to southwestern
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Afric ...
) * '' Moringa hildebrandtii'' Engl. – Hildebrandt's moringa (indigenous to southwestern Madagascar) * '' Moringa longituba'' Engl. (indigenous to
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
and Somalia) * ''
Moringa oleifera ''Moringa oleifera'' is a fast-growing, drought-resistant tree of the family Moringaceae, native to the Indian subcontinent. Common names include moringa, drumstick tree (from the long, slender, triangular seed-pods), horseradish tree (from t ...
'' Lam. (syn. ''M. pterygosperma'') – horseradish tree (indigenous to northwestern
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
) * '' Moringa ovalifolia'' Dinter &
Berger Berger is a surname in both German and French, although there is no etymological connection between the names in the two languages. The French surname is an occupational name for a shepherd, from Old French ''bergier'' (Late Latin ''berbicarius'', ...
(indigenous to
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
and
Angola , national_anthem = "Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordinat ...
) * ''
Moringa peregrina ''Moringa peregrina'' is a species of flowering plant in the family Moringaceae that is native to Arabian Peninsula Horn of Africa and Southern Sinai, Egypt and can be found throughout Arabia, Egypt and as far north as Syria. Moringa peregrin ...
'' ( Forssk.) Fiori indigenous to Arabian Peninsula
Horn of Africa The Horn of Africa (HoA), also known as the Somali Peninsula, is a large peninsula and geopolitical region in East Africa.Robert Stock, ''Africa South of the Sahara, Second Edition: A Geographical Interpretation'', (The Guilford Press; 2004 ...
and in the Southern Sinai,
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a List of transcontinental countries, transcontinental country spanning the North Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via a land bridg ...
* '' Moringa pygmaea'' Verdc. (indigenous to Somalia) * '' Moringa rivae''
Chiov. __NOTOC__ ''Emilio Chiovenda'' (18 May 1871 – 19 February 1941) was an Italian botanist. Chiovenda was born in Rome in 1871 to a family originating from rural Piedmont. He was educated at the Collegio Rosmini in Stresa and Domodossola College be ...
(indigenous to Kenya and Ethiopia) * '' Moringa ruspoliana'' Engl. (indigenous to Ethiopia) * ''
Moringa stenopetala ''Moringa stenopetala'', commonly known as the African Moringa or cabbage tree, is a deciduous tree in the plant genus ''Moringa'', native to Kenya and Ethiopia. A drought-resistant species, it is characterized by its bottle-shaped trunk, long ...
'' ( Baker f.) Cufod. (indigenous to Kenya and Ethiopia)


References


External links


Moringa Home Page
Instituto de Biología, Universidad Nacional Autónoma de México
The International Moringa Germplasm Collection
{{Taxonbar, from=Q310468 Brassicales genera Edible plants