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Miklos Dezso Ferenc Udvardy (March 23, 1919 – January 27, 1998) was a Hungarian-American
biologist A biologist is a scientist who conducts research in biology. Biologists are interested in studying life on Earth, whether it is an individual cell, a multicellular organism, or a community of interacting populations. They usually specialize ...
and university instructor. He contributed to a wide variety of disciplines, such as
biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
,
evolutionary biology Evolutionary biology is the subfield of biology that studies the evolutionary processes (natural selection, common descent, speciation) that produced the diversity of life on Earth. It is also defined as the study of the history of life ...
,
ornithology Ornithology is a branch of zoology that concerns the "methodological study and consequent knowledge of birds with all that relates to them." Several aspects of ornithology differ from related disciplines, due partly to the high visibility and t ...
, and
vegetation classification Vegetation classification is the process of classifying and mapping the vegetation over an area of the earth's surface. Vegetation classification is often performed by state based agencies as part of land use, resource and environmental management. ...
, contributing 191 papers, 8 books, and 3 maps to the scientific literature.


Early life and career

Udvardy was born on March 23, 1919, in
Debrecen Debrecen ( , is Hungary's second-largest city, after Budapest, the regional centre of the Northern Great Plain region and the seat of Hajdú-Bihar County. A city with county rights, it was the largest Hungarian city in the 18th century and ...
,
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Cr ...
into a noble family. His father was the nobleman Miklos Udvardy de Udvardy et Básth, and his mother was Elizabeth Komlossy de Komlos. Despite an early interest in birds, his father encouraged him to study law, but he later went on to earn a doctorate in
biology Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditary ...
from the
University of Debrecen ThUniversity of Debrecen( hu, Debreceni Egyetem) is a university located in Debrecen, Hungary. It is the oldest continuously operating institution of higher education in Hungary ever since its establishment in 1538 by Suleiman the Magnificent f ...
in 1942. His first position was as research biologist at the Tihanyi Biological Station on
Lake Balaton Lake Balaton () is a freshwater lake in the Transdanubian region of Hungary. It is the largest lake in Central Europe, and one of the region's foremost tourist destinations. The Zala River provides the largest inflow of water to the lake, and ...
in western Hungary. Udvardy left Hungary in 1948, and secured a postdoctoral fellowship at
Helsinki University The University of Helsinki ( fi, Helsingin yliopisto, sv, Helsingfors universitet, abbreviated UH) is a public research university located in Helsinki, Finland since 1829, but founded in the city of Turku (in Swedish ''Åbo'') in 1640 as the ...
in Finland with Professor Pontus Palmgren ( fi), an ornithologist and zoologist. He met his future wife Maud Björklund in 1950, and moved to
Uppsala Uppsala (, or all ending in , ; archaically spelled ''Upsala'') is the county seat of Uppsala County and the fourth-largest city in Sweden, after Stockholm, Gothenburg, and Malmö. It had 177,074 inhabitants in 2019. Located north of the ca ...
in Sweden. He was Curator of Marine Invertebrates at the
Swedish Museum of Natural History The Swedish Museum of Natural History ( sv, Naturhistoriska riksmuseet, literally, the National Museum of Natural History), in Stockholm, is one of two major museums of natural history in Sweden, the other one being located in Gothenburg. The ...
in Stockholm for a brief time.


Later career

Miklos and Maud married in 1951 and emigrated to Canada. Udvardy arrived at the
University of British Columbia The University of British Columbia (UBC) is a public research university with campuses near Vancouver and in Kelowna, British Columbia. Established in 1908, it is British Columbia's oldest university. The university ranks among the top thr ...
in
Vancouver, British Columbia Vancouver ( ) is a major city in western Canada, located in the Lower Mainland region of British Columbia. As the most populous city in the province, the 2021 Canadian census recorded 662,248 people in the city, up from 631,486 in 2016. The ...
in 1952 and was appointed Assistant Professor in the Department of Zoology in 1953, where he lectured in comparative anatomy and
ornithology Ornithology is a branch of zoology that concerns the "methodological study and consequent knowledge of birds with all that relates to them." Several aspects of ornithology differ from related disciplines, due partly to the high visibility and t ...
until 1966. He was a visiting professor at the
University of Hawaii A university () is an institution of higher (or tertiary) education and research which awards academic degrees in several academic disciplines. Universities typically offer both undergraduate and postgraduate programs. In the United States, th ...
during the 1958–1959 academic year, and was Lida Scott Brown Lecturer in Ornithology at the
University of California, Los Angeles The University of California, Los Angeles (UCLA) is a public land-grant research university in Los Angeles, California. UCLA's academic roots were established in 1881 as a teachers college then known as the southern branch of the Californ ...
in the 1963–1964 academic year. He served as Professor of Biological Sciences at
California State University, Sacramento California State University, Sacramento (CSUS, Sacramento State, or informally Sac State) is a public university in Sacramento, California. Founded in 1947 as Sacramento State College, it is the eleventh oldest school in the 23-campus California ...
(CSUS) from 1967 until his retirement in 1991. There he sponsored eight graduate students, and spent a year as Visiting Professor at the
University of Bonn The Rhenish Friedrich Wilhelm University of Bonn (german: Rheinische Friedrich-Wilhelms-Universität Bonn) is a public research university located in Bonn, North Rhine-Westphalia, Germany. It was founded in its present form as the ( en, Rhine ...
in Germany, and a year as Fulbright Lecturer at the National Autonomous University of Honduras. Udvardy served as a mentor to an impressive array of graduate students, including at UBC Peter Grant (Princeton), Jean Bedard (Universite Laval, Quebec), Spencer Sealy (University of Manitoba), Kees Vermeer (Canadian Wildlife Service), and at CSUS Raymond Pierotti (University of Kansas). He was an eclectic scholar, teaching biogeography, evolution, ornithology, speciation, among other courses at CSUS. A graduate student award at CSUS was created to honor Udvardy's memory. Udvardy's work in
biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
was seminal to the field, and his system of
biogeographic realm A biogeographic realm or ecozone is the broadest biogeographic division of Earth's land surface, based on distributional patterns of terrestrial organisms. They are subdivided into bioregions, which are further subdivided into ecoregions. ...
s is the basis for the system widely used today. His ''Map of the Biogeographic Provinces of the World'' was an important contribution to thinking in biogeography. Working with
UNESCO The United Nations Educational, Scientific and Cultural Organization is a List of specialized agencies of the United Nations, specialized agency of the United Nations (UN) aimed at promoting world peace and security through international coope ...
, Udvardy produced a crucial document for conservation entitled ''A Classification of the Biogeographic Provinces of the World'', which was meant to complement the map. Udvardy came up with the idea of
competitive exclusion In ecology, the competitive exclusion principle, sometimes referred to as Gause's law, is a proposition that two species which compete for the same limited resource cannot coexist at constant population values. When one species has even the sligh ...
, but did not use the term by which this idea later became known, so his accomplishment is not widely recognized. He was one of the first to emphasize the role of humans in species dispersal and in introducing species into new environments and the possible consequences of such acts. Udvardy coined the term "
zoogeography Zoogeography is the branch of the science of biogeography that is concerned with geographic distribution (present and past) of animal species. As a multifaceted field of study, zoogeography incorporates methods of molecular biology, genetics, mor ...
" to identify studies of the distribution of animals, and published the book ''Dynamic Zoogeography'' on the topic. Miklos Udvardy died of surgical complications in
Sacramento, California ) , image_map = Sacramento County California Incorporated and Unincorporated areas Sacramento Highlighted.svg , mapsize = 250x200px , map_caption = Location within Sacramento C ...
, on January 27, 1998. He was survived by his wife, Maud, and their three children and two grandchildren.


See also

*
Alfred Russel Wallace Alfred Russel Wallace (8 January 1823 – 7 November 1913) was a British natural history, naturalist, explorer, geographer, anthropologist, biologist and illustrator. He is best known for independently conceiving the theory of evolution thro ...
, who also played a large role in developing the idea of
biogeographic realm A biogeographic realm or ecozone is the broadest biogeographic division of Earth's land surface, based on distributional patterns of terrestrial organisms. They are subdivided into bioregions, which are further subdivided into ecoregions. ...
s


Selected publications

* Udvardy, M. D. F. (1975). ''A classification of the biogeographical provinces of the world''. IUCN Occasional Paper no. 18. Morges, Switzerland: IUCN

* Udvardy, Miklos D. F. (1975) "World Biogeographical Provinces" (Map). The CoEvolution Quarterly, Sausalito, California.


External links


In memoriam: Miklos D. F. Udvardy, 1919–1998
{{DEFAULTSORT:Udvardy, Miklos 1919 births 1998 deaths 20th-century American zoologists 20th-century Hungarian zoologists American ornithologists Biogeographers Hungarian ornithologists Universidad Nacional Autónoma de Honduras faculty University of British Columbia faculty University of California, Los Angeles faculty